203 lines
20 KiB
XML
203 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.953.49557" ID-GBIF-Dataset="735ba058-8e23-4d05-b35b-b677731b20c9" ID-PMC="PMC7398948" ID-Pensoft-Pub="1313-2970-953-105" ID-Pensoft-UUID="9D3BBD894B7752108D2406E4A6D088DC" ID-PubMed="32821198" ID-ZooBank="0AA9A04BAA0A4FE3896D07BBC005373D" ModsDocID="1313-2970-953-105" checkinTime="1595913798767" checkinUser="pensoft" docAuthor="Qian, Xingyang & Zhang, Xiao" docDate="2020" docId="9868E2F5778E5DFE8F1C650100A87C67" docLanguage="en" docName="ZooKeys 953: 105-118" docOrigin="ZooKeys 953" docSource="http://dx.doi.org/10.3897/zookeys.953.49557" docTitle="Geranomyia jiulongensis Qian & Zhang 2020, sp. nov." docType="treatment" docUuid="2E46B202-AA91-4C18-9421-6355DE078719" docUuidSource="ZooBank" docVersion="4" id="9D3BBD894B7752108D2406E4A6D088DC" lastPageNumber="105" masterDocId="9D3BBD894B7752108D2406E4A6D088DC" masterDocTitle="Two new Geranomyia Haliday (Diptera, Limoniidae) crane flies from Mount Jiulong in China, with an updated key to Chinese species" masterLastPageNumber="118" masterPageNumber="105" pageNumber="105" updateTime="1668169160721" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Two new Geranomyia Haliday (Diptera, Limoniidae) crane flies from Mount Jiulong in China, with an updated key to Chinese species</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Qian, Xingyang</mods:namePart>
|
|
<mods:affiliation>Key Lab of Integrated Crop Pest Management of Shandong Province, College of Plant Health and Medicine, Qingdao Agricultural University, Qingdao 266109, China</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Zhang, Xiao</mods:namePart>
|
|
<mods:affiliation>Key Lab of Integrated Crop Pest Management of Shandong Province, College of Plant Health and Medicine, Qingdao Agricultural University, Qingdao 266109, China</mods:affiliation>
|
|
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8054-0968</mods:nameIdentifier>
|
|
<mods:nameIdentifier type="email">779696635@qq.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2020</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>953</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>105</mods:start>
|
|
<mods:end>118</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.953.49557</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.953.49557</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-953-105</mods:identifier>
|
|
<mods:identifier type="ZooBank">0AA9A04BAA0A4FE3896D07BBC005373D</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">9D3BBD894B7752108D2406E4A6D088DC</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="166016769" LSID="urn:lsid:zoobank.org:act:2E46B202-AA91-4C18-9421-6355DE078719" httpUri="http://treatment.plazi.org/id/9868E2F5778E5DFE8F1C650100A87C67" lastPageNumber="105" pageId="0" pageNumber="105">
|
|
<subSubSection pageId="0" pageNumber="105" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<taxonomicName LSID="9868E2F5-778E-5DFE-8F1C-650100A87C67" authority="Qian & Zhang, 2020" authorityName="Qian & Zhang" authorityYear="2020" class="Insecta" family="Limoniidae" genus="Geranomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geranomyia jiulongensis" order="Diptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="jiulongensis" status="sp. nov.">Geranomyia jiulongensis</taxonomicName>
|
|
<taxonomicNameLabel pageId="0" pageNumber="105">sp. nov.</taxonomicNameLabel>
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">Figures 1</figureCitation>
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geranomyia jiulongensis sp. nov. a male hypopygium, dorsal view b male hypopygium, ventral view. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.953.49557.figure2" httpUri="https://binary.pensoft.net/fig/436439" pageId="0" pageNumber="105">, 2</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="105">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">Prescutum yellow with three broad, dark-brown longitudinal stripes. Pleuron of thorax yellow, with a broad, dark-brown stripe. Wing with seven large spots on costal region; Sc1 ending near fork of Rs, basal section of CuA1 nearly its length before fork of M. Lobe of gonostylus large with an arched rostral prolongation armed with two basal spines arising from two tubercles.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="description">
|
|
<paragraph pageId="0" pageNumber="105">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">Male.</emphasis>
|
|
Body length 5.0-5.3 mm, wing length 5.3-5.5 mm, mouthparts length 2.2-2.3 mm.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
Head (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">1b</figureCitation>
|
|
). Black. Setae on head black. Antenna length 1.2-1.3 mm, brownish black. Scape cylindrical. Pedicel nearly globose. Flagellomeres oval, terminal flagellomere with tip knob-like. Mouthparts brownish black with black setae.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105" start="Figure 1" startId="F1">
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">Figure 1.</emphasis>
|
|
<taxonomicName authorityName="Qian & Zhang" authorityYear="2020" class="Insecta" family="Limoniidae" genus="Geranomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geranomyia jiulongensis" order="Diptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="jiulongensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="105">Geranomyia jiulongensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="105">a</emphasis>
|
|
Male habitus, lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="105">b</emphasis>
|
|
head, lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="105">c</emphasis>
|
|
thorax, dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="105">d</emphasis>
|
|
wing
|
|
<emphasis bold="true" pageId="0" pageNumber="105">e</emphasis>
|
|
ovipositor, lateral view. Scale bars: 1.0 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="105">a</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="105">b-d</emphasis>
|
|
); 0.2 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="105">e</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
Thorax (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">1c</figureCitation>
|
|
). Pronotum yellow with a broad dark brown median stripe. Prescutum yellow, with three broad, dark-brown longitudinal stripes, each lateral stripe about 1/2 length of median stripe. Scutum pale yellow, with a dark-brown longitudinal stripe at middle area, each lobe with a large, dark-brown spot. Scutellum yellow, with two sides and anterior region dark brown, posterior region with a dark-brown spot. Mediotergite brownish black. Pleuron of thorax (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">1a</figureCitation>
|
|
) yellow, with a broad, dark-brown stripe extending from cervical region to mediotergite. Setae on thorax brownish black. Coxae pale yellow; trochanters pale yellow; femora brownish yellow, with fore femur paler; tibiae brownish yellow; tarsi brownish yellow, with terminal three segments darker. Setae on legs brownish black. Wing (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">1d</figureCitation>
|
|
) tinged pale brownish with a brownish-black pattern: seven large spots on costal region; seams along cord, m-m and basal section of M3; a spot at fork of Rs; a very light spot at sub-tip of A2. Veins brownish yellow, darker in clouded areas. Venation: Sc long, Sc1 ending near fork of Rs, Sc2 at its tip; basal section of CuA1 nearly its length before fork of M. Halter length 1.1-1.2 mm, pale yellow with base of stem dark brown.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
Abdomen (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">1a</figureCitation>
|
|
). Tergites brownish yellow with caudal halves dark brown. Sternites pale yellow. Setae on tergites brown and on sternites white.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
Hypopygium (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geranomyia jiulongensis sp. nov. a male hypopygium, dorsal view b male hypopygium, ventral view. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.953.49557.figure2" httpUri="https://binary.pensoft.net/fig/436439" pageId="0" pageNumber="105">2</figureCitation>
|
|
). Posterior margin of ninth tergite slightly emarginate. Gonocoxite slender with an elongate and blunt-apexed ventromesal lobe. Clasper of gonostylus arched at 2/3 of length, tip acute. Lobe of gonostylus large with an arched rostral prolongation armed with two basal spines arising from two tubercles. Paramere stout, wide at base, distal part trianglar. Aedeagus long, with two apical lobes.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.953.49557.figure2" httpUri="https://binary.pensoft.net/fig/436439" pageId="0" pageNumber="105" start="Figure 2" startId="F2">
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">Figure 2.</emphasis>
|
|
<taxonomicName authorityName="Qian & Zhang" authorityYear="2020" class="Insecta" family="Limoniidae" genus="Geranomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geranomyia jiulongensis" order="Diptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="jiulongensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="105">Geranomyia jiulongensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="105">a</emphasis>
|
|
male hypopygium, dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="105">b</emphasis>
|
|
male hypopygium, ventral view. Scale bars: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">Female.</emphasis>
|
|
Body length 5.5-6.4 mm, wing length 5.0-5.8 mm, mouthparts length 2.0-2.5 mm. Similar to male, but tenth tergite (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">1e</figureCitation>
|
|
) brown. Cercus brownish yellow with basal 1/2 brown. Hypogynial valve brownish yellow with tip slightly darker, tip near 2/3 way along cercus.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="type material">
|
|
<paragraph pageId="0" pageNumber="105">Type material.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="105">Holotype</emphasis>
|
|
: male (QAU), China: Zhejiang, Suichang, Mount Jiulong, Luohanyuan (
|
|
<geoCoordinate degrees="28" direction="north" minutes="23" orientation="latitude" precision="15" seconds="24" value="28.39">28°23'24"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="118" direction="east" minutes="51" orientation="longitude" precision="15" seconds="00" value="118.85">118°51'00"E</geoCoordinate>
|
|
, 517 m), 2019.VII.26, Xingyang Qian.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="105">Paratypes</emphasis>
|
|
: 10 males 5 females (QAU), same data as holotype. 1 female (QAU), China: Zhejiang, Suichang, Mount Jiulong, Longkoucun (
|
|
<geoCoordinate degrees="28" direction="north" minutes="18" orientation="latitude" precision="15" seconds="11" value="28.303055">28°18'11"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="118" direction="east" minutes="56" orientation="longitude" precision="15" seconds="42" value="118.945">118°56'42"E</geoCoordinate>
|
|
, 305 m), 2019.VII.24, Xingyang Qian. 1 male 5 females (QAU), China: Zhejiang, Suichang, Mt. Jiulong, Xikengli (
|
|
<geoCoordinate degrees="28" direction="north" minutes="20" orientation="latitude" precision="15" seconds="10" value="28.336111">28°20'10"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="118" direction="east" minutes="55" orientation="longitude" precision="15" seconds="00" value="118.916664">118°55'00"E</geoCoordinate>
|
|
, 732 m), 2019.VII.25, Xingyang Qian. 1 male (QAU), China: Zhejiang, Suichang, Mount Jiulong, Yanping (
|
|
<geoCoordinate degrees="28" direction="north" minutes="22" orientation="latitude" precision="15" seconds="23" value="28.373056">28°22'23"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="118" direction="east" minutes="53" orientation="longitude" precision="15" seconds="48" value="118.89667">118°53'48"E</geoCoordinate>
|
|
, 667 m), 2019.VII.26, Xingyang Qian. 1 female (QAU), China: Zhejiang, Suichang, Mount Jiulong, Zuobieyuan (
|
|
<geoCoordinate degrees="28" direction="north" minutes="17" orientation="latitude" precision="15" seconds="10" value="28.28611">28°17'10"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="118" direction="east" minutes="46" orientation="longitude" precision="15" seconds="42" value="118.778336">118°46'42"E</geoCoordinate>
|
|
, 640 m), 2019.VII.28, Xingyang Qian.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="distribution">
|
|
<paragraph pageId="0" pageNumber="105">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">China (Zhejiang).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="etymology">
|
|
<paragraph pageId="0" pageNumber="105">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">The species is named after the type locality, Mount Jiulong.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="remarks">
|
|
<paragraph pageId="0" pageNumber="105">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
This species is very similar to
|
|
<taxonomicName lsidName="G. radialis" pageId="0" pageNumber="105" rank="species" species="radialis">
|
|
<emphasis italics="true" pageId="0" pageNumber="105">G. radialis</emphasis>
|
|
</taxonomicName>
|
|
but can be distinguished by the prescutum of the thorax having three broad, dark-brown stripes (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">1c</figureCitation>
|
|
) and the lobe of the gonostylus being long and more than twice the length of the gonostylus clasper (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geranomyia jiulongensis sp. nov. a male hypopygium, dorsal view b male hypopygium, ventral view. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.953.49557.figure2" httpUri="https://binary.pensoft.net/fig/436439" pageId="0" pageNumber="105">2a</figureCitation>
|
|
). In
|
|
<taxonomicName lsidName="G. radialis" pageId="0" pageNumber="105" rank="species" species="radialis">
|
|
<emphasis italics="true" pageId="0" pageNumber="105">G. radialis</emphasis>
|
|
</taxonomicName>
|
|
, the prescutum has a narrow brown median stripe, and the lobe of the gonostylus is short and slightly exceeds the gonostylus clasper (Alexander 1930). This new species is also somewhat similar to
|
|
<taxonomicName lsidName="G. immobilis" pageId="0" pageNumber="105" rank="species" species="immobilis">
|
|
<emphasis italics="true" pageId="0" pageNumber="105">G. immobilis</emphasis>
|
|
</taxonomicName>
|
|
(Alexander, 1932) from the Philippines in its pattern and wing venation but can be easily distinguished from it by the pleuron of the thorax being yellow with a broad dark brown stripe extending from the cervical region to the mediotergite (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geranomyia jiulongensis sp. nov. a Male habitus, lateral view b head, lateral view c thorax, dorsal view d wing e ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.5 mm (b-d); 0.2 mm (e)." figureDoi="10.3897/zookeys.953.49557.figure1" httpUri="https://binary.pensoft.net/fig/436438" pageId="0" pageNumber="105">1a</figureCitation>
|
|
) and the lobe of the gonostylus with two spines arising from two tubercles. In
|
|
<taxonomicName lsidName="G. immobilis" pageId="0" pageNumber="105" rank="species" species="immobilis">
|
|
<emphasis italics="true" pageId="0" pageNumber="105">G. immobilis</emphasis>
|
|
</taxonomicName>
|
|
, the pleuron of the thorax is chiefly dark brown, with the sternopleurite light yellow, and the lobe of the gonostylus has two spines arising from a common tubercle (
|
|
<bibRefCitation author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="105" pagination="597 - 638" refId="B11" refString="Alexander, CP, 1932. New or little-known Tipulidae from the Philippines (Diptera). XV. Philippine Journal of Science 48: 597 - 638" title="New or little-known Tipulidae from the Philippines (Diptera). XV." volume="48" year="1932">Alexander 1932</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |