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<document id="F0A124DA7EC1A22DF4E27D0545F1AD28" ID-DOI="10.11646/zootaxa.4747.2.1" ID-GBIF-Dataset="55ab330e-0468-42ed-aeb0-7850751cf989" ID-ISSN="1175-5326" ID-Zenodo-Dep="3694740" ID-ZooBank="6AA8F5DC-8D70-42B0-B016-6F9C4211C471" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583220371855" checkinUser="plazi" docAuthor="Dick, Matthew H., Ngai, Nguyen Danh &amp; Doan, Hung Dinh" docDate="2020" docId="CD3D2E24CC6BFFD0B191FBD4FCE118D8" docLanguage="en" docName="zootaxa.4747.2.1.pdf" docOrigin="Zootaxa 4747 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhynchozoon latiavicularium Dick &amp; Ngai &amp; Doan 2020, n. sp." docType="treatment" docUuid="D543ACD0-6CEA-46F5-8624-F9FD808E0A37" docUuidSource="ZooBank" docVersion="7" lastPageNumber="239" masterDocId="3104565CCC4FFFF6B106FFB1FFC61C7B" masterDocTitle="Taxonomy and diversity of coelobite bryozoans from drift coral cobbles on Co To Island, northern Vietnam" masterLastPageNumber="252" masterPageNumber="201" pageNumber="237" updateTime="1698724047848" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A3CEFC6383BA8A401DE9025A389C6725">Taxonomy and diversity of coelobite bryozoans from drift coral cobbles on Co To Island, northern Vietnam</mods:title>
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<mods:namePart id="5E8E0906F8B3FDA4FD5C3F69FA8366C7">Dick, Matthew H.</mods:namePart>
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<mods:namePart id="FDD09244A630828CFDE4D5165D06EDB7">Ngai, Nguyen Danh</mods:namePart>
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<mods:namePart id="C3AA7211A9721E3CDBB2CF707F51C770">Doan, Hung Dinh</mods:namePart>
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<mods:title id="60EF6284842B8EF891F19919C779A225">Zootaxa</mods:title>
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<treatment id="CD3D2E24CC6BFFD0B191FBD4FCE118D8" ID-DOI="http://doi.org/10.5281/zenodo.3703704" ID-GBIF-Taxon="162381130" ID-Zenodo-Dep="3703704" LSID="urn:lsid:zoobank.org:act:D543ACD0-6CEA-46F5-8624-F9FD808E0A37" httpUri="http://treatment.plazi.org/id/CD3D2E24CC6BFFD0B191FBD4FCE118D8" lastPageId="38" lastPageNumber="239" pageId="36" pageNumber="237">
<subSubSection id="0D8ECCB9CC6BFFD2B191FBD4FD8E1804" box="[151,584,1125,1152]" pageId="36" pageNumber="237" type="nomenclature">
<paragraph id="452B9F32CC6BFFD2B191FBD4FD8E1804" blockId="36.[151,975,1125,1223]" box="[151,584,1125,1152]" pageId="36" pageNumber="237">
<heading id="1E63285ECC6BFFD2B191FBD4FD8E1804" bold="true" box="[151,584,1125,1152]" fontSize="11" level="1" pageId="36" pageNumber="237" reason="1">
<emphasis id="77E04320CC6BFFD2B191FBD4FD8E1804" bold="true" box="[151,584,1125,1152]" pageId="36" pageNumber="237">
<taxonomicName id="8294E4B1CC6BFFD2B191FBD4FE3918FB" authority="Dick &amp; Ngai &amp; Doan, 2020" authorityName="Dick &amp; Ngai &amp; Doan" authorityYear="2020" box="[151,511,1125,1152]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="36" pageNumber="237" phylum="Bryozoa" rank="species" species="latiavicularium" status="sp. nov.">
<emphasis id="77E04320CC6BFFD2B191FBD4FE3918FB" bold="true" box="[151,511,1125,1152]" italics="true" pageId="36" pageNumber="237">Rhynchozoon latiavicularium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CD3FE5BCC6BFFD2B300FBD4FD8E1804" box="[518,584,1125,1151]" pageId="36" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC6BFFD2B191FB38FC0918BC" pageId="36" pageNumber="237" type="description">
<paragraph id="452B9F32CC6BFFD2B191FB38FF3F18D8" blockId="36.[151,975,1125,1223]" box="[151,249,1161,1187]" pageId="36" pageNumber="237">
(
<figureCitation id="DDAF83B7CC6BFFD2B1A6FB38FF3718D8" box="[160,241,1161,1187]" captionStart="FIGURE 17" captionStartId="37.[151,250,1687,1712]" captionTargetBox="[151,1436,345,1663]" captionTargetId="figure@37.[151,1436,345,1663]" captionTargetPageId="37" captionText="FIGURE 17. Rhynchozoon latiavicularium n. sp. AC. Paratype, VNMN-0265; DF. Holotype, VNMN-0264. A. Colony. B. Marginal autozooids in frontodistal view, showing orifices and suboral avicularia. C. Enlargement of orifices. D. Colony surface, showing numerous frontal avicularia; ooecia also abundant. E. Suboral and frontal avicularia; note that avicularia can have rounded (arrows) or more acute (white arrowheads) proximal end of rostrum; also note V-shaped crossbar elements (black arrowheads) of avicularia. F. Immersed ooecia. Scale bars: A, 1 mm; B, D, 500 µm; C, 100 µm; E, 125 µm; F, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3694774" httpUri="https://zenodo.org/record/3694774/files/figure.png" pageId="36" pageNumber="237">Fig. 17</figureCitation>
)
</paragraph>
<paragraph id="452B9F32CC6BFFD2B191FB1DFC0918BC" blockId="36.[151,975,1125,1223]" box="[151,975,1196,1223]" pageId="36" pageNumber="237">
<uri id="31059330CC6BFFD2B191FB1DFC0918BC" box="[151,975,1196,1223]" pageId="36" pageNumber="237">
urn:lsid:zoobank.org:act:
<uuid id="3132A5E7CC6BFFD2B0AEFB1DFC0918BC" box="[424,975,1196,1223]" pageId="36" pageNumber="237">D543ACD0-6CEA-46F5-8624-F9FD808E0A37</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC6BFFD2B191FB44FE5A1948" pageId="36" pageNumber="237" type="etymology">
<paragraph id="452B9F32CC6BFFD2B191FB44FE5A1948" blockId="36.[151,1437,1269,2015]" pageId="36" pageNumber="237">
<emphasis id="77E04320CC6BFFD2B191FB44FED81974" bold="true" box="[151,286,1269,1295]" pageId="36" pageNumber="237">Etymology.</emphasis>
The specific name combines the Latin word
<taxonomicName id="8294E4B1CC6BFFD2B220FB44FC941974" authorityName="Busk" authorityYear="1852" box="[806,850,1269,1295]" class="Gymnolaemata" family="Candidae" genus="Caberea" kingdom="Animalia" order="Cheilostomatida" pageId="36" pageNumber="237" phylum="Bryozoa" rank="species" species="lata">
<emphasis id="77E04320CC6BFFD2B220FB44FC941974" box="[806,850,1269,1295]" italics="true" pageId="36" pageNumber="237">lata</emphasis>
</taxonomicName>
(wide) with
<emphasis id="77E04320CC6BFFD2B2E1FB44FBA81974" box="[999,1134,1269,1295]" italics="true" pageId="36" pageNumber="237">avicularium</emphasis>
, referring to the relatively wide frontal avicularia.
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC6BFFD2B1C1FA8DFDA21900" pageId="36" pageNumber="237" type="materials_examined">
<paragraph id="452B9F32CC6BFFD2B1C1FA8DFDA21900" blockId="36.[151,1437,1269,2015]" pageId="36" pageNumber="237">
<emphasis id="77E04320CC6BFFD2B1C1FA8DFE76192C" bold="true" box="[199,432,1340,1367]" pageId="36" pageNumber="237">Material examined.</emphasis>
<materialsCitation id="F5FC956FCC6BFFD2B0B1FA8CFDA21900" ID-GBIF-Occurrence="2573311943" pageId="36" pageNumber="237" typeStatus="holotype">
<typeStatus id="9A2F2190CC6BFFD2B0B1FA8CFDE4192C" box="[439,546,1341,1367]" pageId="36" pageNumber="237" type="holotype">Holotype</typeStatus>
, VNMN-0264 (CT-14);
<typeStatus id="9A2F2190CC6BFFD2B237FA8CFC59192C" box="[817,927,1341,1367]" pageId="36" pageNumber="237" type="paratype">paratypes</typeStatus>
, VNMN-0265 (CT-4), VNMN-0266 (CT-25), VNMN-0267 (CT-34), all on SEM stubs.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC6BFFD2B1C1FA34FF731A70" pageId="36" pageNumber="237" type="description">
<paragraph id="452B9F32CC6BFFD2B1C1FA34FF731A70" blockId="36.[151,1437,1269,2015]" pageId="36" pageNumber="237">
<emphasis id="77E04320CC6BFFD2B1C1FA34FE7E19E4" bold="true" box="[199,440,1412,1439]" pageId="36" pageNumber="237">Measurements. AzL</emphasis>
, 0.450.63 (0.536 ± 0.062);
<emphasis id="77E04320CC6BFFD2B3F9FA35FCFA19E4" bold="true" box="[767,828,1412,1439]" pageId="36" pageNumber="237">AzW</emphasis>
, 0.300.54 (0.399 ± 0.056);
<emphasis id="77E04320CC6BFFD2B587FA35FB7019E4" bold="true" box="[1153,1206,1412,1439]" pageId="36" pageNumber="237">OrL</emphasis>
, 0.100.14 (0.121 ± 0.010);
<emphasis id="77E04320CC6BFFD2B1E9FA19FEEB19B8" bold="true" box="[239,301,1448,1475]" pageId="36" pageNumber="237">OrW</emphasis>
, 0.100.12 (0.106 ± 0.006) (n = 15, 2).
<emphasis id="77E04320CC6BFFD2B3F1FA19FCEB19B8" bold="true" box="[759,813,1448,1475]" pageId="36" pageNumber="237">OvL</emphasis>
, 0.140.21 (0.173 ± 0.023);
<emphasis id="77E04320CC6BFFD2B573FA19FB7319B8" bold="true" box="[1141,1205,1448,1475]" pageId="36" pageNumber="237">OvW</emphasis>
, 0.190.23 (0.208 ± 0.013) (n = 15, 1).
<emphasis id="77E04320CC6BFFD2B06EFA7CFDCB199C" bold="true" box="[360,525,1484,1511]" pageId="36" pageNumber="237">Frontal AvRL</emphasis>
, 0.140.18 (0.157 ± 0.014);
<emphasis id="77E04320CC6BFFD2B24AFA7DFC34199C" bold="true" box="[844,1010,1484,1511]" pageId="36" pageNumber="237">frontal AvRW</emphasis>
,
<geoCoordinate id="20A0F9F5CC6BFFD2B506FA7CFBF9199C" box="[1024,1087,1485,1511]" degrees="0.066" direction="north" orientation="latitude" pageId="36" pageNumber="237" precision="55" value="0.066">0.066</geoCoordinate>
<geoCoordinate id="20A0F9F5CC6BFFD2B538FA7CFB4A199C" box="[1086,1164,1485,1511]" degrees="0.102" direction="west" orientation="longitude" pageId="36" pageNumber="237" precision="55" value="-0.102">0.102</geoCoordinate>
(0.080 ± 0.008) (n = 30, 2).
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC6BFFD2B1C1F9A5FE831B7C" pageId="36" pageNumber="237" type="diagnosis">
<paragraph id="452B9F32CC6BFFD2B1C1F9A5FE831B7C" blockId="36.[151,1437,1269,2015]" pageId="36" pageNumber="237">
<emphasis id="77E04320CC6BFFD2B1C1F9A5FE841A54" bold="true" box="[199,322,1556,1583]" pageId="36" pageNumber="237">Diagnosis.</emphasis>
Suboral sinus U-shaped, moderately shallow, narrow, flanked by straight orificial margin and broad, semicircular condyles. Oral denticles indistinct. Oral spines lacking. Zooids with suboral avicularium and single frontal avicularium. Suboral avicularium relatively broad; uncinus scarcely evident. Frontal avicularia relatively short, broad; weakly diamond shaped or semicircular proximally; scarcely decurved, with triangular mandibular portion lacking setiform end; paired crossbar elements V-shaped. Ooecium with transversely oval endooecial panel in peristome; labellum present but not markedly broad. Cylindrical projections around secondary orifice in older parts of colony.
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC6BFFD0B1C1F8A1FCB21E44" lastPageId="38" lastPageNumber="239" pageId="36" pageNumber="237" type="description">
<paragraph id="452B9F32CC6BFFD2B1C1F8A1FE4E1B34" blockId="36.[151,1437,1269,2015]" pageId="36" pageNumber="237">
<emphasis id="77E04320CC6BFFD2B1C1F8A1FE9F1B50" bold="true" box="[199,345,1808,1835]" pageId="36" pageNumber="237">Description.</emphasis>
Colony encrusting, sheet-like, initially unilaminar, developing bilaminar zones due to frontal budding, but not nodular.
</paragraph>
<paragraph id="452B9F32CC6BFFD3B1C1F8E8FBC61D38" blockId="36.[151,1437,1269,2015]" lastBlockId="37.[151,1436,153,323]" lastPageId="37" lastPageNumber="238" pageId="36" pageNumber="237">Zooids irregularly hexagonal but boundaries distinct only at colony margin. Frontal wall at margin smooth, convex, imperforate, with 710 small, circular areolae along each lateral margin; with thickening frontal calcification, areolar openings become larger and fewer, five or six along each lateral margin. Primary orifice with anter D-shaped or nearly circular; median sinus variable, ranging from narrowly U-shaped to more broadly rounded-V-shaped, flanked on each side by small shelf, with rounded condyle filling angle between shelf and lateral orifical margin; condyles not directed distally. Oral spines lacking. With increased calcification, scattered low, rounded tubercles appearing on frontal wall, though surface texture remaining smooth; from two to seven short, blunt cylindrical processes forming around secondary orifice as outgrowths from the frontal walls of surrounding zooids; and deep, asymmetrical secondary orifical sinus forming to one side of midline or other.</paragraph>
<caption id="11EBCFBACC6AFFD3B191F926FA8B1B3B" ID-DOI="http://doi.org/10.5281/zenodo.3694774" ID-Zenodo-Dep="3694774" httpUri="https://zenodo.org/record/3694774/files/figure.png" pageId="37" pageNumber="238" startId="37.[151,250,1687,1712]" targetBox="[151,1436,345,1663]" targetPageId="37">
<paragraph id="452B9F32CC6AFFD3B191F926FA8B1B3B" blockId="37.[151,1437,1687,1856]" pageId="37" pageNumber="238">
<emphasis id="77E04320CC6AFFD3B191F926FEE51ACB" bold="true" box="[151,291,1687,1712]" pageId="37" pageNumber="238">FIGURE 17.</emphasis>
<taxonomicName id="8294E4B1CC6AFFD3B02BF929FD9D1ACB" authority="Dick &amp; Ngai &amp; Doan, 2020" authorityName="Dick &amp; Ngai &amp; Doan" authorityYear="2020" box="[301,603,1688,1712]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="37" pageNumber="238" phylum="Bryozoa" rank="species" species="latiavicularium" status="sp. nov.">
<emphasis id="77E04320CC6AFFD3B02BF929FD9D1ACB" box="[301,603,1688,1712]" italics="true" pageId="37" pageNumber="238">Rhynchozoon latiavicularium</emphasis>
</taxonomicName>
<emphasis id="77E04320CC6AFFD3B363F929FD671ACB" bold="true" box="[613,673,1688,1712]" pageId="37" pageNumber="238">
<taxonomicNameLabel id="6CD3FE5BCC6AFFD3B363F929FD671ACB" box="[613,673,1688,1712]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
AC. Paratype, VNMN-0265; DF. Holotype, VNMN-0264. A. Colony. B. Marginal autozooids in frontodistal view, showing orifices and suboral avicularia. C. Enlargement of orifices. D. Colony surface, showing numerous frontal avicularia; ooecia also abundant. E. Suboral and frontal avicularia; note that avicularia can have rounded (arrows) or more acute (white arrowheads) proximal end of rostrum; also note V-shaped crossbar elements (black arrowheads) of avicularia. F. Immersed ooecia. Scale bars: A, 1 mm; B, D, 500 µm; C, 100 µm; E, 125 µm; F, 250 µm.
</paragraph>
</caption>
<paragraph id="452B9F32CC6AFFD0B1C1F8DFFDFA1DA8" blockId="37.[151,1437,1902,2037]" lastBlockId="38.[151,1437,153,1187]" lastPageId="38" lastPageNumber="239" pageId="37" pageNumber="238">Zooids with suboral avicularium, usually 2025% longer than orifice width; poster broadly semicircular, with smooth cryptocyst. Rostrum directed laterally, with rostral plane also tilted slightly toward distal end of zooid; mandibular portion asymmetrical, forming right triangle, or symmetrical, forming short-equilateral triangle; opesial margin of mandibular portion irregular with one or two coarse, nodular denticles on each side. Hinge element on each side of rostrum narrowly V-shaped, apex pointing medially, with or without complete crossbar extending between opposing elements. Suboral avicularium occasionally larger, twice as long as orifice width, with rostral plane convex in frontal direction. Most zooids have single frontal avicularium located in proximal half of frontal shield, directed distolaterally, laterally, proximolaterally, or proximally; frontal avicularia similar in size and shape to suboral avicularia; ratio of average rostrum length to rostrum width = 1.9; proximal end of rostrum usually acute (rostrum diamond-shaped), but sometimes rounded as in suboral avicularia; mandibular portion usually symmetrical and long-triangular, sometimes asymmetrical and comprising right triangle. Opesial margin of mandibular portion irregular, with up to six coarse, rounded denticles on each side. Rostrum slightly decurved at end, indicating mandible is probably hooked distally.</paragraph>
<paragraph id="452B9F32CC69FFD0B1C1FE6CFCB21E44" blockId="38.[151,1437,153,1187]" pageId="38" pageNumber="239">Ooecium immersed with age; non-calcified ectooecium leaving circular or oval zone of exposed endooecium vertically oriented in sunken peristome, with broad labellum along proximal ooecial margin; secondary calcification atop ooecium typically giving rise to one or two coarse tubercles.</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC69FFD0B1C1FDF8FEA01804" pageId="38" pageNumber="239" type="discussion">
<paragraph id="452B9F32CC69FFD0B1C1FDF8FA971FD3" blockId="38.[151,1437,153,1187]" pageId="38" pageNumber="239">
<emphasis id="77E04320CC69FFD0B1C1FDF8FEFD1E18" bold="true" box="[199,315,585,611]" pageId="38" pageNumber="239">Remarks.</emphasis>
<taxonomicName id="8294E4B1CC69FFD0B044FDF8FD4B1E18" authority="Dick &amp; Ngai &amp; Doan, 2020" authorityName="Dick &amp; Ngai &amp; Doan" authorityYear="2020" box="[322,653,585,611]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="38" pageNumber="239" phylum="Bryozoa" rank="species" species="latiavicularium" status="sp. nov.">
<emphasis id="77E04320CC69FFD0B044FDF8FD4B1E18" box="[322,653,585,611]" italics="true" pageId="38" pageNumber="239">Rhynchozoon latiavicularium</emphasis>
</taxonomicName>
<emphasis id="77E04320CC69FFD0B395FDF8FD141E18" bold="true" box="[659,722,585,611]" pageId="38" pageNumber="239">
<taxonomicNameLabel id="6CD3FE5BCC69FFD0B395FDF8FD141E18" box="[659,722,585,611]" pageId="38" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is quite similar to
<taxonomicName id="8294E4B1CC69FFD0B2A4FDFBFBA91E18" authority="Dick &amp; Ngai &amp; Doan, 2020" authorityName="Dick &amp; Ngai &amp; Doan" authorityYear="2020" box="[930,1135,585,611]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="38" pageNumber="239" phylum="Bryozoa" rank="species" species="setiavicularium" status="sp. nov.">
<emphasis id="77E04320CC69FFD0B2A4FDFBFBA91E18" box="[930,1135,585,611]" italics="true" pageId="38" pageNumber="239">R. setiavicularium</emphasis>
</taxonomicName>
<emphasis id="77E04320CC69FFD0B570FDF8FB731E18" bold="true" box="[1142,1205,585,611]" pageId="38" pageNumber="239">
<taxonomicNameLabel id="6CD3FE5BCC69FFD0B570FDF8FB731E18" box="[1142,1205,585,611]" pageId="38" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
above, but differs in the following characters (character state of
<taxonomicName id="8294E4B1CC69FFD0B378FDDFFC8A1EFC" authority="Dick &amp; Ngai &amp; Doan, 2020" authorityName="Dick &amp; Ngai &amp; Doan" authorityYear="2020" box="[638,844,621,647]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="38" pageNumber="239" phylum="Bryozoa" rank="species" species="setiavicularium" status="sp. nov.">
<emphasis id="77E04320CC69FFD0B378FDDFFC8A1EFC" box="[638,844,621,647]" italics="true" pageId="38" pageNumber="239">R. setiavicularium</emphasis>
</taxonomicName>
<emphasis id="77E04320CC69FFD0B255FDDCFC541EFC" bold="true" box="[851,914,621,647]" pageId="38" pageNumber="239">
<taxonomicNameLabel id="6CD3FE5BCC69FFD0B255FDDCFC541EFC" box="[851,914,621,647]" pageId="38" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in parentheses).
<taxonomicName id="8294E4B1CC69FFD0B557FDDCFA5B1EFC" authority="Dick &amp; Ngai &amp; Doan, 2020" authorityName="Dick &amp; Ngai &amp; Doan" authorityYear="2020" box="[1105,1437,621,647]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="38" pageNumber="239" phylum="Bryozoa" rank="species" species="latiavicularium" status="sp. nov.">
<emphasis id="77E04320CC69FFD0B557FDDCFA5B1EFC" box="[1105,1437,621,647]" italics="true" pageId="38" pageNumber="239">Rhynchozoon latiavicularium</emphasis>
</taxonomicName>
<emphasis id="77E04320CC69FFD0B191FD20FF131ED0" bold="true" box="[151,213,657,683]" pageId="38" pageNumber="239">
<taxonomicNameLabel id="6CD3FE5BCC69FFD0B191FD20FF131ED0" box="[151,213,657,683]" pageId="38" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has larger zooids, average ZL × ZW = 0.54 ×
<quantity id="826C32D7CC69FFD0B3C8FD20FCF71ED0" box="[718,817,657,683]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="38" pageNumber="239" unit="mm" value="0.4">0.40 mm</quantity>
(0.46 ×
<quantity id="826C32D7CC69FFD0B28AFD20FC341ED0" box="[908,1010,657,683]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="38" pageNumber="239" unit="mm" value="0.32">0.32 mm</quantity>
); the orifical condyles are not discrete, filling angle between lateral and proximal margins of orifice (discrete, distally directed knobs); the frontal avicularian rostrum is wider relative to length, with an average L/W ratio ≈ 2.0, with a triangular mandible not distinctly nar- row distally (L/W ratio ≈ 3.0, tapering and narrow distally); the frontal avicularian rostrum is not markedly decurved (rostrum often markedly decurved); the poster in the frontal avicularia varies from acute and triangular to rounded and semicircular (sharply acute and triangular); the avicularian hinge element comprising each side of the crossbar is sharply V-shaped (hinge elements uniform in width, not V-shaped); the endooecial panel on the proximal side of the ooecium is smaller, and transversely oval or semicircular in outline (proportionally larger, semicircular).
</paragraph>
<paragraph id="452B9F32CC69FFD0B1C1FC00FEA01804" blockId="38.[151,1437,153,1187]" pageId="38" pageNumber="239">
<taxonomicName id="8294E4B1CC69FFD0B1C1FC00FDD41FB0" authority="Dick &amp; Ngai &amp; Doan, 2020" authorityName="Dick &amp; Ngai &amp; Doan" authorityYear="2020" box="[199,530,945,971]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="38" pageNumber="239" phylum="Bryozoa" rank="species" species="setiavicularium" status="sp. nov.">
<emphasis id="77E04320CC69FFD0B1C1FC00FDD41FB0" box="[199,530,945,971]" italics="true" pageId="38" pageNumber="239">Rhynchozoon setiavicularium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CD3FE5BCC69FFD0B31EFC00FD671FB0" box="[536,673,945,971]" pageId="38" pageNumber="239" rank="subSpecies">
<emphasis id="77E04320CC69FFD0B31EFC00FD931FB0" bold="true" box="[536,597,945,971]" pageId="38" pageNumber="239">n. sp.</emphasis>
and
<taxonomicName id="8294E4B1CC69FFD0B38FFC03FD671FB0" authority="Dick &amp; Ngai &amp; Doan, 2020" authorityName="Dick &amp; Ngai &amp; Doan" authorityYear="2020" box="[649,673,946,971]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="38" pageNumber="239" phylum="Bryozoa" rank="species" species="latiavicularium" status="sp. nov.">
<emphasis id="77E04320CC69FFD0B38FFC03FD671FB0" box="[649,673,946,971]" italics="true" pageId="38" pageNumber="239">R.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="77E04320CC69FFD0B3A1FC00FC921FB0" box="[679,852,945,971]" italics="true" pageId="38" pageNumber="239">latiavicularium</emphasis>
<emphasis id="77E04320CC69FFD0B25FFC00FC511FB0" bold="true" box="[857,919,945,971]" pageId="38" pageNumber="239">
<taxonomicNameLabel id="6CD3FE5BCC69FFD0B25FFC00FC511FB0" box="[857,919,945,971]" pageId="38" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are probably closely related, as the differences between them are slight.
<bibRefCitation id="2105E2C3CC69FFD0B0A9FC64FD001F94" author="Dick, M. H. &amp; Mawatari, S. F." box="[431,710,980,1007]" pageId="38" pageNumber="239" pagination="344 - 354" refId="ref28961" refString="Dick, M. H. &amp; Mawatari, S. F. (2005) Morphological and molecular concordance of Rhynchozoon clades (Bryozoa, Cheilostomata) from Alaska. Invertebrate Biology, 124, 344 - 354. https: // doi. org / 10.1111 / j. 1744 - 7410.2005.00032. x" type="journal article" year="2005">Dick &amp; Mawatari (2005)</bibRefCitation>
documented a parallel case for two similar
<taxonomicName id="8294E4B1CC69FFD0B5A8FC64FA831F94" authorityName="Hincks" authorityYear="1895" box="[1198,1349,981,1007]" class="Gymnolaemata" family="Phidoloporidae" genus="Rhynchozoon" kingdom="Animalia" order="Cheilostomatida" pageId="38" pageNumber="239" phylum="Bryozoa" rank="genus">
<emphasis id="77E04320CC69FFD0B5A8FC64FA831F94" box="[1198,1349,981,1007]" italics="true" pageId="38" pageNumber="239">Rhynchozoon</emphasis>
</taxonomicName>
species co-occurring at a locality in southeastern
<collectingRegion id="875051D0CC69FFD0B368FC48FD791868" box="[622,703,1017,1043]" country="United States of America" name="Alaska" pageId="38" pageNumber="239">Alaska</collectingRegion>
. In that study, a mitochondrial DNA (mtDNA) analysis detected two mitochondrial clades, and a morphological analysis showed slight but consistent qualitative and quantitative, clade-specific differences. One species had already been described and the other was eventually described as new (Dick
<emphasis id="77E04320CC69FFD0B1D8FBD7FEC91804" box="[222,271,1125,1151]" italics="true" pageId="38" pageNumber="239">et al</emphasis>
. 2005).
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC69FFD0B1C1FB39FCE118D8" box="[199,807,1160,1187]" pageId="38" pageNumber="239" type="distribution">
<paragraph id="452B9F32CC69FFD0B1C1FB39FCE118D8" blockId="38.[151,1437,153,1187]" box="[199,807,1160,1187]" pageId="38" pageNumber="239">
<emphasis id="77E04320CC69FFD0B1C1FB39FEA718D8" bold="true" box="[199,353,1160,1187]" pageId="38" pageNumber="239">Distribution.</emphasis>
Co To Island is the only known locality.
</paragraph>
</subSubSection>
</treatment>
</document>