treatments-xml/data/F3/5D/BC/F35DBC37BC2B59A697C1975D50E9F452.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1104.83264" ID-Pensoft-Pub="1313-2970-1104-115" ID-Pensoft-UUID="269D5675E1335FAE864664066101ABE8" ID-ZooBank="400FB40356244BFA8C17F12208C3B564" ModsDocID="1313-2970-1104-115" checkinTime="1654946543406" checkinUser="pensoft" docAuthor="Zhan, Yi, Sivayyapram, Varat, Liu, Fengxiang, Li, Daiqin &amp; Xu, Xin" docDate="2022" docId="F35DBC37BC2B59A697C1975D50E9F452" docLanguage="en" docName="ZooKeys 1104: 115-128" docOrigin="ZooKeys 1104" docPubDate="2022-06-10" docSource="http://dx.doi.org/10.3897/zookeys.1104.83264" docTitle="Liphistius hatyai Zhan &amp; Xu 2022, sp. nov." docType="treatment" docUuid="0C8153A0-51E6-42A0-A009-BEDA4DB4D77E" docUuidSource="ZooBank" docVersion="1" id="269D5675E1335FAE864664066101ABE8" lastPageNumber="115" masterDocId="269D5675E1335FAE864664066101ABE8" masterDocTitle="Three new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae) from Thailand" masterLastPageNumber="128" masterPageNumber="115" pageNumber="115" updateTime="1654946543406" updateUser="pensoft">
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<mods:title>Three new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae) from Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhan, Yi</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sivayyapram, Varat</mods:namePart>
<mods:affiliation>Center of Excellence in Entomology and Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Fengxiang</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocatalysis and Enzyme Engineering, School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Li, Daiqin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8269-7734</mods:nameIdentifier>
<mods:affiliation>Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
<mods:nameIdentifier type="email">dbslidq@nus.edu.sg</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Xu, Xin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5632-6622</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China</mods:affiliation>
<mods:nameIdentifier type="email">xuxin_09@163.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-06-10</mods:number>
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<mods:number>1104</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1104.83264</mods:url>
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<mods:classification>journal article</mods:classification>
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<treatment LSID="urn:lsid:zoobank.org:act:0C8153A0-51E6-42A0-A009-BEDA4DB4D77E" httpUri="http://treatment.plazi.org/id/F35DBC37BC2B59A697C1975D50E9F452" lastPageNumber="115" pageId="0" pageNumber="115">
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<paragraph pageId="0" pageNumber="115">
<taxonomicName LSID="http://zoobank.org/0C8153A0-51E6-42A0-A009-BEDA4DB4D77E" authority="Zhan &amp; Xu" authorityName="Zhan &amp; Xu" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius hatyai" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="hatyai" status="sp. nov.">Liphistius hatyai Zhan &amp; Xu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="115">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="description">
<paragraph pageId="0" pageNumber="115">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="type material">
<paragraph pageId="0" pageNumber="115">Type material.</paragraph>
<paragraph pageId="0" pageNumber="115">
<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Holotype</emphasis>
: Thailand • 1 ♂; Songkhla Province, Hat Yai District, Kho Hong, 
<geoCoordinate degrees="7.04" direction="north" orientation="latitude" precision="555" value="7.04">7.04°N</geoCoordinate>
, 
<geoCoordinate degrees="100.50" direction="east" orientation="longitude" precision="555" value="100.5">100.50°E</geoCoordinate>
; alt. 25 m; 13 November 2016; N. Warrit, V. Sivayyapram, N. Chatthanabun, P. Traiyasut leg.; XUX-2017-492. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Paratype</emphasis>
: Thailand • 1 ♀, same data as for the holotype; XUX-2017-493.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="diagnosis">
<paragraph pageId="0" pageNumber="115">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="115">
The male of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. hatyai" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="hatyai">
<emphasis italics="true" pageId="0" pageNumber="115">L. hatyai</emphasis>
</taxonomicName>
sp. nov. resembles males of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. albipes" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="albipes">
<emphasis italics="true" pageId="0" pageNumber="115">L. albipes</emphasis>
</taxonomicName>
Schwendinger, 1995 and 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. yangae" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="yangae">
<emphasis italics="true" pageId="0" pageNumber="115">L. yangae</emphasis>
</taxonomicName>
Platnick &amp; Sedgwick, 1984 in having a rounded, scale-like paraembolic plate (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A, E</figureCitation>
) but can be distinguished in having the tegulum with three transverse ridges in retrolateral view, while the latter two species have only one transverse ridge (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3C, G</figureCitation>
); from males of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. bicoloripes" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="bicoloripes">
<emphasis italics="true" pageId="0" pageNumber="115">L. bicoloripes</emphasis>
</taxonomicName>
Ono, 1988, 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. castaneus" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis italics="true" pageId="0" pageNumber="115">L. castaneus</emphasis>
</taxonomicName>
Schwendinger, 1995, and 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. niphanae" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="niphanae">
<emphasis italics="true" pageId="0" pageNumber="115">L. niphanae</emphasis>
</taxonomicName>
Ono, 1988 in having the cumulus slightly elevated (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C</figureCitation>
); from the male of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. inthanon" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="inthanon">
<emphasis italics="true" pageId="0" pageNumber="115">L. inthanon</emphasis>
</taxonomicName>
sp. nov. in having the tibial apophysis with fewer and longer setae (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C</figureCitation>
), the cumulus slightly elevated (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C</figureCitation>
), the subtegular apophysis absent (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A, F</figureCitation>
), and the embolic parts detached (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-G</figureCitation>
); from the male of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. keeratikiati" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="keeratikiati">
<emphasis italics="true" pageId="0" pageNumber="115">L. keeratikiati</emphasis>
</taxonomicName>
sp. nov. in having the sclerotised embolic part with three longitudinal ridges reaching the apex (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A</figureCitation>
) and the tegulum with three transverse ridges distally (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3C, G</figureCitation>
); from males of other 
<taxonomicName authorityName="Schiodte" authorityYear="1849" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Liphistius</emphasis>
</taxonomicName>
species in having the cumulus slightly elevated, the sclerotised embolic part with three longitudinal ridges in prolateral view, and the subtegular apophysis absent (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A, B, E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="115">
The female of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. hatyai" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="hatyai">
<emphasis italics="true" pageId="0" pageNumber="115">L. hatyai</emphasis>
</taxonomicName>
sp. nov. differs from females of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. albipes" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="albipes">
<emphasis italics="true" pageId="0" pageNumber="115">L. albipes</emphasis>
</taxonomicName>
and 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. castaneus" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis italics="true" pageId="0" pageNumber="115">L. castaneus</emphasis>
</taxonomicName>
in having a slightly narrower V-shaped posterior stalk (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3H, I</figureCitation>
); from females of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. bicoloripes" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="bicoloripes">
<emphasis italics="true" pageId="0" pageNumber="115">L. bicoloripes</emphasis>
</taxonomicName>
and 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. castaneus" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis italics="true" pageId="0" pageNumber="115">L. castaneus</emphasis>
</taxonomicName>
in having poreplate lacking anterolateral lobes (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3I</figureCitation>
); from the female of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. niphanae" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="niphanae">
<emphasis italics="true" pageId="0" pageNumber="115">L. niphanae</emphasis>
</taxonomicName>
in having the anterior margin of the poreplate straight (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3H, I</figureCitation>
); from females of 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. yangae" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="yangae">
<emphasis italics="true" pageId="0" pageNumber="115">L. yangae</emphasis>
</taxonomicName>
, 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. inthanon" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="inthanon">
<emphasis italics="true" pageId="0" pageNumber="115">L. inthanon</emphasis>
</taxonomicName>
sp. nov., and 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. keeratikiati" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="keeratikiati">
<emphasis italics="true" pageId="0" pageNumber="115">L. keeratikiati</emphasis>
</taxonomicName>
sp. nov. in having the poreplate almost squared and with a slightly V-shaped posterior stalk (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3H, I</figureCitation>
); from females of other 
<taxonomicName authorityName="Schiodte" authorityYear="1849" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Liphistius</emphasis>
</taxonomicName>
species in having a hair at the center of posterior stalk dorsally (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="description">
<paragraph pageId="0" pageNumber="115">Description.</paragraph>
<paragraph pageId="0" pageNumber="115">
<emphasis bold="true" pageId="0" pageNumber="115">Male.</emphasis>
Carapace reddish-brown, with a few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 2-6 larger than others, fifth largest; chelicerae robust, promargin of cheliceral groove with 12 denticles of variable size; labium yellow and fused with sternum; sternum yellow, with a few short setae on the anterior tip and many long setae on the elongated posterior tip; legs yellowish brown, with strong hairs and spines, without distinct annulations, with 3 tarsal claws; 8 spinnerets. Measurements: BL 17.6, CL 8.81, CW 8.42, OL 8.33, OW 7.31; eye sizes and interdistances: AME 0.06, ALE 0.77, PME 0.47, PLE 0.59, AME-AME 0.13, AME-ALE 0.21, PME-PME 0.08, PME-PLE 0.09, ALE-PLE 0.12, ALE-ALE 0.17, PLE-PLE 0.39, AME-PME 0.12. Labium 0.55 long and 0.70 wide. Sternum 3.74 long and 1.04 wide. Leg I 22.55 (7.11 + 2.57 + 4.90 + 5.64 + 2.33), leg II 22.85 (6.76 + 1.73 + 5.56 + 6.06 + 2.74), leg III 26.62 (6.89 + 3.55 + 5.63 + 7.51 + 3.04), leg IV 30.54 (9.07 + 3.83 + 7.50 + 10.25 + 2.63).
</paragraph>
<paragraph pageId="0" pageNumber="115">
<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Palp</emphasis>
: tibial apophysis with four setae of same length, stouter basally and slender distally (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C</figureCitation>
); paracymbium with many setae situated at tip (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C</figureCitation>
); several tapering spines on slightly elevated cumulus (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C</figureCitation>
); contrategulum with a triangular process, and an arched smooth edge with a sharp projection (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3B, C, G</figureCitation>
); tegulum with a serrated edge proximally and 3 transverse ridges distally (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3B, C, G</figureCitation>
); embolic parts detached (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C, G</figureCitation>
), paraembolic plate semicircular, scale-like (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A, E</figureCitation>
); embolus with 3 distinct longitudinal ridges reaching the tip prolaterally, with a few denticulations on apex, and with a flat membranous opening (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C, G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="115">
<emphasis bold="true" pageId="0" pageNumber="115">Female.</emphasis>
Carapace orange, with few short, scattered bristles; opisthosoma gray, with 12 brown tergites, close to each other, with gray patches, 2-6 larger than others, fifth largest; 8 eyes on dark ocular tubercle; chelicerae robust, reddish brown; promargin of cheliceral groove with 10 denticles of variable size; labium yellow, fused with sternum; sternum yellow with several setae; legs with strong setae and spines, without distinct annulations, with 3 tarsal claws; 8 spinnerets. Measurements: BL 23.8, CL 11.42, CW 10.33, OL 11.03, OW 10.06; eye sizes and interdistances: AME 0.14, ALE 1.00, PME 0.52, PLE 0.61, AME-AME 0.12, AME-ALE 0.25, PME-PME 0.10, PME-PLE 0.12, ALE-PLE 0.15, ALE-ALE 0.10, PLE-PLE 0.60, AME-PME 0.15. Labium 2.45 long and 1.26 wide. Sternum 5.15 long, 1.81 wide. Palp 20.19 (7.41 + 3.65 + 4.72 +4.41), leg I 25.15 (8.71 + 4.49 + 4.59 + 4.74 + 2.62), leg II 26.62 (9.09 + 4.52 + 4.81 + 5.24 + 2.96), leg III 28.10 (9.25 + 4.28 + 5.56 + 5.89 + 3.12), leg IV 37.53 (11.06 + 4.80 + 7.70 + 9.97 + 4.00).
</paragraph>
<paragraph pageId="0" pageNumber="115">
<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Genitalia</emphasis>
: poreplate almost squared; posterior stalk slightly V-shaped, with a hair situated in the center dorsally; central dorsal opening small, situated in center of poreplate; receptacular cluster spherical (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3H, I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="etymology">
<paragraph pageId="0" pageNumber="115">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="115">
The species epithet 
<normalizedToken originalValue="“hatyai”">&quot;hatyai&quot;</normalizedToken>
refers to the location of the type locality in Hat Yai District.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="distribution">
<paragraph pageId="0" pageNumber="115">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="115">
Southern Thailand (Songkhla Province) (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map of Thailand showing the localities of three new Liphistius species described." figureDoi="10.3897/zookeys.1104.83264.figure2" httpUri="https://binary.pensoft.net/fig/694475" pageId="0" pageNumber="115">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="remarks">
<paragraph pageId="0" pageNumber="115">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName authorityName="Zhan &amp; Xu" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius hatyai" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="hatyai">
<emphasis italics="true" pageId="0" pageNumber="115">Liphistius hatyai</emphasis>
</taxonomicName>
sp. nov. can be assigned to the 
<taxonomicName authorityName="Platnick &amp; Sedgwick" authorityYear="1984" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius trang" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="trang">
<emphasis italics="true" pageId="0" pageNumber="115">Liphistius trang</emphasis>
</taxonomicName>
-group according to the morphology of male palp and female genitalia. In males, the sclerotised part of embolus has two or three longitudinal ridges reaching the tip, the cumulus is slightly elevated, and the subtegulum is lacking an apophysis (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-G</figureCitation>
). In females, the poreplate has a small central dorsal opening and a receptacular cluster (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3H, I</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="115">
Specifically, 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. hatyai" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="hatyai">
<emphasis italics="true" pageId="0" pageNumber="115">L. hatyai</emphasis>
</taxonomicName>
sp. nov. belongs to the species complex D of the 
<taxonomicName authorityName="Platnick &amp; Sedgwick" authorityYear="1984" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius trang" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="trang">
<emphasis italics="true" pageId="0" pageNumber="115">Liphistius trang</emphasis>
</taxonomicName>
-group (sensu 
<bibRefCitation DOI="https://doi.org/10.1111/j.1463-6409.1998.tb00426.x" author="Schwendinger, PJ" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="115" pagination="17 - 30" refId="B8" refString="Schwendinger, PJ, 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30, DOI: https://doi.org/10.1111/j.1463-6409.1998.tb00426.x" title="Five new Liphistius species (Araneae, Mesothelae) from Thailand." url="https://doi.org/10.1111/j.1463-6409.1998.tb00426.x" volume="27" year="1998">Schwendinger 1998</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.35929/RSZ.0017" author="Schwendinger, PJ" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="115" pagination="321 - 353" refId="B10" refString="Schwendinger, PJ, Syuhadah, N, Lehmann-Graber, C, Price, L, Huber, S, Hashim, R, Bhassu, S, Monod, L, 2019. A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in Peninsular Malaysia; part 2. Revue Suisse de Zoologie 126 (2): 321 - 353, DOI: https://doi.org/10.35929/RSZ.0017" title="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in Peninsular Malaysia; part 2." url="https://doi.org/10.35929/RSZ.0017" volume="126" year="2019">Schwendinger et al. 2019</bibRefCitation>
) based on male palp and female genital morphology. In males, the slightly elevated cumulus possesses long, needle-like spines (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A-C</figureCitation>
), and the sclerotised part of embolus carries three longitudinal ridges reaching the tip (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3A</figureCitation>
). Female genitalia consist of a nearly squared poreplate, and a narrow, slightly V-shaped posterior stalk (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp and female genitalia of Liphistius hatyai Zhan &amp; Xu, sp. nov. A palp, prolateral view B palp, ventral view C palp, retrolateral view D-G palp, distal views H vulva, dorsal view I vulva, ventral view A-G XUX- 2017 - 492 (holotype) H, I XUX- 2017 - 493. Abbreviations used: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; RC = receptacular cluster; CT = contrategulum; Cu = cumulus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1104.83264.figure3" httpUri="https://binary.pensoft.net/fig/694476" pageId="0" pageNumber="115">3H, I</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="115">
Species complex D includes species distributed in southern Thailand, western Peninsular Malaysia, and Sumatra. This species complex in Thailand includes 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. albipes" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="albipes">
<emphasis italics="true" pageId="0" pageNumber="115">L. albipes</emphasis>
</taxonomicName>
, 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. bicoloripes" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="bicoloripes">
<emphasis italics="true" pageId="0" pageNumber="115">L. bicoloripes</emphasis>
</taxonomicName>
, 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. castaneus" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis italics="true" pageId="0" pageNumber="115">L. castaneus</emphasis>
</taxonomicName>
, 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. niphanae" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="niphanae">
<emphasis italics="true" pageId="0" pageNumber="115">L. niphanae</emphasis>
</taxonomicName>
, 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. trang" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="trang">
<emphasis italics="true" pageId="0" pageNumber="115">L. trang</emphasis>
</taxonomicName>
Platnick &amp; Sedgwick, 1984, and 
<taxonomicName class="Arachnida" genus="L." kingdom="Animalia" lsidName="L. yangae" order="Araneae" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="yangae">
<emphasis italics="true" pageId="0" pageNumber="115">L. yangae</emphasis>
</taxonomicName>
(for details see 
<bibRefCitation DOI="https://doi.org/10.1111/j.1463-6409.1998.tb00426.x" author="Schwendinger, PJ" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="115" pagination="17 - 30" refId="B8" refString="Schwendinger, PJ, 1998. Five new Liphistius species (Araneae, Mesothelae) from Thailand. Zoologica Scripta 27 (1): 17 - 30, DOI: https://doi.org/10.1111/j.1463-6409.1998.tb00426.x" title="Five new Liphistius species (Araneae, Mesothelae) from Thailand." url="https://doi.org/10.1111/j.1463-6409.1998.tb00426.x" volume="27" year="1998">Schwendinger 1998</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.35929/RSZ.0017" author="Schwendinger, PJ" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="115" pagination="321 - 353" refId="B10" refString="Schwendinger, PJ, Syuhadah, N, Lehmann-Graber, C, Price, L, Huber, S, Hashim, R, Bhassu, S, Monod, L, 2019. A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in Peninsular Malaysia; part 2. Revue Suisse de Zoologie 126 (2): 321 - 353, DOI: https://doi.org/10.35929/RSZ.0017" title="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in Peninsular Malaysia; part 2." url="https://doi.org/10.35929/RSZ.0017" volume="126" year="2019">Schwendinger et al. 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>