183 lines
24 KiB
XML
183 lines
24 KiB
XML
<document id="3680A10D5CA896D240775FE3799361FF" ID-CLB-Dataset="37321" ID-DOI="10.11646/zootaxa.4209.1.1" ID-GBIF-Dataset="2de5b7bf-3975-458a-b200-07ec69f4e013" ID-ISSN="1175-5326" ID-Zenodo-Dep="272646" ID-ZooBank="849BAB5C-464A-4B4A-A586-5742411EDC01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484407904318" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Castro, Peter" docDate="2016" docId="F16BFB33FFF5FFACFF6AFB01FD68FF4E" docLanguage="en" docName="zootaxa.4209.1.1.pdf.imf" docOrigin="Zootaxa 4209 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Megaesthesiinae Stevcic 2005" docType="treatment" docVersion="8" lastPageNumber="87" masterDocId="0D52834BFFA3FFFBFFFDFFD1FFCDFFB9" masterDocTitle="Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="87" updateTime="1698435566492" updateUser="ExternalLinkService">
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<mods:title id="CEAFE10A42F707EC45877F6AC66AC9BA">Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)</mods:title>
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<mods:namePart id="052D246D7827A5B3998EE459767B3924">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="3266EA432E88A2CCED063B5937A67828">Castro, Peter</mods:namePart>
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<mods:title id="8A0D8253FE8D2A0798F19ED09D8D01E1">Zootaxa</mods:title>
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<mods:date id="4FF82452BE74C9AE60A24A9F360ECE06">2016</mods:date>
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<mods:number id="20B80F6209B6858E2D81D5CF4E4A6879">4209</mods:number>
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<treatment id="F16BFB33FFF5FFACFF6AFB01FD68FF4E" ID-DOI="http://doi.org/10.5281/zenodo.5617213" ID-GBIF-Taxon="127419530" ID-Zenodo-Dep="5617213" LSID="urn:lsid:plazi:treatment:F16BFB33FFF5FFACFF6AFB01FD68FF4E" httpUri="http://treatment.plazi.org/id/F16BFB33FFF5FFACFF6AFB01FD68FF4E" lastPageId="87" lastPageNumber="87" pageId="86" pageNumber="87">
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<paragraph id="797D4A25FFF5FFADFF6AFB01FD40FB53" blockId="86.[151,653,1232,1258]" box="[151,653,1232,1258]" pageId="86" pageNumber="87">
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<heading id="2235FD49FFF5FFADFF6AFB01FD40FB53" bold="true" box="[151,653,1232,1258]" fontSize="11" level="1" pageId="86" pageNumber="87" reason="1">
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<emphasis id="4BB69637FFF5FFADFF6AFB01FD40FB53" bold="true" box="[151,653,1232,1258]" pageId="86" pageNumber="87">
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Subfamily
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<taxonomicName id="BEC231A6FFF5FFADFEE2FB01FD40FB53" ID-CoL="847GM" authority="Stevcic 2005" authorityName="Stevcic" authorityYear="2005" box="[287,653,1232,1258]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="86" pageNumber="87" phylum="Arthropoda" rank="subFamily" subFamily="Megaesthesiinae">
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Megaesthesiinae
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<bibRefCitation id="1D5337D4FFF5FFADFE0EFB01FD40FB53" author="Stevcic" box="[499,653,1232,1258]" pageId="86" pageNumber="87" refString="Stevcic, Z. (2005) The reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Natura Croatica (Fauna Croatica), 14, supplement 1, 1 - 159." type="journal article" year="2005">Števčić 2005</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="31D819AEFFF5FFADFF6AFAC7FE48FAF2" pageId="86" pageNumber="87" type="reference_group">
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<paragraph id="797D4A25FFF5FFADFF6AFAC7FE48FAF2" blockId="86.[151,1436,1302,1355]" pageId="86" pageNumber="87">
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<treatmentCitationGroup id="59D26D0BFFF5FFADFF6AFAC7FE48FAF2" pageId="86" pageNumber="87">
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Subfamily
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<taxonomicName id="BEC231A6FFF5FFADFEFBFAC7FDA3FA95" ID-CoL="847GM" authority="Stevcic 2005: 108" authorityName="Stevcic" authorityPageNumber="108" authorityYear="2005" box="[262,622,1302,1324]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="86" pageNumber="87" phylum="Arthropoda" rank="subFamily" subFamily="Megaesthesiinae">
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Megaesthesiinae
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<treatmentCitation id="F8636C34FFF5FFADFE48FAC7FDA3FA95" author="Stevcic" box="[437,622,1302,1324]" page="108" pageId="86" pageNumber="87" year="2005">
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<bibRefCitation id="1D5337D4FFF5FFADFE48FAC7FDF1FA95" author="Stevcic" box="[437,572,1302,1324]" pageId="86" pageNumber="87" refString="Stevcic, Z. (2005) The reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Natura Croatica (Fauna Croatica), 14, supplement 1, 1 - 159." type="journal article" year="2005">Števčić 2005</bibRefCitation>
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: 108
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</treatmentCitation>
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</taxonomicName>
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; 2013: 186.—Ng
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<emphasis id="4BB69637FFF5FFADFCDFFAC9FC98FA95" box="[802,853,1302,1325]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
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2008: 27, 76.—
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<treatmentCitation id="F8636C34FFF5FFADFC0AFAC7FB3DFA95" author="De" box="[1015,1264,1302,1325]" page="33" pageId="86" pageNumber="87" year="2009">
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<bibRefCitation id="1D5337D4FFF5FFADFC0AFAC7FB03FA95" author="De" box="[1015,1230,1302,1325]" pageId="86" pageNumber="87" refString="De Grave, S., Pentcheff, N. D., Ahyong, S. T., Chan, T. - Y., Crandall, K. A., Dworschak, P. C., Felder, D. L., Feldmann, R. M., Fransen, C. H. J. M., Goulding, L. Y. D., Lemaitre, R., Low, M. E. Y., Martin, J. W., Ng, P. K. L., Schweitzer, C. E., Tan, S. H., Tshudy, D. & Wetzer, R. (2009) A classification of living and fossil genera of decapod crustaceans. Raffles Bulletin of Zoology, suppl. 21, 1 - 109." type="journal article" year="2009">
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De Grave
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<emphasis id="4BB69637FFF5FFADFBA2FAC9FB5FFA95" box="[1119,1170,1302,1325]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
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2009
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</bibRefCitation>
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: 33
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||
</treatmentCitation>
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[in list].—
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<treatmentCitation id="F8636C34FFF5FFADFAA0FAC7FE4CFAF2" author="Davie" page="1082" pageId="86" pageNumber="87" year="2015">
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<bibRefCitation id="1D5337D4FFF5FFADFAA0FAC7FE8CFAF2" author="Davie" pageId="86" pageNumber="87" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 c) Systematics and classification of Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. & Von Vaupel Klein, C. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, complementary to the volumes translated from the French of the Traite de Zoologie, 9 (C) (I), Decapoda: Brachyura (Part 2), Brill, Leiden, pp. 1049 - 1130. http: // dx. doi. org / 10.1163 / 9789004190832 _ 021" type="journal article" year="2015" yearSuffix="c">
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Davie
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<emphasis id="4BB69637FFF5FFADFF3AFAE7FF37FAF2" box="[199,250,1333,1355]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
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2015c
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</bibRefCitation>
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||
: 1082
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="31D819AEFFF5FFADFF6AFAA6FE6AFA61" pageId="86" pageNumber="87" type="materials_examined">
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<paragraph id="797D4A25FFF5FFADFF6AFAA6FE11FA0D" blockId="86.[151,636,1399,1496]" pageId="86" pageNumber="87">
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<emphasis id="4BB69637FFF5FFADFF6AFAA6FED1FA29" bold="true" box="[151,284,1399,1424]" pageId="86" pageNumber="87">Type genus</emphasis>
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.
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<taxonomicName id="BEC231A6FFF5FFADFED6FAA6FEC2FA0C" authority="Rathbun, 1909 Other" authorityName="Rathbun, 1909 Other" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus">
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<emphasis id="4BB69637FFF5FFADFED6FAA6FE1DFA29" box="[299,464,1399,1424]" italics="true" pageId="86" pageNumber="87">Megaesthesius</emphasis>
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<bibRefCitation id="1D5337D4FFF5FFADFE2AFAA6FDB1FA29" author="Rathbun" box="[471,636,1399,1424]" pageId="86" pageNumber="87" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
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<emphasis id="4BB69637FFF5FFADFF3AFA4DFEC2FA0C" bold="true" box="[199,271,1436,1461]" pageId="86" pageNumber="87">Other</emphasis>
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</taxonomicName>
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||
<emphasis id="4BB69637FFF5FFADFEE8FA4DFE1EFA0C" bold="true" box="[277,467,1436,1461]" pageId="86" pageNumber="87">genera included</emphasis>
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:
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</paragraph>
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||
<paragraph id="797D4A25FFF5FFADFF3AFA6EFE6AFA61" blockId="86.[151,636,1399,1496]" box="[199,423,1471,1496]" pageId="86" pageNumber="87">
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<taxonomicName id="BEC231A6FFF5FFADFF3AFA6EFE9FFA61" box="[199,338,1471,1496]" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="4BB69637FFF5FFADFF3AFA6EFE9FFA61" box="[199,338,1471,1496]" italics="true" pageId="86" pageNumber="87">Alainthesius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="50852B4CFFF5FFADFEA5FA6EFE6AFA61" box="[344,423,1471,1496]" pageId="86" pageNumber="87" rank="genus">
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<emphasis id="4BB69637FFF5FFADFEA5FA6EFE6AFA61" bold="true" box="[344,423,1471,1496]" pageId="86" pageNumber="87">n. gen.</emphasis>
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||
</taxonomicNameLabel>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="31D819AEFFF5FFADFF6AF9D6FCF9F935" pageId="86" pageNumber="87" type="diagnosis">
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||
<paragraph id="797D4A25FFF5FFADFF6AF9D6FCF9F935" blockId="86.[151,1437,1543,2037]" pageId="86" pageNumber="87">
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<emphasis id="4BB69637FFF5FFADFF6AF9D6FEC7F999" bold="true" box="[151,266,1543,1568]" pageId="86" pageNumber="87">Diagnosis</emphasis>
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. Minute size, carapace subquadrate, high. Eye peduncle conspicuously long, immobile, cornea long, eye reduced, eye pigmentation reduced. Antennule greatly enlarged (particularly in males), cannot be folded. Buccal cavern conspicuously arched, sunken. Chelae nearly equal in both sexes. Outer (ventral) surface of penis calcified, resembling plate between “supplementary plate”, sternite 8.
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</paragraph>
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</subSubSection>
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<subSubSection id="31D819AEFFF5FFACFF3AF946FD68FF4E" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87" type="discussion">
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<paragraph id="797D4A25FFF5FFACFF3AF946FD68FF4E" blockId="86.[151,1437,1543,2037]" lastBlockId="87.[151,1436,151,248]" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
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<emphasis id="4BB69637FFF5FFADFF3AF946FEF9F909" bold="true" box="[199,308,1687,1712]" pageId="86" pageNumber="87">Remarks</emphasis>
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. The structure of the penis and the calcified plate covering it is of interest. Like all chasmocarcinids, megaesthesiines possess a distinctive “supplementary plate” between thoracic sternites 7 and 8. This “plate” is an invagination of sternite 8, and in chasmocarcinids the elongated penis is positioned just below this plate and the rest of sternite 8. In megaesthesiines there is a wider space between the “supplementary plate” and the rest of sternite 8 where the penis sits, and most of the penis (except for the distal part) is surrounded by a calcified tube, with the upper part forming an elongated plate (
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<figureCitation id="E1F956A0FFF5FFADFD51F89DFD36F8DD" box="[684,763,1868,1893]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C ‒ F, Megaesthesius yokoyai (after Takeda & Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G ‒ L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M ‒ R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S ‒ V, posterior margin of epistome; W ‒ Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="86" pageNumber="87">Fig. 19</figureCitation>
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W‒Y). This penis plate sits exactly in the space between the “supplementary plate” and the rest of sternite 8. In
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<taxonomicName id="BEC231A6FFF5FFADFD13F8BEFC34F831" box="[750,1017,1903,1928]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="sagedae">
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<emphasis id="4BB69637FFF5FFADFD13F8BEFC34F831" box="[750,1017,1903,1928]" italics="true" pageId="86" pageNumber="87">Megaesthesius sagedae</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="BEC231A6FFF5FFADFBF7F8BEFB36F831" box="[1034,1275,1903,1928]" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="signatus" status="sp. nov.">
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<emphasis id="4BB69637FFF5FFADFBF7F8BEFB36F831" box="[1034,1275,1903,1928]" italics="true" pageId="86" pageNumber="87">Alainthesius signatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="50852B4CFFF5FFADFAF8F8BEFA8AF831" box="[1285,1351,1903,1928]" pageId="86" pageNumber="87" rank="species">
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<emphasis id="4BB69637FFF5FFADFAF8F8BEFA8AF831" bold="true" box="[1285,1351,1903,1928]" pageId="86" pageNumber="87">n. sp.</emphasis>
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</taxonomicNameLabel>
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and
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<taxonomicName id="BEC231A6FFF5FFADFA79F8A1FECCF814" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="bertrandi" status="sp. nov.">
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<emphasis id="4BB69637FFF5FFADFA79F8A1FECCF814" italics="true" pageId="86" pageNumber="87">A. bertrandi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="50852B4CFFF5FFADFEF4F845FE84F814" box="[265,329,1940,1965]" pageId="86" pageNumber="87" rank="species">
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<emphasis id="4BB69637FFF5FFADFEF4F845FE84F814" bold="true" box="[265,329,1940,1965]" pageId="86" pageNumber="87">n. sp.</emphasis>
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</taxonomicNameLabel>
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, this penis plate does not move at all when the coxa of the fifth pereiopod (from which the penis emerges) is engaged and is effectively immobile, although the margins do not appear to be fused with the thoracic sternite (
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<figureCitation id="E1F956A0FFF5FFADFF06F80DFE81F84D" box="[251,332,2012,2037]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C ‒ F, Megaesthesius yokoyai (after Takeda & Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G ‒ L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M ‒ R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S ‒ V, posterior margin of epistome; W ‒ Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="86" pageNumber="87">Fig. 19</figureCitation>
|
||
W‒Y). In
|
||
<taxonomicName id="BEC231A6FFF5FFADFE3EF80DFD0AF84D" box="[451,711,2012,2037]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
|
||
<emphasis id="4BB69637FFF5FFADFE3EF80DFD0AF84D" box="[451,711,2012,2037]" italics="true" pageId="86" pageNumber="87">Megaesthesius migmus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="50852B4CFFF5FFADFD2DF80DFCDCF84C" box="[720,785,2012,2037]" pageId="86" pageNumber="87" rank="species">
|
||
<emphasis id="4BB69637FFF5FFADFD2DF80DFCDCF84C" bold="true" box="[720,785,2012,2037]" pageId="86" pageNumber="87">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, however, the penis plate is mobile (
|
||
<figureCitation id="E1F956A0FFF5FFADFB47F80DFAC0F84D" box="[1210,1293,2012,2037]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C ‒ F, Megaesthesius yokoyai (after Takeda & Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G ‒ L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M ‒ R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S ‒ V, posterior margin of epistome; W ‒ Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="86" pageNumber="87">
|
||
Fig.
|
||
<date id="0D7C6CE5FFF5FFADFB0DF80DFAC0F84D" box="[1264,1293,2012,2036]" pageId="86" pageNumber="87" value="1919-10">19</date>
|
||
</figureCitation>
|
||
X). This is a feature that will need further study to determine if it is affected by growth or if it varies. The lack of additional male specimens of
|
||
<taxonomicName id="BEC231A6FFF4FFACFECFFF6DFDF8FF6D" box="[306,565,188,213]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
|
||
<emphasis id="4BB69637FFF4FFACFECFFF6DFDF8FF6D" box="[306,565,188,213]" italics="true" pageId="87" pageNumber="88">Megaesthesius migmus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4BB69637FFF4FFACFDC1FF6DFD4EFF6C" bold="true" box="[572,643,188,213]" pageId="87" pageNumber="88">
|
||
<taxonomicNameLabel id="50852B4CFFF4FFACFDC1FF6DFDB0FF6C" box="[572,637,188,213]" pageId="87" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="BEC231A6FFF4FFACFD76FF6DFCB4FF6D" box="[651,889,188,213]" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="signatus" status="sp. nov.">
|
||
<emphasis id="4BB69637FFF4FFACFD76FF6DFCB4FF6D" box="[651,889,188,213]" italics="true" pageId="87" pageNumber="88">Alainthesius signatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="50852B4CFFF4FFACFC7CFF6DFC0DFF6C" box="[897,960,188,213]" pageId="87" pageNumber="88" rank="species">
|
||
<emphasis id="4BB69637FFF4FFACFC7CFF6DFC0DFF6C" bold="true" box="[897,960,188,213]" pageId="87" pageNumber="88">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="BEC231A6FFF4FFACFC02FF6DFB44FF6C" box="[1023,1161,188,213]" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="bertrandi" status="sp. nov.">
|
||
<emphasis id="4BB69637FFF4FFACFC02FF6DFB44FF6C" box="[1023,1161,188,213]" italics="true" pageId="87" pageNumber="88">A. bertrandi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="50852B4CFFF4FFACFB6DFF6DFB02FF6C" box="[1168,1231,188,213]" pageId="87" pageNumber="88" rank="species">
|
||
<emphasis id="4BB69637FFF4FFACFB6DFF6DFB02FF6C" bold="true" box="[1168,1231,188,213]" pageId="87" pageNumber="88">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(only one of each is known) poses limitations to study at present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |