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<document id="955EE4A4EBF0FEF591EF7F2F79231E40" ID-DOI="10.11646/zootaxa.4747.2.1" ID-GBIF-Dataset="55ab330e-0468-42ed-aeb0-7850751cf989" ID-ISSN="1175-5326" ID-Zenodo-Dep="3694740" ID-ZooBank="6AA8F5DC-8D70-42B0-B016-6F9C4211C471" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583220371855" checkinUser="plazi" docAuthor="Dick, Matthew H., Ngai, Nguyen Danh &amp; Doan, Hung Dinh" docDate="2020" docId="CD3D2E24CC59FFE1B191F919FC1C1E88" docLanguage="en" docName="zootaxa.4747.2.1.pdf" docOrigin="Zootaxa 4747 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hippopodina adunca Tilbrook 2006" docType="treatment" docVersion="7" lastPageNumber="224" masterDocId="3104565CCC4FFFF6B106FFB1FFC61C7B" masterDocTitle="Taxonomy and diversity of coelobite bryozoans from drift coral cobbles on Co To Island, northern Vietnam" masterLastPageNumber="252" masterPageNumber="201" pageNumber="223" updateTime="1698724047848" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DEFEA5A53D6ABDFACF793BF0052CF0CD">Taxonomy and diversity of coelobite bryozoans from drift coral cobbles on Co To Island, northern Vietnam</mods:title>
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<mods:namePart id="8A7C9F43497DB0C5C14A509DE431D1BF">Dick, Matthew H.</mods:namePart>
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<mods:namePart id="C4A333DF6DD8C60ED9445FFDDD3886EA">Ngai, Nguyen Danh</mods:namePart>
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<mods:namePart id="C2F8E52D0C4DCE52B7FAB298C1A6FF77">Doan, Hung Dinh</mods:namePart>
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<treatment id="CD3D2E24CC59FFE1B191F919FC1C1E88" ID-DOI="http://doi.org/10.5281/zenodo.3703686" ID-GBIF-Taxon="162381107" ID-Zenodo-Dep="3703686" LSID="urn:lsid:plazi:treatment:CD3D2E24CC59FFE1B191F919FC1C1E88" httpUri="http://treatment.plazi.org/id/CD3D2E24CC59FFE1B191F919FC1C1E88" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223">
<subSubSection id="0D8ECCB9CC59FFE0B191F919FD931AB9" box="[151,597,1704,1731]" pageId="22" pageNumber="223" type="nomenclature">
<paragraph id="452B9F32CC59FFE0B191F919FD931AB9" blockId="22.[151,597,1704,1766]" box="[151,597,1704,1731]" pageId="22" pageNumber="223">
<heading id="1E63285ECC59FFE0B191F919FD931AB9" bold="true" box="[151,597,1704,1731]" fontSize="11" level="1" pageId="22" pageNumber="223" reason="1">
<taxonomicName id="8294E4B1CC59FFE0B191F919FD931AB9" ID-CoL="3M2FY" authority="Tilbrook, 2006" authorityName="Tilbrook" authorityYear="2006" box="[151,597,1704,1731]" class="Gymnolaemata" family="Hippopodinidae" genus="Hippopodina" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="223" phylum="Bryozoa" rank="species" species="adunca">
<emphasis id="77E04320CC59FFE0B191F919FD931AB9" bold="true" box="[151,597,1704,1731]" pageId="22" pageNumber="223">
<emphasis id="77E04320CC59FFE0B191F919FE511AB8" bold="true" box="[151,407,1704,1731]" italics="true" pageId="22" pageNumber="223">Hippopodina adunca</emphasis>
<bibRefCitation id="2105E2C3CC59FFE0B09BF919FD931AB9" author="Tilbrook, K. J." box="[413,597,1704,1730]" pageId="22" pageNumber="223" refId="ref32691" refString="Tilbrook, K. J. (2006) Cheilostomatous Bryozoa of the Solomon Islands. Santa Barbara Museum of Natural History, Santa Barbara, 386 pp." type="book" year="2006">Tilbrook, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC59FFE0B191F97DFECB1A9D" box="[151,269,1740,1766]" pageId="22" pageNumber="223" type="description">
<paragraph id="452B9F32CC59FFE0B191F97DFECB1A9D" blockId="22.[151,597,1704,1766]" box="[151,269,1740,1766]" pageId="22" pageNumber="223">
(
<figureCitation id="DDAF83B7CC59FFE0B1A6F97DFEC31A9D" box="[160,261,1740,1766]" captionStart="FIGURE 10" captionStartId="24.[151,250,1248,1273]" captionTargetBox="[151,1436,345,1223]" captionTargetId="figure@24.[151,1436,345,1223]" captionTargetPageId="24" captionText="FIGURE 10. A. Hippopodina adunca Tilbrook, 2006, VNMN-0239, autozooids, with one ooecium. BD. Calloporina sigillata Canu &amp; Bassler, 1929, VNMN-0242. B. Colony. C. Autozooid with ooecium. D. Ancestrula (bottom right) and periancestrular zooids. Scale bars: A, 500 µm; B, 1 mm; C, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3694760" httpUri="https://zenodo.org/record/3694760/files/figure.png" pageId="22" pageNumber="223">Fig. 10A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC59FFE0B191F8A1FC861B3C" pageId="22" pageNumber="223" type="reference_group">
<paragraph id="452B9F32CC59FFE0B191F8A1FC791B52" blockId="22.[151,959,1808,1863]" box="[151,959,1808,1833]" pageId="22" pageNumber="223">
<taxonomicName id="8294E4B1CC59FFE0B191F8A1FD9F1B53" ID-CoL="3M2FY" authority="Tilbrook, 2006, p. 248" authorityName="Tilbrook" authorityPageNumber="248" authorityYear="2006" box="[151,601,1808,1833]" class="Gymnolaemata" family="Hippopodinidae" genus="Hippopodina" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="223" phylum="Bryozoa" rank="species" species="adunca">
<emphasis id="77E04320CC59FFE0B191F8A1FEA81B53" box="[151,366,1808,1832]" italics="true" pageId="22" pageNumber="223">Hippopodina adunca</emphasis>
<bibRefCitation id="2105E2C3CC59FFE0B072F8A1FDCB1B52" author="Tilbrook, K. J." box="[372,525,1808,1833]" pageId="22" pageNumber="223" refId="ref32691" refString="Tilbrook, K. J. (2006) Cheilostomatous Bryozoa of the Solomon Islands. Santa Barbara Museum of Natural History, Santa Barbara, 386 pp." type="book" year="2006">Tilbrook, 2006</bibRefCitation>
, p. 248
</taxonomicName>
, pl. 54C, D, and synonyms therein.
</paragraph>
<paragraph id="452B9F32CC59FFE0B191F89EFC861B3C" blockId="22.[151,959,1808,1863]" box="[151,832,1838,1863]" pageId="22" pageNumber="223">
<taxonomicName id="8294E4B1CC59FFE0B191F89EFD0E1B3C" ID-CoL="3M2FY" authority=": Dick &amp; Grischenko 2017, p. 211" authorityName=": Dick &amp; Grischenko" authorityPageNumber="211" authorityYear="2017" box="[151,712,1838,1863]" class="Gymnolaemata" family="Hippopodinidae" genus="Hippopodina" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="223" phylum="Bryozoa" rank="species" species="adunca">
<emphasis id="77E04320CC59FFE0B191F89EFEA81B3C" box="[151,366,1839,1863]" italics="true" pageId="22" pageNumber="223">Hippopodina adunca</emphasis>
:
<bibRefCitation id="2105E2C3CC59FFE0B07DF89FFDBB1B3C" author="Dick, M. H. &amp; Grischenko, A. V." box="[379,637,1838,1863]" pageId="22" pageNumber="223" pagination="141 - 266" refId="ref28853" refString="Dick, M. H. &amp; Grischenko, A. V. (2017) Rocky-intertidal cheilostome bryozoans from the vicinity of the Sesoko Biological Station, west-central Okinawa, Japan. Journal of Natural History, 51, 141 - 266. https: // doi. org / 10.1080 / 00222933.2016.1253797" type="journal article" year="2017">Dick &amp; Grischenko 2017</bibRefCitation>
, p. 211
</taxonomicName>
, fig. 24a, b.
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC59FFE0B191F8C4FB0B1BF4" box="[151,1229,1909,1936]" pageId="22" pageNumber="223" type="materials_examined">
<paragraph id="452B9F32CC59FFE0B191F8C4FB0B1BF4" blockId="22.[151,1436,1909,2044]" box="[151,1229,1909,1936]" pageId="22" pageNumber="223">
<emphasis id="77E04320CC59FFE0B191F8C4FE471BEB" bold="true" box="[151,385,1909,1936]" pageId="22" pageNumber="223">Material examined.</emphasis>
VNMN-0239 (CT-17), on SEM stub; VNMN-0240, two colony fragments.
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC59FFE1B1C1F82BFDB81DA9" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223" type="description">
<paragraph id="452B9F32CC59FFE0B1C1F82BFDF01B87" blockId="22.[151,1436,1909,2044]" pageId="22" pageNumber="223">
<emphasis id="77E04320CC59FFE0B1C1F82BFE711BCF" bold="true" box="[199,439,1945,1972]" pageId="22" pageNumber="223">Measurements. AzL</emphasis>
, 0.731.00 (0.850 ± 0.080);
<emphasis id="77E04320CC59FFE0B3F8F828FCFD1BCF" bold="true" box="[766,827,1945,1972]" pageId="22" pageNumber="223">AzW</emphasis>
, 0.560.84 (0.670 ± 0.079);
<emphasis id="77E04320CC59FFE0B587F828FB701BCF" bold="true" box="[1153,1206,1945,1972]" pageId="22" pageNumber="223">OrL</emphasis>
, 0.240.26 (0.248 ± 0.008);
<emphasis id="77E04320CC59FFE0B1E8F80CFEEA1BA3" bold="true" box="[238,300,1981,2008]" pageId="22" pageNumber="223">OrW</emphasis>
, 0.190.24 (0.213 ± 0.015) (n = 15, 1).
<emphasis id="77E04320CC59FFE0B3E1F80CFC161BA3" bold="true" box="[743,976,1981,2008]" pageId="22" pageNumber="223">OrL, ooeciate zooid</emphasis>
, 0.224;
<emphasis id="77E04320CC59FFE0B52DF80CFADA1BA3" bold="true" box="[1067,1308,1981,2008]" pageId="22" pageNumber="223">OrW, ooeciate zooid</emphasis>
, 0.263 (n = 1).
<emphasis id="77E04320CC59FFE0B1BBF850FF351B87" bold="true" box="[189,243,2017,2044]" pageId="22" pageNumber="223">OvL</emphasis>
, 0.602;
<emphasis id="77E04320CC59FFE0B049F850FE491B87" bold="true" box="[335,399,2017,2044]" pageId="22" pageNumber="223">OvW</emphasis>
, 0.619 (n = 1).
</paragraph>
<paragraph id="452B9F32CC58FFE1B1C1FF29FAAB1CC9" blockId="23.[151,1437,152,755]" box="[199,1389,152,179]" pageId="23" pageNumber="224">
<emphasis id="77E04320CC58FFE1B1C1FF29FE9F1CC8" bold="true" box="[199,345,152,179]" pageId="23" pageNumber="224">Description.</emphasis>
Colony encrusting, sheet-like, unilaminar but occasionally bilaminar due to self-overgrowth.
</paragraph>
<paragraph id="452B9F32CC58FFE1B1C1FF0DFE761D39" blockId="23.[151,1437,152,755]" pageId="23" pageNumber="224">Zooids large, rectangular, delineated by groove. Frontal wall convex, perforated by numerous tiny pseudopores. Orifice hoof-shaped; poster same width as anter or slightly narrower, separated by sharp condyles; proximal margin of orifice straight or markedly concave. Orifices dimorphic, with those of ooecium-bearing zooids wider than those of zooids lacking ooecia.</paragraph>
<paragraph id="452B9F32CC58FFE1B1C1FEFDFAFE1DF1" blockId="23.[151,1437,152,755]" pageId="23" pageNumber="224">Adventitious avicularia usually paired, sometimes single; each with large chamber lateral to orifice, rostrum long, tapering, directed distomedially and slightly curving around distal end of orifice, crossbar complete.</paragraph>
<paragraph id="452B9F32CC58FFE1B1C1FE25FDB81DA9" blockId="23.[151,1437,152,755]" pageId="23" pageNumber="224">Ooecium prominent, resting on frontal wall of next-distal zooid; circular in outline, perforated over entire surface with tiny pseudopores. Spines lacking.</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC58FFE1B1C1FE6CFA4F1ED1" pageId="23" pageNumber="224" type="discussion">
<paragraph id="452B9F32CC58FFE1B1C1FE6CFA4F1ED1" blockId="23.[151,1437,152,755]" pageId="23" pageNumber="224">
<emphasis id="77E04320CC58FFE1B1C1FE6CFEFD1D8C" bold="true" box="[199,315,477,503]" pageId="23" pageNumber="224">Remarks.</emphasis>
The large avicularia having setiform mandibles directed distomedially are typical of
<taxonomicName id="8294E4B1CC58FFE1B41BFE6CFA531D8D" authorityName=": Dick &amp; Grischenko" authorityYear="2017" box="[1309,1429,476,502]" class="Gymnolaemata" family="Hippopodinidae" genus="Hippopodina" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="224" phylum="Bryozoa" rank="species" species="adunca">
<emphasis id="77E04320CC58FFE1B41BFE6CFA531D8D" box="[1309,1429,476,502]" italics="true" pageId="23" pageNumber="224">H. adunca</emphasis>
</taxonomicName>
, which, however, also typically has the proximal margin of the orifice straight. While some zooids in the specimens from Co To have the proximal orificial margin nearly straight, most have a concave, distinctly rounded margin. In addition, the only ooeciate zooid observed has a sharp, raised suboral collar. A key character that helps differentiate among
<taxonomicName id="8294E4B1CC58FFE1B013FDDDFE6F1EFD" authorityName="Levinsen" authorityYear="1909" box="[277,425,620,646]" class="Gymnolaemata" family="Hippopodinidae" genus="Hippopodina" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="224" phylum="Bryozoa" rank="genus">
<emphasis id="77E04320CC58FFE1B013FDDDFE6F1EFD" box="[277,425,620,646]" italics="true" pageId="23" pageNumber="224">Hippopodina</emphasis>
</taxonomicName>
species is the ancestrular complex, which forms a triad in in some species (including
<taxonomicName id="8294E4B1CC58FFE1B487FDDCFF2F1ED1" authorityName=": Dick &amp; Grischenko" authorityYear="2017" class="Gymnolaemata" family="Hippopodinidae" genus="Hippopodina" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="224" phylum="Bryozoa" rank="species" species="adunca">
<emphasis id="77E04320CC58FFE1B487FDDCFF2F1ED1" italics="true" pageId="23" pageNumber="224">H. adunca</emphasis>
</taxonomicName>
) but a tetrad in others (
<bibRefCitation id="2105E2C3CC58FFE1B0E8FD21FD551ED0" author="Tilbrook, K. J." box="[494,659,656,683]" pageId="23" pageNumber="224" pagination="449 - 456" refId="ref32630" refString="Tilbrook, K. J. (1999) Description of Hippopodina feegeensis and three other species of Hippopodina Levinsen, 1909 (Bryozoa: Cheilostomatida). Journal of the Zoological Society of London, 247, 449 - 456. https: // doi. org / 10.1111 / j. 1469 - 7998.1999. tb 01008. x" type="journal article" year="1999">Tilbrook 1999</bibRefCitation>
, 2006). No ancestrular complex was found in the Co To specimens.
</paragraph>
</subSubSection>
<subSubSection id="0D8ECCB9CC58FFE1B1C1FD05FC1C1E88" pageId="23" pageNumber="224" type="distribution">
<paragraph id="452B9F32CC58FFE1B1C1FD05FC1C1E88" blockId="23.[151,1437,152,755]" pageId="23" pageNumber="224">
<emphasis id="77E04320CC58FFE1B1C1FD05FEA71EB4" bold="true" box="[199,353,692,719]" pageId="23" pageNumber="224">Distribution.</emphasis>
Broadly distributed in the Indo-West Pacific, ranging from
<collectingCountry id="3D83DFA2CC58FFE1B506FD05FBAB1EB5" box="[1024,1133,692,718]" name="Mauritius" pageId="23" pageNumber="224">Mauritius</collectingCountry>
westward to
<collectingCountry id="3D83DFA2CC58FFE1B402FD05FAED1EB5" box="[1284,1323,692,718]" name="Fiji" pageId="23" pageNumber="224">Fiji</collectingCountry>
(
<bibRefCitation id="2105E2C3CC58FFE1B43FFD05FF171E88" author="Tilbrook, K. J." pageId="23" pageNumber="224" refId="ref32691" refString="Tilbrook, K. J. (2006) Cheilostomatous Bryozoa of the Solomon Islands. Santa Barbara Museum of Natural History, Santa Barbara, 386 pp." type="book" year="2006">Tilbrook 2006</bibRefCitation>
) and northward as far as
<collectingRegion id="875051D0CC58FFE1B0EAFD69FD921E89" box="[492,596,728,754]" country="Japan" name="Okinawa" pageId="23" pageNumber="224">Okinawa</collectingRegion>
,
<collectingCountry id="3D83DFA2CC58FFE1B359FD68FD591E88" box="[607,671,729,755]" name="Japan" pageId="23" pageNumber="224">Japan</collectingCountry>
(
<bibRefCitation id="2105E2C3CC58FFE1B3ABFD69FC081E88" author="Dick, M. H. &amp; Grischenko, A. V." box="[685,974,728,755]" pageId="23" pageNumber="224" pagination="141 - 266" refId="ref28853" refString="Dick, M. H. &amp; Grischenko, A. V. (2017) Rocky-intertidal cheilostome bryozoans from the vicinity of the Sesoko Biological Station, west-central Okinawa, Japan. Journal of Natural History, 51, 141 - 266. https: // doi. org / 10.1080 / 00222933.2016.1253797" type="journal article" year="2017">Dick &amp; Grischenko 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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