152 lines
16 KiB
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152 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. & Tishechkin, Alexey K." docDate="2013" docId="FDA8DB0E18A2EAD5202C8D20AC7E6E1F" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus subviridis Caterino & Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="293" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="289" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Caterino, Michael S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>271</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>401</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040844" LSID="urn:lsid:zoobank.org:act:079F7050-05B2-430F-BB3F-293B4177AB41" httpUri="http://treatment.plazi.org/id/FDA8DB0E18A2EAD5202C8D20AC7E6E1F" lastPageId="292" lastPageNumber="293" pageId="288" pageNumber="289">
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<subSubSection pageId="288" pageNumber="289" type="nomenclature">
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<paragraph pageId="288" pageNumber="289">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:079F7050-05B2-430F-BB3F-293B4177AB41" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subviridis" order="Coleoptera" pageId="288" pageNumber="289" phylum="Arthropoda" rank="species" species="subviridis">Operclipygus subviridis</taxonomicName>
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<taxonomicNameLabel pageId="288" pageNumber="289">sp. n.</taxonomicNameLabel>
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Figs 77B, D, G, I78
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<normalizedToken originalValue="A–BMap">A-BMap</normalizedToken>
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28
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</paragraph>
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</subSubSection>
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<subSubSection pageId="288" pageNumber="289" type="type locality">
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<paragraph pageId="288" pageNumber="289">Type locality.</paragraph>
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<paragraph pageId="288" pageNumber="289">
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COSTA RICA: Heredia: La Selva Biological Station [
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.433333">10°26'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-84.01667">84°01'W</geoCoordinate>
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].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="289" pageNumber="290" type="type material">
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<paragraph pageId="289" pageNumber="290">
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<pageBreakToken pageId="289" pageNumber="290" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="289" pageNumber="290">
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Holotype male: "Costa Rica: Heredia, Est. Biol. La Selva. 10.26'[°]N 84.01'[°]W. F.I.T. 24 June 1998, C.Carlton & A.Tishechkin leg." /
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<normalizedToken originalValue="“LSAM0046225”">"LSAM0046225"</normalizedToken>
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(FMNH). Paratypes (71): 54: same data as type, except as noted: 2: 19.vi.1998 (LSAM), 4: 21.vi.1998 (LSAM, MSCC, AKTC), 1: 22.vi.1998 (LSAM), 4: 23.vi.1998 (LSAM), 1: 24.vi.1998 (LSAM), 4: 25.vi.1998 (LSAM, FMNH), 3: 26.vi.1998 (LSAM), 1: 27.vi.1998 (LSAM), 3: 28.vi.1998 (LSAM), 1: 29.vi.1998 (LSAM), 1: i.1992, FIT, W. Bell (SEMC), 1: 25.i.1992 (SEMC), 2: 3.ii.1992 (SEMC), 2: 9.ii.1992 (SEMC), 3: 17.ii.1992 (SEMC), 9: 19.ii.1992 (SEMC), 1: 24.ii.1992 (SEMC), 2: 28.ii.1992 (SEMC), 2: 6.iii.1992 (SEMC), 2: 11.iii.1992 (SEMC), 1: 17.iii.1992 (SEMC), 2: 21.iii.1992 (SEMC), 1: 27.iii.1992 (SEMC), 1: 8.vi.2012, OTS Beetle Course, DNA Extract MSC-2303 (LSAM), 1: 21-28.iii.1988, W.E. Steiner, J.M. Hill, J.M. Swearingen, J.M. Mitchell (USNM); 3: Parque Nac. Braulio Carrillo, Est. El Ceibo, 400-600m, xi.1989, R. Aguilar & M. Zumbado, (INBIO);
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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: 2: Area Cons. Tortuguero, Sector Cerro Cocori, Fca. de E. Rojas, 150m, iii.1992, E. Rojas, (INBIO), 1: iii.1993 (INBIO), 1: ix.1991 (INBIO), 1: v.1993 (INBIO), 3: vi.1991 (INBIO), 1: xii.1992 (INBIO), 1: 10-30.ix.1992 (INBIO). NICARAGUA: Rio San Juan: 3: El Castillo loc. Bartola, iii.2000, I.G. Trezzi (CHFP, MEL); 1: Refugio Bartola, 60km SE San Carlos,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="10.973333">10°58.40'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-84.33833">84°20.30'W</geoCoordinate>
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, 100m, 25-28.v.2002, FIT, R. Brooks, Z. Falin, S. Chatzimanolis (SEMC).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="291" lastPageNumber="292" pageId="289" pageNumber="290" type="other material">
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<paragraph pageId="289" pageNumber="290">Other material.</paragraph>
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<paragraph lastPageId="291" lastPageNumber="292" pageId="289" pageNumber="290">
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COSTA RICA: Guanacaste: 1: Rio San Lorenzo, Tierras Morenas, Z.P. Tenorio, 1050m, ii.1993, C. Rodriguez, (INBIO), 1: ix.1995, G. Rodriguez, (INBIO); 1: Parque Nac. Guanacaste, 9km S Santa Cecilia, Est. Pitilla, 700m,
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<pageBreakToken pageId="290" pageNumber="291" start="start">viii</pageBreakToken>
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.1994, C. Moraga, (INBIO), 1: x.1994, C. Moraga, (INBIO); Cartago: 1: Parque Nac. Barbilla, Turrialba, 2km despues del Rio Dantas, 400m, 17.xi.2000, 28.xi.2000, FIT, W. Arana (INBIO); 1: Parque Nac. Barbilla, Sendero El Felino, 1.5km SO de estqcion, 690m, 20.v.2001, 24.v.2001, W. Arana (INBIO); Alajuela, 3:
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<normalizedToken originalValue="Peñas">Penas</normalizedToken>
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Blancas, 800m, 19.v.1989, FIT, J. Ashe, R. Brooks, R. Leschen (SEMC);
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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: 1: Res Biol Hitoy Cerere, Est. Hitoy Cerere, Valle de la Estrella, 160m, 28.ix-11.x.1999, FIT, W. Arana (INBIO), 1: 100m,
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<normalizedToken originalValue="v–vi">v-vi</normalizedToken>
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.1992, malaise trap, (INBIO); 1: R.B. Hitoy Cerere, Send. a Tepezcuintle, 100-200m, 30.iv-2.v.2002, pitfall, W. Arana (INBIO); 1: Amubri, 70m, 1-22.x.1994, G. Gallardo, (INBIO); Puntarenas: 1: Res. Biol. Carara, Est. Quebrada Bonita, 50m, xi.1994, R. Guzman, (INBIO); 1: Est. Biol.
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<pageBreakToken pageId="291" pageNumber="292" start="start">Las</pageBreakToken>
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Cruces, Coto Brus,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.783334">8°47'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-82.95">82°57'W</geoCoordinate>
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, 1000m, 1-2.iv.2002, FIT, A. Cline & A. Tishechkin (LSAM); 1: Monteverde, 1240m, 21.v.1989, FIT, J. Ashe, R. Brooks, R. Leschen (CHSM). HONDURAS:
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<normalizedToken originalValue="Cortés">Cortes</normalizedToken>
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: 1: Parque Nac. Cusuco, El Cortecito, 361198, 1717050, 1250m, 4-7.vi.2006, FIT, mature broadleaf forest, J. Nunez-Mino (OUMNH). PANAMA:
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<normalizedToken originalValue="Panamá">Panama</normalizedToken>
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: 2: Nusagandi, Ina Igar trail, 18-21.v.1993, pitfall, primary forest, E.G. Riley (TAMU); 1: Barro Colorado Isl.,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.183333">9°11'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-79.85">79°51'W</geoCoordinate>
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, 40m, 25-30.vi.2000, FIT, S. Chatzimanolis (SEMC), 1: 14-18.vi.2000, FIT, S. Chatzimanolis (SEMC).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="292" lastPageNumber="293" pageId="291" pageNumber="292" type="description">
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<paragraph pageId="291" pageNumber="292">Diagnostic description.</paragraph>
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<paragraph lastPageId="292" lastPageNumber="293" pageId="291" pageNumber="292">
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Length: 2.06-2.56 mm, width: 1.81-2.31 mm; body rufobrunneus, generally with a very faint, dull greenish tinge, elongate oval, strongly convex; frons depressed at middle; frontal stria rounded at sides, broadly interrupted above antennal bases, with a short sinuate fragment at middle; supraorbital stria weak, detached from sides of frontal stria; labrum narrow, less than twice as wide as long, weakly emarginate at apex; left mandible untoothed, right with small, subacute basal tooth; pronotal disk weakly depressed basomedially, but lacking distinct prescutellar impression, with very fine, inconspicuous ground punctation and ~10 larger, shallow punctures at sides; marginal pronotal stria interrupted behind head; submarginal pronotal stria continuous across front and along anterior half of lateral margin, obsolete basally; elytron with two complete epipleural striae, outer subhumeral stria absent or represented as very short fragment in apical half, inner subhumeral stria absent, striae 1-2 complete, 3rd stria present in basal third, and as short apical fragment, striae 4-5 represented by short apical fragments only, sutural stria present in apical three-fourths; elytral disk with crenulate, frequently fragmented apical marginal stria; prosternal keel broad, truncate at base, carinal striae converging almost straight to narrow anterior arch; meso- and metaventrite together convex; mesoventrite short, wide, not projecting anteriorly, marginal stria broadly interrupted; mesometaventral stria broadly arched forward to near mesoventral margin; lateral metaventral stria short, not extending past middle of ventrite; 1st abdominal ventrite with single, abbreviated lateral stria, with small fovea behind inner corner of metacoxa; propygidium and pygidium with dense, fine, shallow ground punctation; propygidium with coarse basomedial punctures, sparser laterally and posteriorly; pygidium with few very small, interspersed coarse punctures; apical marginal stria absent. Male genitalia (Figs 77B, D, G, I): accessory sclerites absent; T8 short, sides subparallel in basal two-thirds, convergent to apex, apical emargination narrow, basal emargination deep, broad, nearly reaching basal membrane attachment line, ventrolateral apodemes separated beneath; S9 with sides subparallel to weakly convergent with narrowly rounded apices, ventral halves approximate at base, diverging weakly to apex; T9 subparallel in basal two-thirds, apices narrow; T10 with halves separate; S9 strongly narrowed at middle, with sides subparallel in basal fourth, base narrowly subtruncate, apex with very narrow median emargination, apical flanges interrupted; tegmen widest near base, very gradually narrowed to apex, with little dorsoventral curvature, medioventral process visible from above as a narrow
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<normalizedToken originalValue="‘V’">'V'</normalizedToken>
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-shaped sclerotization, mostly contained in ventral membrane,
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<pageBreakToken pageId="292" pageNumber="293" start="start">very</pageBreakToken>
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weakly projecting beneath; median lobe about half tegmen length, very thin; basal piece about one-third tegmen length.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="292" pageNumber="293" type="remarks">
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<paragraph pageId="292" pageNumber="293">Remarks.</paragraph>
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<paragraph pageId="292" pageNumber="293">This species is very similar to several in this group, having only two complete dorsal elytral striae, with the 3rd stria present basally and apically, and the 4th and 5th present only as apical fragments (Fig. 78A). Although faint, the greenish metallic tinge of most individuals of this species will separate it from all others in this group. This species also frequently has a short outer subhumeral stria in the apical half, a strong apical marginal elytral stria, median part of the frontal stria present, a small (not minute) fovea behind the inner corner of the metacoxa, and small punctures sparsely interspersed with the dense ground punctation of the pygidium (Fig. 78B).</paragraph>
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<paragraph pageId="292" pageNumber="293">We limit the type series to specimens from northeastern Costa Rica and adjacent areas in Nicaragua.</paragraph>
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<caption pageId="292" pageNumber="293">
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<paragraph pageId="292" pageNumber="293">
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Map 28. Records of the
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus impunctipennis" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="impunctipennis">Operclipygus impunctipennis</taxonomicName>
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group.
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</paragraph>
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</caption>
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<caption pageId="292" pageNumber="293">
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<paragraph pageId="292" pageNumber="293">
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Figure 78.
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus impunctipennis" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="impunctipennis">Operclipygus impunctipennis</taxonomicName>
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group. A Dorsal habitus of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subviridis" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="subviridis">Operclipygus subviridis</taxonomicName>
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B Pygidia of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subviridis" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="subviridis">Operclipygus subviridis</taxonomicName>
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C Dorsal habitus of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus vorax" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="vorax">Operclipygus vorax</taxonomicName>
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D Pygidia of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus vorax" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="vorax">Operclipygus vorax</taxonomicName>
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E Dorsal habitus of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus chamelensis" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="chamelensis">Operclipygus chamelensis</taxonomicName>
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F Pygidia of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus chamelensis" order="Coleoptera" pageId="292" pageNumber="293" phylum="Arthropoda" rank="species" species="chamelensis">Operclipygus chamelensis</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="292" pageNumber="293" type="etymology">
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<paragraph pageId="292" pageNumber="293">Etymology.</paragraph>
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<paragraph pageId="292" pageNumber="293">The name of this species refers to its faint metallic greenish tinge.</paragraph>
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</subSubSection>
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</treatment>
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</document> |