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<document id="CDD3812F609B31364672F0A9F87355CF" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A21BB01FEBDE762E5722EAF" docLanguage="en" docName="J_NAT_HIST.53.37–38.2231–2351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) fonsecae" docType="treatment" docVersion="18" lastPageNumber="2298" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2297" updateTime="1698782751744" updateUser="ExternalLinkService">
<mods:mods id="871AB749CAD1A141FCEE24E8BD03F4C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="19D9C6A7E6F6DBA189C098AF214C2AF4">
<mods:title id="57A097A1E28F75CC45D982C7D562F52E">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
</mods:titleInfo>
<mods:name id="7451E72B0828E617CBC30FFA04BB564C" type="personal">
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<mods:namePart id="67EBDC01B894311F0F460C3AF5CAC690">Nunes, Rafael V.</mods:namePart>
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<mods:namePart id="CF5E931FD103B1C26AF01646A7FDB611">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="2C0A53A26EF3EF9AE04C6C3271C55D57">text</mods:typeOfResource>
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<mods:title id="2D672BE348471F63FF53D236E5835C81">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="0D430B3A8FE39B6A44BCC7A0840C1C04">
<mods:date id="A5FF14EA3A20C3B0C11D86AB0F6A1306">2019</mods:date>
<mods:detail id="6447F0DBF0BE2616A1ACAB9B4085500A" type="pubDate">
<mods:number id="DA2733E69F76CC16E7F657D9A0016A5A">2020-01-13</mods:number>
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<mods:number id="1D970C4FB14EC315DCA86627F5FE71A3">53</mods:number>
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<mods:classification id="620D6391C2BE00CF792954A6A06DC721">journal article</mods:classification>
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<treatment id="C740D6092A21BB01FEBDE762E5722EAF" ID-DOI="http://doi.org/10.5281/zenodo.3671884" ID-GBIF-Taxon="162116293" ID-Zenodo-Dep="3671884" LSID="urn:lsid:plazi:treatment:C740D6092A21BB01FEBDE762E5722EAF" httpUri="http://treatment.plazi.org/id/C740D6092A21BB01FEBDE762E5722EAF" lastPageId="68" lastPageNumber="2298" pageId="67" pageNumber="2297">
<subSubSection id="07F334942A21BB06FEBDE762E6B82E7B" pageId="67" pageNumber="2297" type="nomenclature">
<paragraph id="4F56671F2A21BB06FEBDE762E6B82E7B" blockId="67.[287,1028,540,600]" pageId="67" pageNumber="2297">
<heading id="141ED0732A21BB06FEBDE762E6B82E7B" centered="true" fontSize="11" level="2" pageId="67" pageNumber="2297" reason="5">
2.2.:
<taxonomicName id="88E91C9C2A21BB06FEFAE762E0682E15" ID-CoL="6D43J" authority="(Luederwaldt 1926)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1926" box="[344,1028,540,566]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="2297" phylum="Arthropoda" rank="species" species="fonsecae" subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A21BB06FEFAE762E5822E15" box="[344,494,540,566]" italics="true" pageId="67" pageNumber="2297">Dichotomius</emphasis>
(
<emphasis id="7D9DBB0D2A21BB06FE5CE762E6FF2E15" box="[510,659,540,566]" italics="true" pageId="67" pageNumber="2297">Cephagonus</emphasis>
)
<emphasis id="7D9DBB0D2A21BB06FD06E762E7612E15" box="[676,781,540,566]" italics="true" pageId="67" pageNumber="2297">fonsecae</emphasis>
(
<bibRefCitation id="2B781AEE2A21BB06FCB9E762E7902E15" author="Luederwaldt H" box="[795,1020,540,566]" pageId="67" pageNumber="2297" pagination="135 - 138" refId="ref53236" refString="Luederwaldt H. 1926. Novas especies do genero &quot; Pinotus &quot;. Rev Museu Nacional. 14: 135 - 138." type="journal article" year="1926">Luederwaldt 1926</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="D7D27B9A2A21BB06FDF5E740E6A02E7B" box="[599,716,574,600]" captionStart="Figure 21" captionStartId="36.[160,225,1600,1623]" captionTargetBox="[163,1152,142,1569]" captionTargetId="figure@36.[163,1153,142,1569]" captionTargetPageId="36" captionText="Figure 21. D. fonsecae: (a) male dorsal habitus; (b) Holotype dorsal habitus; (c) holotype labels; (d) female dorsal view; (e) female ventral view; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665557" httpUri="https://zenodo.org/record/3665557/files/figure.png" pageId="67" pageNumber="2297">Figure 21</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A21BB06FF02E70AE06C2EF2" pageId="67" pageNumber="2297" type="reference_group">
<paragraph id="4F56671F2A21BB06FF02E70AE7632EAF" blockId="67.[160,1024,628,721]" box="[160,783,628,652]" pageId="67" pageNumber="2297">
<taxonomicName id="88E91C9C2A21BB06FF02E70AE6452EAF" authority="Luederwaldt 1926" authorityName="Luederwaldt" authorityYear="1926" box="[160,553,628,652]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="2297" phylum="Arthropoda" rank="species" species="fonsecae">
<emphasis id="7D9DBB0D2A21BB06FF02E70AE53B2EAF" box="[160,343,628,652]" italics="true" pageId="67" pageNumber="2297">Pinotus fonsecae</emphasis>
<bibRefCitation id="2B781AEE2A21BB06FEFCE70AE6452EAF" author="Luederwaldt H" box="[350,553,628,652]" pageId="67" pageNumber="2297" pagination="135 - 138" refId="ref53236" refString="Luederwaldt H. 1926. Novas especies do genero &quot; Pinotus &quot;. Rev Museu Nacional. 14: 135 - 138." type="journal article" year="1926">Luederwaldt 1926</bibRefCitation>
</taxonomicName>
ı p. 135 (description)
</paragraph>
<paragraph id="4F56671F2A21BB06FF02E7E9E7722E8C" blockId="67.[160,1024,628,721]" box="[160,798,663,687]" pageId="67" pageNumber="2297">
<taxonomicName id="88E91C9C2A21BB06FF02E7E9E5982E8C" authorityName="Luederwaldt" authorityYear="1926" box="[160,500,663,687]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="2297" phylum="Arthropoda" rank="species" species="fonsecae" subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A21BB06FF02E7E9E49C2E8C" box="[160,240,663,687]" italics="true" pageId="67" pageNumber="2297">Pinotus</emphasis>
(
<emphasis id="7D9DBB0D2A21BB06FF5DE7E9E5EA2E8C" box="[255,390,663,687]" italics="true" pageId="67" pageNumber="2297">Cephagonus</emphasis>
)
<emphasis id="7D9DBB0D2A21BB06FE37E7E9E5982E8C" box="[405,500,663,687]" italics="true" pageId="67" pageNumber="2297">fonsecae</emphasis>
</taxonomicName>
ı
<bibRefCitation id="2B781AEE2A21BB06FDA3E7E9E6A02E8C" author="Luederwaldt H" box="[513,716,663,687]" pageId="67" pageNumber="2297" pagination="603 - 776" refId="ref53282" refString="Luederwaldt H. 1929. As especies brasileiras do genero. Pinotus. Rev Museu Paulista. 16: 603 - 776." type="journal article" year="1929">
Luederwaldt
<date id="3B5741DF2A21BB06FD37E7E9E6A02E8C" box="[661,716,663,687]" pageId="67" pageNumber="2297" value="1929-01">1929</date>
</bibRefCitation>
ı p. 110
</paragraph>
<paragraph id="4F56671F2A21BB06FF02E7C7E06C2EF2" blockId="67.[160,1024,628,721]" box="[160,1024,697,721]" pageId="67" pageNumber="2297">
<taxonomicName id="88E91C9C2A21BB06FF02E7C7E5AA2EF2" ID-CoL="8CYB" authority="" authorityName="Burmeister" authorityYear="1846" box="[160,454,697,721]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="2239" phylum="Arthropoda" rank="subGenus" subGenus="Selenocopris">
<emphasis id="7D9DBB0D2A21BB06FF02E7C7E5452EF2" box="[160,297,697,721]" italics="true" pageId="67" pageNumber="2297">Dichotomius</emphasis>
(
<emphasis id="7D9DBB0D2A21BB06FE9AE7C7E5D32EF2" box="[312,447,697,721]" italics="true" pageId="67" pageNumber="2297">selenocopris</emphasis>
)
</taxonomicName>
sp.ı Sarmiento-Garcés &amp;
<bibRefCitation id="2B781AEE2A21BB06FD7CE7C7E7CB2EF2" author="Sarmiento Garces Ri Amat-Garcia G" box="[734,935,697,721]" pageId="67" pageNumber="2297" pagination="4" refId="ref54224" refString="Sarmiento Garces Ri Amat-Garcia G. 2014. Escarabajos del Genero Dichotomius Hope 1838 (scarabaeidae: scarabaeinae) en Colombia. Fauna de Colombia. Monografia n 3 4. Bogota: Universidad Nacional de Colombia." type="journal article" year="2014">
Amat-Garcia
<date id="3B5741DF2A21BB06FCD2E7C7E7CB2EF2" box="[880,935,697,721]" pageId="67" pageNumber="2297" value="2014-01">2014</date>
</bibRefCitation>
ı p. 104.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A21BB06FF02E780E6422852" pageId="67" pageNumber="2297" type="diagnosis">
<paragraph id="4F56671F2A21BB06FF02E780E01A2FC4" blockId="67.[160,1155,766,999]" pageId="67" pageNumber="2297">
<emphasis id="7D9DBB0D2A21BB06FF02E780E5762F34" bold="true" box="[160,282,766,791]" italics="true" pageId="67" pageNumber="2297">Diagnosis.</emphasis>
distinguishable from the other species in the group by the following combination of characteristics: males
<emphasis id="7D9DBB0D2A21BB06FDA2E65FE66A2F1A" box="[512,518,801,825]" italics="true" pageId="67" pageNumber="2297">’</emphasis>
cephalic carina longer than wideı emarginated at apex producing two tubercles (as in
<taxonomicName id="88E91C9C2A21BB06FE5BE63AE6232F7F" authorityName="Haroldi" authorityYear="1867" box="[505,591,836,860]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="2297" phylum="Arthropoda" rank="species" species="fissus">
<emphasis id="7D9DBB0D2A21BB06FE5BE63AE6232F7F" box="[505,591,836,860]" italics="true" pageId="67" pageNumber="2297">D. fissus</emphasis>
</taxonomicName>
of the same group); anterior and posterior portion of the pronotum lacking distinct punctuation (at most some fineı sparse punctures as in
<taxonomicName id="88E91C9C2A21BB06FF02E6F4E49C2F81" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[160,240,906,930]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="2297" phylum="Arthropoda" rank="species" species="seag">
<emphasis id="7D9DBB0D2A21BB06FF02E6F4E49C2F81" box="[160,240,906,930]" italics="true" pageId="67" pageNumber="2297">D. seag</emphasis>
</taxonomicName>
of the same group); males
<emphasis id="7D9DBB0D2A21BB06FDB9E6F7E64D2F82" box="[539,545,905,929]" italics="true" pageId="67" pageNumber="2297">’</emphasis>
pronotal disc with a pair of acute tubercles at anterior portion; pronotal lateral borders almost completely smooth
<emphasis id="7D9DBB0D2A21BB06FCF1E6D2E70C2FE7" box="[851,864,940,964]" italics="true" pageId="67" pageNumber="2297">–</emphasis>
except for a slightly irregular portion near medio-lateral pronotal pit and elytra lacking distinct microsculpture.
</paragraph>
<paragraph id="4F56671F2A21BB06FF1CE16BE6422852" blockId="67.[190,1155,1044,1137]" pageId="67" pageNumber="2297">
<emphasis id="7D9DBB0D2A21BB06FF1CE16BE4A0280A" bold="true" box="[190,204,1045,1065]" pageId="67" pageNumber="2297">●</emphasis>
<typeStatus id="9052D9BD2A21BB06FF7AE16AE53E280F" box="[216,338,1044,1068]" pageId="67" pageNumber="2297" type="holotype">HOLOTYPE</typeStatus>
of
<taxonomicName id="88E91C9C2A21BB06FED7E16AE647280F" authorityName="Luederwaldt" authorityYear="1926" box="[373,555,1044,1068]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="2297" phylum="Arthropoda" rank="species" species="fonsecae">
<emphasis id="7D9DBB0D2A21BB06FED7E16AE647280F" box="[373,555,1044,1068]" italics="true" pageId="67" pageNumber="2297">Pinotus fonsecae</emphasis>
</taxonomicName>
: O.
<collectingCountry id="37FE278F2A21BB06FDF7E16AE6A5280F" box="[597,713,1044,1068]" name="Ecuador" pageId="67" pageNumber="2297">ECUADOR</collectingCountry>
. Macas. Feyer {whiteı printed}/TIPO {red with black marginsı printed}/
<taxonomicName id="88E91C9C2A21BB06FDB9E149E704286C" authority="Lud. Lud Det" authorityName="Lud. Lud Det" box="[539,872,1079,1103]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="2297" phylum="Arthropoda" rank="species" species="fonsecae">
<emphasis id="7D9DBB0D2A21BB06FDB9E149E6BD286C" box="[539,721,1079,1103]" italics="true" pageId="67" pageNumber="2297">Pinotus fonsecae</emphasis>
Lüd. Lüd Det
</taxonomicName>
23. {whiteı handwritten}/ 25904 {whiteı printed} [MZSP].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A21BB06FF02E1E0E07428DF" pageId="67" pageNumber="2297" type="materials_examined">
<paragraph id="4F56671F2A21BB06FF02E1E0E07428DF" blockId="67.[160,1156,1182,1276]" pageId="67" pageNumber="2297">
<emphasis id="7D9DBB0D2A21BB06FF02E1E0E5FA2894" bold="true" box="[160,406,1182,1207]" italics="true" pageId="67" pageNumber="2297">
Non
<typeStatus id="9052D9BD2A21BB06FF47E1E0E5742894" box="[229,280,1182,1207]" pageId="67" pageNumber="2297">type</typeStatus>
material.
</emphasis>
<collectingCountry id="37FE278F2A21BB06FE04E1E1E6482894" box="[422,548,1183,1207]" name="Colombia" pageId="67" pageNumber="2297">COLOMBIA</collectingCountry>
:
<emphasis id="7D9DBB0D2A21BB06FD99E1E1E6F22894" bold="true" box="[571,670,1183,1207]" pageId="67" pageNumber="2297">
<collectingRegion id="8D2DA9FD2A21BB06FD99E1E1E6F22894" box="[571,670,1183,1207]" country="Colombia" name="Caqueta" pageId="67" pageNumber="2297">Caquetá</collectingRegion>
</emphasis>
. Solano. Parque Nacional Chiribiquete. 0³10
<emphasis id="7D9DBB0D2A21BB06FF70E1BCE4B428F9" bold="true" box="[210,216,1218,1242]" italics="true" pageId="67" pageNumber="2297">ʹ</emphasis>
54
<emphasis id="7D9DBB0D2A21BB06FF56E1BCE49128F9" box="[244,253,1218,1242]" italics="true" pageId="67" pageNumber="2297">”</emphasis>
Nı 72
<geoCoordinate id="2ADD01D82A21BB06FE9EE1BCE5C828F9" box="[316,420,1218,1242]" degrees="3" direction="west" minutes="36" orientation="longitude" pageId="67" pageNumber="2297" precision="15" seconds="57" value="-3.6158333">
³36
<emphasis id="7D9DBB0D2A21BB06FEC2E1BCE50A28F9" bold="true" box="[352,358,1218,1242]" italics="true" pageId="67" pageNumber="2297">ʹ</emphasis>
57
<emphasis id="7D9DBB0D2A21BB06FE23E1BCE5E628F9" box="[385,394,1218,1242]" italics="true" pageId="67" pageNumber="2297">”</emphasis>
W
</geoCoordinate>
. 800 mosl. E Gonzalez [1 Ƌ at
<collectionCode id="29F8FFDA2A21BB06FCB9E1BCE70C28F9" box="[795,864,1218,1242]" pageId="67" pageNumber="2297">CEMT</collectionCode>
];
<collectingCountry id="37FE278F2A21BB06FCD1E1BCE7C228F9" box="[883,942,1218,1242]" name="Peru" pageId="67" pageNumber="2297">PERU</collectingCountry>
[1 Ƌ at UNMSM];
<emphasis id="7D9DBB0D2A21BB06FF02E19AE48D28DF" bold="true" box="[160,225,1252,1276]" pageId="67" pageNumber="2297">
<collectingRegion id="8D2DA9FD2A21BB06FF02E19AE48D28DF" box="[160,225,1252,1276]" country="Peru" name="Junin" pageId="67" pageNumber="2297">Júnin</collectingRegion>
</emphasis>
: Satipo. 1100 mosl. Fezes humanas. A Santibañez [3 ƋƋı
<specimenCount id="59EFAC962A21BB06FCCAE19AE7C928DF" box="[872,933,1252,1276]" pageId="67" pageNumber="2297" type="female">2 ♀♀</specimenCount>
at
<collectionCode id="29F8FFDA2A21BB06FC68E19AE06328DF" box="[970,1039,1252,1276]" pageId="67" pageNumber="2297">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A21BB01FF02E057E5C42E07" lastPageId="68" lastPageNumber="2298" pageId="67" pageNumber="2297" type="description">
<paragraph id="4F56671F2A21BB01FF02E057E59F2D2D" blockId="67.[160,1156,1321,1727]" lastBlockId="68.[160,1156,142,270]" lastPageId="68" lastPageNumber="2298" pageId="67" pageNumber="2297">
<emphasis id="7D9DBB0D2A21BB06FF02E057E52A2961" bold="true" box="[160,326,1321,1346]" italics="true" pageId="67" pageNumber="2297">Redescription.</emphasis>
Colour: blackı shinyı with a brown sheen near the elytral base.
<emphasis id="7D9DBB0D2A21BB06FB9DE054E0112961" bold="true" box="[1087,1149,1322,1346]" pageId="67" pageNumber="2297">Head</emphasis>
: fronto-clypeal and genal surface smooth. Inter-ocular surface with fine punctures. Gena borders slightly curved inwards. Cephalic carina longer than wideı emarginate and producing two tubercles at the apex.
<emphasis id="7D9DBB0D2A21BB06FDFAE0ECE6BC2989" bold="true" box="[600,720,1426,1450]" pageId="67" pageNumber="2297">Pronotum</emphasis>
: lateral borders slightly irregular near the median lateral pronotal pit
<emphasis id="7D9DBB0D2A21BB06FDB1E0CAE64C29EF" box="[531,544,1460,1484]" italics="true" pageId="67" pageNumber="2297">–</emphasis>
rest of it smooth. Pronotal disc shinyı having a few spaced fine punctures and delimited anteriorly by a pair of acute tubercles. Anterior portionı including excavation and anterior angles with very few and inconspicuous sparse punctures. Posterior margin lacking punctuation.
<emphasis id="7D9DBB0D2A21BB06FD70E363E71E2A16" bold="true" box="[722,882,1565,1589]" pageId="67" pageNumber="2297">Metasternum</emphasis>
: sides and anterior portion of anterior lobe covered by dense red setaeı setigerous punctures equally spacedı where present.
<emphasis id="7D9DBB0D2A21BB06FEEEE31CE5FE2A59" bold="true" box="[332,402,1634,1658]" pageId="67" pageNumber="2297">Elytra</emphasis>
: striae weakly impressed with weak punctures. Punctures separated by twice their diameter. Interstriae flatı lacking microsculpture.
<emphasis id="7D9DBB0D2A21BB06FC36E3FBE0652ABE" bold="true" box="[916,1033,1669,1693]" pageId="67" pageNumber="2297">Abdomen</emphasis>
: glabrousı punctuation of the anterior margin of the ventrites interrupted medially.
<emphasis id="7D9DBB0D2A21BB06FBAFE3D9E0EF2A9C" bold="true" box="[1037,1155,1703,1727]" pageId="67" pageNumber="2297">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A26BB01FF05E5F0E57D2C85" box="[167,273,142,166]" captionStart="Figure 21" captionStartId="36.[160,225,1600,1623]" captionTargetBox="[163,1152,142,1569]" captionTargetId="figure@36.[163,1153,142,1569]" captionTargetPageId="36" captionText="Figure 21. D. fonsecae: (a) male dorsal habitus; (b) Holotype dorsal habitus; (c) holotype labels; (d) female dorsal view; (e) female ventral view; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665557" httpUri="https://zenodo.org/record/3665557/files/figure.png" pageId="68" pageNumber="2298">Figure 21</figureCitation>
(f-h)): parameraı apexes dorsally roundedı convergent; longitudinal excavation present taking more than 2/3 of the paramera length; basal excavation absent. Laterallyı abrupt declivity towards the apex. Ventrallyı subgenital plate rounded at the base and strongly emarginated at apex.
</paragraph>
<paragraph id="4F56671F2A26BB01FF02E445E5C42E07" blockId="68.[160,1157,315,548]" pageId="68" pageNumber="2298">
<emphasis id="7D9DBB0D2A26BB01FF02E445E5A82D77" bold="true" box="[160,452,315,340]" italics="true" pageId="68" pageNumber="2298">Morphological variation.</emphasis>
males BL: 21
<emphasis id="7D9DBB0D2A26BB01FDFDE445E6002D70" box="[607,620,315,339]" italics="true" pageId="68" pageNumber="2298">–</emphasis>
22 mmı PW:
<quantity id="8811CAFA2A26BB01FD55E442E72B2D77" box="[759,839,316,340]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="68" pageNumber="2298" unit="mm" value="12.0">12 mm</quantity>
. Females BL 18
<emphasis id="7D9DBB0D2A26BB01FC53E445E7922D70" box="[1009,1022,315,339]" italics="true" pageId="68" pageNumber="2298">–</emphasis>
20 mmı PW:
<quantity id="8811CAFA2A26BB01FF02E420E5772D55" box="[160,283,350,374]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="68" pageNumber="2298" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">
10
<emphasis id="7D9DBB0D2A26BB01FF1EE420E4A52D55" box="[188,201,350,374]" italics="true" pageId="68" pageNumber="2298">–</emphasis>
12 mm
</quantity>
. Females distinguishable from males as follows: Head: fronto-clypeal surface transversely wrinkled. Cephalic carina very lowı producing two small tubercles. Pronotum: punctuation coarser than on males but following same distribution pattern. Elytra: striae deeply impressedı punctures separated by twice their diameter. Abdomen: sixth ventrite bearing a pair of triangular projections reaching the apical margin of the pygidium (
<figureCitation id="D7D27B9A2A26BB01FEBEE772E5172E07" box="[284,379,524,548]" captionStart="Figure 7" captionStartId="20.[160,225,1522,1545]" captionTargetBox="[166,1148,142,1488]" captionTargetId="figure@20.[163,1153,142,1490]" captionTargetPageId="20" captionText="Figure 7. (a) Mandible of D. fissus (Harold), inc – incisory lobe (membranous), mo – molar lobe. (b) Maxilla and maxillary palp of same species, mxp – maxillary palp, ga – galea, las – lacinia. (c) Epipharynx of same species. (d) Labrum and labial palpomere of same species. (e) Ventral clypeal process of D. fissceps indicated by white arrow.(f) Antena of same species.esc – escapo, lam – lamella. (g) Macropterous hind wig of D. fissiceps. (h) Brachypterous hindwing of D. ingens." figureDoi="http://doi.org/10.5281/zenodo.3665533" httpUri="https://zenodo.org/record/3665533/files/figure.png" pageId="68" pageNumber="2298">Figure 7</figureCitation>
(b)).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A26BB01FF02E72EE5722EAF" pageId="68" pageNumber="2298" type="distribution">
<paragraph id="4F56671F2A26BB01FF02E72EE5722EAF" blockId="68.[160,1155,592,652]" pageId="68" pageNumber="2298">
<emphasis id="7D9DBB0D2A26BB01FF02E72EE55F2E4A" bold="true" box="[160,307,592,617]" italics="true" pageId="68" pageNumber="2298">Distribution.</emphasis>
Macası West
<collectingCountry id="37FE278F2A26BB01FE77E72FE65F2E4A" box="[469,563,593,617]" name="Ecuador" pageId="68" pageNumber="2298">Equador</collectingCountry>
(type-locality)ı
<collectingCountry id="37FE278F2A26BB01FD40E72FE7792E4A" box="[738,789,593,617]" name="Peru" pageId="68" pageNumber="2298">Peru</collectingCountry>
(
<collectingRegion id="8D2DA9FD2A26BB01FC81E72FE70E2E4A" box="[803,866,593,617]" country="Peru" name="Junin" pageId="68" pageNumber="2298">Junín</collectingRegion>
) and
<collectingCountry id="37FE278F2A26BB01FC01E72FE0632E4A" box="[931,1039,593,617]" name="Colombia" pageId="68" pageNumber="2298">Colombia</collectingCountry>
(
<collectingRegion id="8D2DA9FD2A26BB01FBBFE72FE0122E4A" box="[1053,1150,593,617]" country="Colombia" name="Caqueta" pageId="68" pageNumber="2298">Caquetá</collectingRegion>
) (
<figureCitation id="D7D27B9A2A26BB01FF05E70AE57D2EAF" box="[167,273,628,652]" captionStart="Figure 17" captionStartId="32.[160,225,1651,1674]" captionTargetBox="[164,1153,143,1619]" captionTargetId="figure@32.[163,1153,142,1619]" captionTargetPageId="32" captionText="Figure 17. D. mundus: (a) male dorsal habitus; (b) female dorsal habitus; (c) female ventral habitus; (d) paramera dorsal view; (e) paramera ventral view; (f) paramera lateral view; (g) type labels." figureDoi="http://doi.org/10.5281/zenodo.3665549" httpUri="https://zenodo.org/record/3665549/files/figure.png" pageId="68" pageNumber="2298">Figure 17</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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