318 lines
43 KiB
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318 lines
43 KiB
XML
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<mods:title id="92D246F810083A7B00481F6DBA22590E">Notes on Oriental Clivinini and the description of two new species (Coleoptera: Carabidae: Clivinini)</mods:title>
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<taxonomicName id="82BD99DCFFE5053AEFD2FAC1EEECFAC6" ID-CoL="8MJGT" authority="LESNE, 1896" box="[178,416,1287,1309]" class="Insecta" family="Carabidae" genus="Rugiluclivina" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="alutacea">
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<emphasis id="77C93E4DFFE5053AEFD2FAC1EEECFAC6" bold="true" box="[178,416,1287,1309]" italics="true" pageId="8" pageNumber="205">Rugiluclivina alutacea</emphasis>
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<bibRefCitation id="212C9FAEFFE5053AEEC5FAC1EEA6FAC7" author="LESNE P" box="[421,490,1287,1308]" pageId="8" pageNumber="205" pagination="238 - 259" refId="ref9588" refString="LESNE P. (1896): Cicindelides et Carabides Indo-Chinois recuellis par M. Pavie. Diagnoses des especes nouvelles et d'un genre nouveau. - Bulletin du Muse um d'Histoire Naturelle de Paris 1896: 238 - 259." type="journal article" year="1896">LESNE</bibRefCitation>
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,
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1896 comb.nov. (
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<figureCitation id="DD86FEDAFFE5053AEDCDFAC1EC5CFAC6" box="[685,784,1287,1309]" captionStart="Figs 8-11" captionStartId="10.[178,218,1513,1535]" captionTargetBox="[186,1081,216,1463]" captionTargetId="figure@10.[178,1085,204,1510]" captionText="Figs 8-11: Rugiluclivina alutacea LESNE: (8) Holotype, habitus, with original mounting card and pin; (9) Original lables; (10) Aedeagus and parameres, dorsal view, specimen from Cambodia; (11) Female coxostylus, holotype." figureDoi="http://doi.org/10.5281/zenodo.3998153" httpUri="https://zenodo.org/record/3998153/files/figure.png" pageId="8" pageNumber="205">Figs 8-11</figureCitation>
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)
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<emphasis id="77C93E4DFFE5053AEFD2FAEBEE05FA99" box="[178,329,1325,1346]" italics="true" pageId="8" pageNumber="205">Clivina alutacea</emphasis>
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<emphasis id="77C93E4DFFE5053AEFD2FA81EE05FA87" box="[178,329,1351,1372]" italics="true" pageId="8" pageNumber="205">Clivina alutacea</emphasis>
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<bibRefCitation id="212C9FAEFFE5053AEE2EFA8EEEA4FA86" author="LESNE P" box="[334,488,1351,1373]" pageId="8" pageNumber="205" pagination="1879 - 1895" refId="ref9629" refString="LESNE P. (1904): Famille des Carabides. In: Pavie A. Mission Pavie Indo-Chine 1879 - 1895. Etudes diverses, III Recherches sur l'histoire Naturelle de Indo-Chine Orientale. - Paris: Ernst Leroux, pp. 549." type="journal article" year="1904">LESNE, 1904: 67</bibRefCitation>
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<emphasis id="77C93E4DFFE5053AEFD2FAA7EE05FAAD" box="[178,329,1377,1398]" italics="true" pageId="8" pageNumber="205">Clivina alutacea</emphasis>
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<bibRefCitation id="212C9FAEFFE5053AEE2EFAA4EE8AFAAC" author="LESNE P" box="[334,454,1377,1399]" pageId="8" pageNumber="205" pagination="238 - 259" refId="ref9588" refString="LESNE P. (1896): Cicindelides et Carabides Indo-Chinois recuellis par M. Pavie. Diagnoses des especes nouvelles et d'un genre nouveau. - Bulletin du Muse um d'Histoire Naturelle de Paris 1896: 238 - 259." type="journal article" year="1896">LESNE, 1896</bibRefCitation>
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<emphasis id="77C93E4DFFE5053AEFD2FABAEE05FA4A" box="[178,329,1404,1425]" italics="true" pageId="8" pageNumber="205">Clivina alutacea</emphasis>
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<bibRefCitation id="212C9FAEFFE5053AEE2EFABAEE8AFA4A" author="LESNE P" box="[334,454,1403,1425]" pageId="8" pageNumber="205" pagination="238 - 259" refId="ref9588" refString="LESNE P. (1896): Cicindelides et Carabides Indo-Chinois recuellis par M. Pavie. Diagnoses des especes nouvelles et d'un genre nouveau. - Bulletin du Muse um d'Histoire Naturelle de Paris 1896: 238 - 259." type="journal article" year="1896">LESNE, 1896</bibRefCitation>
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,
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<taxonomicName id="82BD99DCFFE5053AEFD2FA50EE8AFA77" ID-CoL="8LBVD" authority="LESNE, 1896" authorityName="Lesne" authorityYear="1896" box="[178,454,1430,1452]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="206" phylum="Arthropoda" rank="species" species="alutacea">
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<emphasis id="77C93E4DFFE5053AEFD2FA50EE05FA70" box="[178,329,1430,1451]" italics="true" pageId="8" pageNumber="205">Clivina alutacea</emphasis>
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<bibRefCitation id="212C9FAEFFE5053AEE2EFA50EE8AFA77" author="LESNE P" box="[334,454,1430,1452]" pageId="8" pageNumber="205" pagination="238 - 259" refId="ref9588" refString="LESNE P. (1896): Cicindelides et Carabides Indo-Chinois recuellis par M. Pavie. Diagnoses des especes nouvelles et d'un genre nouveau. - Bulletin du Muse um d'Histoire Naturelle de Paris 1896: 238 - 259." type="journal article" year="1896">LESNE, 1896</bibRefCitation>
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,
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y p e m a t e r i a l:
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<typeStatus id="9A065CFDFFE5053AEECDFA73ED4DFA10" box="[429,513,1461,1483]" pageId="8" pageNumber="205" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="53BB29D6FFE5053AED6BFA7DED50FA0B" box="[523,540,1467,1488]" pageId="8" pageNumber="205" type="female">♀</specimenCount>
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, with labels and data: beige, circular, handwritten with black ink "2579 86", backside yellow / light-brown, rectangular, printed "
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<collectionCode id="23AC7A9AFFE5053AECEBFA08EB71FA38" box="[907,1085,1486,1507]" pageId="8" pageNumber="205">MUSEUM PARIS</collectionCode>
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A.
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<specimenCode id="151B4A24FFE5053AEE1FFA20EEBFFA27" box="[383,499,1510,1532]" collectionCode="PAVIE" pageId="8" pageNumber="205">PAVIE 1886</specimenCode>
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" / beige, rectangular, handwritten with black ink "
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<taxonomicName id="82BD99DCFFE5053AECCEFA20EE5FF9C8" ID-CoL="8LBVD" authority="Lesne" authorityName="Lesne" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="alutacea">Clivina alutacea Lesne</taxonomicName>
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Type" / beige, rectangular, handwritten with black ink and printed "
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<taxonomicName id="82BD99DCFFE5053AEC1AFA38ECB2F9C8" ID-CoL="8LBVD" authority="Lesne" authorityName="Lesne" box="[890,1022,1534,1556]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="alutacea">alutacea Lesne</taxonomicName>
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det.
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<determinerName id="B0DE0E9BFFE5053AEE64F9D0EE0AF9F7" box="[260,326,1558,1580]" pageId="8" pageNumber="205">K. Kult</determinerName>
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57"; labels as in
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<figureCitation id="DD86FEDAFFE5053AEEBAF9D0ED40F9F0" box="[474,524,1558,1580]" captionStart="Figs 8-11" captionStartId="10.[178,218,1513,1535]" captionTargetBox="[186,1081,216,1463]" captionTargetId="figure@10.[178,1085,204,1510]" captionText="Figs 8-11: Rugiluclivina alutacea LESNE: (8) Holotype, habitus, with original mounting card and pin; (9) Original lables; (10) Aedeagus and parameres, dorsal view, specimen from Cambodia; (11) Female coxostylus, holotype." figureDoi="http://doi.org/10.5281/zenodo.3998153" httpUri="https://zenodo.org/record/3998153/files/figure.png" pageId="8" pageNumber="205">
|
|
<collectorName id="E8488789FFE5053AEEBAF9D0EEB1F9F7" box="[474,509,1558,1580]" pageId="8" pageNumber="205">Fig.</collectorName>
|
|
9
|
|
</figureCitation>
|
|
(
|
|
<collectionCode id="23AC7A9AFFE5053AED78F9D0ED2CF9F7" box="[536,608,1558,1580]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="205">MNHN</collectionCode>
|
|
).
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0DA7B1D4FFE4053EEFD2FF1DEE14FCDD" lastPageId="12" lastPageNumber="209" pageId="9" pageNumber="206" type="description">
|
|
<paragraph id="4502E25FFFE4053BEFD2FF1DEB72FE19" blockId="9.[178,1087,219,618]" pageId="9" pageNumber="206">
|
|
<materialsCitation id="F5D5E802FFE4053BEFD2FF1DECB0FE92" ID-GBIF-Occurrence="2848386307" collectionCode="R" pageId="9" pageNumber="206" specimenCount="2" typeStatus="holotype">
|
|
<collectionCode id="23AC7A9AFFE4053BEFD2FF1DEF8FFF2B" box="[178,195,219,240]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="206">R</collectionCode>
|
|
e m a r k: The
|
|
<typeStatus id="9A065CFDFFE4053BEE17FF1DEE9CFF2A" box="[375,464,219,241]" pageId="9" pageNumber="206" type="holotype">holotype</typeStatus>
|
|
is not in a good condition. The pieces of the body were mounted together some time ago with black glue. The same is true for some of the specimens from the additional historic material. In the
|
|
<typeStatus id="9A065CFDFFE4053BED85FED3EC72FEF0" box="[741,830,277,299]" pageId="9" pageNumber="206" type="holotype">holotype</typeStatus>
|
|
the following parts are missing: Left antennae from joint three onwards, right one from joint six onwards
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="F5D5E802FFE4053BEB67FEF5EB76FE19" ID-GBIF-Occurrence="2848386304" collectionCode="LESNE" collectorName="The" pageId="9" pageNumber="206" specimenCount="1" typeStatus="holotype">
|
|
movable spur of left front tibia, terminal three tarsalia of left intermediate leg and the complete right hind leg.
|
|
<collectorName id="E8488789FFE4053BEEF2FEABEEF5FE58" box="[402,441,365,387]" pageId="9" pageNumber="206">The</collectorName>
|
|
right elytron and the dorsal integument shows a big whole where the original not sharpened pin was placed. A d d i t i o n a l m a t e r i a l seen by
|
|
<collectionCode id="23AC7A9AFFE4053BED38FE6AEDDAFE1A" box="[600,662,428,449]" pageId="9" pageNumber="206">LESNE</collectionCode>
|
|
:
|
|
<specimenCount id="53BB29D6FFE4053BEDC5FE6AEDABFE1A" box="[677,743,428,449]" pageId="9" pageNumber="206" type="generic">1 spec.</specimenCount>
|
|
with labels and data: beige, circular
|
|
</materialsCitation>
|
|
,
|
|
</paragraph>
|
|
<paragraph id="4502E25FFFE4053BEFBFFE02EC18FDB1" blockId="9.[178,1087,219,618]" pageId="9" pageNumber="206">
|
|
handwritten with black ink "1036 75" backside yellow / light-brown, rectangular, printed "MUSEUM PARIS COCHINCHINE JULIEN 1875" / beige, rectangular, handwritten with black ink "
|
|
<taxonomicName id="82BD99DCFFE4053BEE6FFE32EEAAFDD2" authority="Lesne" authorityName="Lesne" box="[271,486,500,522]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="206" phylum="Arthropoda" rank="species" species="alutacea">Clivina alutacea Lesne</taxonomicName>
|
|
" (MNHN).
|
|
<specimenCount id="53BB29D6FFE4053BED3DFE32ED36FDD2" box="[605,634,500,521]" pageId="9" pageNumber="206" type="male">1♂</specimenCount>
|
|
,
|
|
<specimenCount id="53BB29D6FFE4053BEDEBFE32ED9DFDD2" box="[651,721,500,521]" pageId="9" pageNumber="206" type="generic">1 spec.</specimenCount>
|
|
, with labels and data: beige, circular, handwritten with black ink "2579 86" backside yellow / light-brown, rectangular, printed "MUSEUM PARIS
|
|
<collectingCountry id="3DAAA2CFFFE4053BEEFAFDE2ED55FDE2" box="[410,537,548,569]" name="Cambodia" pageId="9" pageNumber="206">CAMBODGE</collectingCountry>
|
|
A. PAVIE 1886" (MNHN). 2 specs., with labels and data: beige, circular, handwritten with black ink "1036 75" backside yellow / light-brown, rectangular, printed "MUSEUM PARIS COCHINCHINE JULIEN 1875" (MNHN).
|
|
</paragraph>
|
|
<paragraph id="4502E25FFFE4053BEFD2FDB1EC9DFCEF" blockId="9.[178,1085,631,821]" pageId="9" pageNumber="206">
|
|
D i a g n o s i s: A medium sized black
|
|
<taxonomicName id="82BD99DCFFE4053BED2EFDB1ED9BFD57" box="[590,727,631,652]" class="Insecta" family="Carabidae" genus="Rugiluclivina" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="206" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="77C93E4DFFE4053BED2EFDB1ED9BFD57" box="[590,727,631,652]" italics="true" pageId="9" pageNumber="206">Rugiluclivina</emphasis>
|
|
</taxonomicName>
|
|
species with multiple fine carinae on the frons of the head, shorter and completely reticulated elytra. Distinguished from the most similar species
|
|
<taxonomicName id="82BD99DCFFE4053BEECEFD76ED61FD1F" authorityName="Balkenohl" authorityYear="1996" box="[430,557,687,709]" class="Insecta" family="Carabidae" genus="Rugiluclivina" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="206" phylum="Arthropoda" rank="species" species="reticulata">
|
|
<emphasis id="77C93E4DFFE4053BEECEFD76ED61FD1F" box="[430,557,687,709]" italics="true" pageId="9" pageNumber="206">R. reticulata</emphasis>
|
|
</taxonomicName>
|
|
BALKENOHL by the completely convex and smooth reflexed lateral margin of the pronotum. In addition, the intervals of the elytra are convex, the second setigerous puncture on interval three is situated behind middle, and the striae one and two, three and four, and five and six joining at apex. Moreover, the aedeagus and the apical setae of the male parameres exhibit a different pattern.
|
|
</paragraph>
|
|
<paragraph id="4502E25FFFE4053BEFD2FC85EEC7FC83" blockId="9.[178,395,835,857]" box="[178,395,835,857]" pageId="9" pageNumber="206">R e d e s c r i p t i o n:</paragraph>
|
|
<paragraph id="4502E25FFFE4053BEFD2FCA1EDD3FC43" blockId="9.[178,1085,871,921]" pageId="9" pageNumber="206">
|
|
Measurements:
|
|
<typeStatus id="9A065CFDFFE4053BEE36FCA1EEF4FCA6" box="[342,440,871,893]" pageId="9" pageNumber="206" type="holotype">Holotype</typeStatus>
|
|
: Body length
|
|
<quantity id="82454FBAFFE4053BED2CFCA1EDE7FCA7" box="[588,683,871,892]" metricMagnitude="-3" metricUnit="m" metricValue="8.12" pageId="9" pageNumber="206" unit="mm" value="8.12">8.12 mm</quantity>
|
|
; width
|
|
<quantity id="82454FBAFFE4053BED9BFCA1EC17FCA7" box="[763,859,871,892]" metricMagnitude="-3" metricUnit="m" metricValue="2.31" pageId="9" pageNumber="206" unit="mm" value="2.31">2.31 mm</quantity>
|
|
; ratio length/width of pronotum 0.89; ratio length/width of elytra 1.90.
|
|
</paragraph>
|
|
<paragraph id="4502E25FFFE4053BEFD2FC61ED7FFC2E" blockId="9.[178,1085,934,1013]" pageId="9" pageNumber="206">
|
|
Other material: Body length
|
|
<quantity id="82454FBAFFE4053BEEBFFC61ED3CFC67" box="[479,624,935,956]" metricMagnitude="-3" metricUnit="m" metricValue="7.779999999999999" metricValueMax="8.12" metricValueMin="7.44" pageId="9" pageNumber="206" unit="mm" value="7.779999999999999" valueMax="8.12" valueMin="7.44">7.44-8.12 mm</quantity>
|
|
(ẍ =
|
|
<quantity id="82454FBAFFE4053BEDD0FC61EC5FFC66" box="[688,787,935,957]" metricMagnitude="-3" metricUnit="m" metricValue="7.79" pageId="9" pageNumber="206" unit="mm" value="7.79">7.79 mm</quantity>
|
|
*); width
|
|
<quantity id="82454FBAFFE4053BEC14FC61EB49FC67" box="[884,1029,935,956]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" metricValueMax="2.4" metricValueMin="2.14" pageId="9" pageNumber="206" unit="mm" value="2.27" valueMax="2.4" valueMin="2.14">2.14-2.40 mm</quantity>
|
|
(ẍ =
|
|
<quantity id="82454FBAFFE4053BEFD2FC05EE58FC02" box="[178,276,963,985]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="9" pageNumber="206" unit="mm" value="2.25">2.25 mm</quantity>
|
|
*); ratio length/width of pronotum 0.89-0.91 (ẍ = 0.89*); ratio length/width of elytra 1.88-1.95 (ẍ = 1.90*); (*n = 6).
|
|
</paragraph>
|
|
<paragraph id="4502E25FFFE4053BEFD2FBC5EE5AFB8A" blockId="9.[178,1085,1027,1105]" pageId="9" pageNumber="206">Colour: Dorsal and ventral surface dull; nigricans; mandibles niger; mouthparts, intermediate and hind legs fuscus; antennae with scapus and pedicellus fuscus, other parts nigricans.</paragraph>
|
|
<paragraph id="4502E25FFFE40538EFD2FB99EC30FA06" blockId="9.[178,1086,1119,1541]" lastBlockId="10.[834,892,1466,1501]" lastPageId="10" lastPageNumber="207" pageId="9" pageNumber="206">
|
|
Head (
|
|
<figureCitation id="DD86FEDAFFE4053BEF97FB99EE7AFBAD" box="[247,310,1119,1142]" captionStart="Figs 8-11" captionStartId="10.[178,218,1513,1535]" captionTargetBox="[186,1081,216,1463]" captionTargetId="figure@10.[178,1085,204,1510]" captionText="Figs 8-11: Rugiluclivina alutacea LESNE: (8) Holotype, habitus, with original mounting card and pin; (9) Original lables; (10) Aedeagus and parameres, dorsal view, specimen from Cambodia; (11) Female coxostylus, holotype." figureDoi="http://doi.org/10.5281/zenodo.3998153" httpUri="https://zenodo.org/record/3998153/files/figure.png" pageId="9" pageNumber="206">Fig. 8</figureCitation>
|
|
): One fourth smaller than pronotum. Outline subsquare. Clypeus relatively wide, straight anteriorly, with two teeth anteriorly, reflexed margined, separated from frons by deep transverse furrow, distinctly separated from wings by notches, surface with five carinae in form of an inverted V. Wings somewhat wider than teeth of clypeus, as far projecting anteriorly as teeth of clypeus, surface with few longitudinal rugae. Supraantennal plates vaulted, laterally amargined, projecting widely laterally up to mideye level, with small obtuse rounded angle anteriorly, projecting anteriorly less far as wings, separated from wings by distinct notches, surface with irregular longitudinal carinae. Furrows between clypeus, wings, supraantennal plates joining posteriorly. Two clypeal setae situated at joining point of furrows, joined furrows joining with transverse furrow at level of mid-eye. Supraorbital furrow deep, wide, diverging posteriorly and running to level of posterior third of eye. Supraorbital carina invisible, bilaterally with two distinct supraorbital setigerous punctures. Frons distinctly convex, surface with irregular carinae laterally and at level of transverse furrow, completely irregular and fine at middle. Neck constriction slight but well visible laterally. Eyes subglobose, projecting
|
|
<emphasis id="77C93E4DFFE70538EC22FA7CEC30FA06" bold="true" box="[834,892,1466,1501]" pageId="10" pageNumber="207">10</emphasis>
|
|
</paragraph>
|
|
<caption id="11C2B2D7FFE70538EFD2FA2FEEF1F9F6" ID-DOI="http://doi.org/10.5281/zenodo.3998153" ID-Zenodo-Dep="3998153" httpUri="https://zenodo.org/record/3998153/files/figure.png" pageId="10" pageNumber="207" startId="10.[178,218,1513,1535]" targetBox="[186,1081,216,1463]" targetPageId="10">
|
|
<paragraph id="4502E25FFFE70538EFD2FA2FEEF1F9F6" blockId="10.[178,1085,1513,1581]" pageId="10" pageNumber="207">
|
|
<emphasis id="77C93E4DFFE70538EFD2FA2FEE47FA25" bold="true" box="[178,267,1513,1535]" pageId="10" pageNumber="207">Figs 8-11</emphasis>
|
|
:
|
|
<taxonomicName id="82BD99DCFFE70538EE7AFA2FEEA6FA25" box="[282,490,1513,1534]" class="Insecta" family="Carabidae" genus="Rugiluclivina" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="207" phylum="Arthropoda" rank="species" species="alutacea">
|
|
<emphasis id="77C93E4DFFE70538EE7AFA2FEEA6FA25" box="[282,490,1513,1534]" italics="true" pageId="10" pageNumber="207">Rugiluclivina alutacea</emphasis>
|
|
</taxonomicName>
|
|
<materialsCitation id="F5D5E802FFE70538EE92FA2CEEF1F9F6" ID-GBIF-Occurrence="2848386303" collectionCode="LESNE" country="Cambodia" county="Female" location="Original" municipality="Aedeagus" pageId="10" pageNumber="207" specimenCount="2" typeStatus="holotype">
|
|
<collectionCode id="23AC7A9AFFE70538EE92FA2CED63FA25" box="[498,559,1514,1535]" pageId="10" pageNumber="207">LESNE</collectionCode>
|
|
: (
|
|
<emphasis id="77C93E4DFFE70538ED24FA2FED03FA25" bold="true" box="[580,591,1513,1534]" pageId="10" pageNumber="207">8</emphasis>
|
|
)
|
|
<typeStatus id="9A065CFDFFE70538ED3FFA2FEDFAFA24" box="[607,694,1513,1535]" pageId="10" pageNumber="207" type="holotype">Holotype</typeStatus>
|
|
, habitus, with original mounting card and pin; (
|
|
<emphasis id="77C93E4DFFE70538EF83F9C6EFA2F9CE" bold="true" box="[227,238,1536,1557]" pageId="10" pageNumber="207">9</emphasis>
|
|
)
|
|
<location id="4062B484FFE70538EF9CF9C6EE04F9CD" LSID="urn:lsid:plazi:treatment:CD145349FFE5053EEFD2FAC1EE14FCDD:4062B484FFE70538EF9CF9C6EE04F9CD" box="[252,328,1536,1558]" country="Cambodia" county="Female" municipality="Aedeagus" name="Original" pageId="10" pageNumber="207">Original</location>
|
|
lables; (
|
|
<emphasis id="77C93E4DFFE70538EEF6F9C6EEE1F9CE" bold="true" box="[406,429,1536,1557]" pageId="10" pageNumber="207">10</emphasis>
|
|
)
|
|
<collectingMunicipality id="A5667825FFE70538EEDBF9C6ED58F9CD" box="[443,532,1536,1558]" pageId="10" pageNumber="207">Aedeagus</collectingMunicipality>
|
|
and parameres, dorsal view, specimen from
|
|
<collectingCountry id="3DAAA2CFFFE70538ECCDF9C6EB41F9CD" box="[941,1037,1536,1558]" name="Cambodia" pageId="10" pageNumber="207">Cambodia</collectingCountry>
|
|
; (
|
|
<emphasis id="77C93E4DFFE70538EB7FF9C6EB7AF9CE" bold="true" box="[1055,1078,1536,1557]" pageId="10" pageNumber="207">11</emphasis>
|
|
)
|
|
<collectingCounty id="AC639AD3FFE70538EFD2F9D1EFB9F9F6" box="[178,245,1559,1581]" pageId="10" pageNumber="207">Female</collectingCounty>
|
|
coxostylus,
|
|
<typeStatus id="9A065CFDFFE70538EE07F9D1EEF5F9F6" box="[359,441,1559,1581]" pageId="10" pageNumber="207" type="holotype">holotype</typeStatus>
|
|
.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="4502E25FFFE60539EFD2FF1CED91FE30" blockId="11.[178,1085,218,492]" pageId="11" pageNumber="208">laterally, genae indistinct. Antenna elongated, ratio length/width of segments four and five 1.82, scapus with longitudinal reticulation, seta on scapus situated dorso-frontally at apical tip, segment two attached eccentrically, segment three pubescent in apical twothirds, fully pubescent from segment four onwards. Labrum wider as clypeus, bilobed, conspicuously excised at middle, amargined, six-setose, ciliate laterally, surface with transverse reticulation in basal part. Mandible distinctly shorter than head, slender in apical third, acute, apical half bent dorsally, with few longitudinal carinae. Maxillary palpus slender, two apical segments bent laterally. Apical segment of labial palpomere fusiform, segment two tumid, with one seta. Ligula long, wide, spatulate, with small tooth at each side, with one apical setigerous puncture.</paragraph>
|
|
<paragraph id="4502E25FFFE60539EFD2FE3DECE5FCF5" blockId="11.[178,1085,506,814]" pageId="11" pageNumber="208">
|
|
Pronotum (
|
|
<figureCitation id="DD86FEDAFFE60539EE44FE3CEE2FFDCB" box="[292,355,506,528]" captionStart="Figs 8-11" captionStartId="10.[178,218,1513,1535]" captionTargetBox="[186,1081,216,1463]" captionTargetId="figure@10.[178,1085,204,1510]" captionText="Figs 8-11: Rugiluclivina alutacea LESNE: (8) Holotype, habitus, with original mounting card and pin; (9) Original lables; (10) Aedeagus and parameres, dorsal view, specimen from Cambodia; (11) Female coxostylus, holotype." figureDoi="http://doi.org/10.5281/zenodo.3998153" httpUri="https://zenodo.org/record/3998153/files/figure.png" pageId="11" pageNumber="208">Fig. 8</figureCitation>
|
|
): Subquadrate, slightly convex on disc (lateral view), distinctly convex in frontal view. Anterior angles pointed, slightly rounded and projecting. Posterior angles distinct, obtusely toothed. Reflexed lateral margin distinctly convex, more rounded to angles in anterior and posterior quarter. Lateral channel narrow, of same size in whole length, anterior setigerous puncture connected with channel, posterior seta removed from channel by diameter of pore. Base slightly produced at middle. Reflexed margin smooth, thin in whole length, running from posterior angles to base as straight line. Anterior transverse line deep, reaching anterior margin, not joining lateral margin. Median line distinct, complete, joining with anterior transverse line, narrow towards base. Dorsal surface conspicuously coreaceate, lateral transverse rugae indistinct or missing, with isodiametric reticulation baso-laterally. Basal impressions short, indistinct.
|
|
</paragraph>
|
|
<paragraph id="4502E25FFFE60539EFD2FCFAEE61FB57" blockId="11.[178,1086,828,1164]" pageId="11" pageNumber="208">
|
|
Elytra (
|
|
<figureCitation id="DD86FEDAFFE60539EE60FCFAEE0CFC89" box="[256,320,828,850]" captionStart="Figs 8-11" captionStartId="10.[178,218,1513,1535]" captionTargetBox="[186,1081,216,1463]" captionTargetId="figure@10.[178,1085,204,1510]" captionText="Figs 8-11: Rugiluclivina alutacea LESNE: (8) Holotype, habitus, with original mounting card and pin; (9) Original lables; (10) Aedeagus and parameres, dorsal view, specimen from Cambodia; (11) Female coxostylus, holotype." figureDoi="http://doi.org/10.5281/zenodo.3998153" httpUri="https://zenodo.org/record/3998153/files/figure.png" pageId="11" pageNumber="208">Fig. 8</figureCitation>
|
|
): Cylindriform, slightly transversally depressed in anterior third in lateral view, distinctly convex in frontal view. Slightly dilated in apical half, slightly contracted to humerus. Humerus rounded, rectangular. Base concave at declivity, finely margined from humerus to peduncle, declivity slightly overhanging over parts of peduncle, setigerous puncture at base of first stria, with one small tubercle at base of each free interval. Scutellar stria indistinct, without tubercle-like carina. Striae punctuate-striate, one to four free at base, two to six not reaching tip of apex, one and two, three and four, and five and six joining at apex. Intervals flattened to moderately convex, seventh carinate at humerus and apex, eights completely carinate. Interval three with four setigerous punctures, situated at the third stria, second setigerous puncture situated distinctly behind middle of elytra. Reflexed lateral margin smooth. Whole surface covered with dense isodiametric reticulation.
|
|
</paragraph>
|
|
<paragraph id="4502E25FFFE60539EFD2FB5CEEACFB6B" blockId="11.[178,480,1178,1200]" box="[178,480,1178,1200]" pageId="11" pageNumber="208">Hind wings: Fully developed.</paragraph>
|
|
<paragraph id="4502E25FFFE60539EFD2FB79EC43FAF0" blockId="11.[178,1085,1215,1323]" pageId="11" pageNumber="208">Ventral surface: Proepisternum covered with distinct isodiametric reticulation and some transverse wrinkles, submarginal furrow complete. Epipleuron with distinct isodiametric reticulation. Abdomen covered with distinct isodiametric reticulation, the two apical setigerous punctures on terminal segment widely separated.</paragraph>
|
|
<paragraph id="4502E25FFFE60539EFD2FAFCEE00FA06" blockId="11.[178,1085,1337,1502]" pageId="11" pageNumber="208">Legs: Protibia with distinct and complete carina and sulcus dorsally, lateral upper spine ensiform, turned distinctly ventrally and slightly laterally; movable spur ensiform, turned indistinctly ventrally. Three preapical lateral denticles distinct, obliquely truncated. First segment of protarsus longer than segments two to four together, with rows of setae dorsally. Intermediate tibia with distinct spur preapically, furnished with seta, inner side with few setae.</paragraph>
|
|
<paragraph id="4502E25FFFE60539EFD2FA2AEDC1F9E7" blockId="11.[178,1085,1516,1596]" pageId="11" pageNumber="208">Sexual differences: Male: Terminal sternite fuscus; apical half with two blunt carinae, with irregular grain-like rugae. Female: Apical half of terminal sternite with irregular to transverse rugae, free of reticulation at middle.</paragraph>
|
|
<paragraph id="4502E25FFFE1053EEFD2FF1CEC30FE9E" blockId="12.[178,1085,218,326]" pageId="12" pageNumber="209">
|
|
Male genitalia (
|
|
<figureCitation id="DD86FEDAFFE1053EEE37FF1CEEEAFF2B" box="[343,422,218,240]" captionStart="Figs 8-11" captionStartId="10.[178,218,1513,1535]" captionTargetBox="[186,1081,216,1463]" captionTargetId="figure@10.[178,1085,204,1510]" captionText="Figs 8-11: Rugiluclivina alutacea LESNE: (8) Holotype, habitus, with original mounting card and pin; (9) Original lables; (10) Aedeagus and parameres, dorsal view, specimen from Cambodia; (11) Female coxostylus, holotype." figureDoi="http://doi.org/10.5281/zenodo.3998153" httpUri="https://zenodo.org/record/3998153/files/figure.png" pageId="12" pageNumber="209">Fig. 10</figureCitation>
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): Median lobe slender, nearly straight in basal half, moderately and equally arcuate in apical part. Endophallus densely folded. Both parameres slender, moderately arcuate, somewhat distorted, dorsal one with two longer and two smaller and ventral one with two longer and one smaller nematiform setae at apex.
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</paragraph>
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<paragraph id="4502E25FFFE1053EEFD2FE92ECC8FE78" blockId="12.[178,1085,340,419]" pageId="12" pageNumber="209">
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Female genitalia (
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<figureCitation id="DD86FEDAFFE1053EEE17FE92EE85FEB1" box="[375,457,340,362]" captionStart="Figs 8-11" captionStartId="10.[178,218,1513,1535]" captionTargetBox="[186,1081,216,1463]" captionTargetId="figure@10.[178,1085,204,1510]" captionText="Figs 8-11: Rugiluclivina alutacea LESNE: (8) Holotype, habitus, with original mounting card and pin; (9) Original lables; (10) Aedeagus and parameres, dorsal view, specimen from Cambodia; (11) Female coxostylus, holotype." figureDoi="http://doi.org/10.5281/zenodo.3998153" httpUri="https://zenodo.org/record/3998153/files/figure.png" pageId="12" pageNumber="209">Fig. 11</figureCitation>
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): Coxostylus slightly sigmoid, dorso-ventrally moderately depressed, gently curved to apex, apex asetose, with one big robust and three medium sized nematiform setae in middle part, laterally with one long thin seta.
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</paragraph>
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<paragraph id="4502E25FFFE1053EEFD2FE74EC87FE13" blockId="12.[178,971,434,456]" box="[178,971,434,456]" pageId="12" pageNumber="209">
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Variation: In the
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<typeStatus id="9A065CFDFFE1053EEE00FE74EEF5FE13" box="[352,441,434,456]" pageId="12" pageNumber="209" type="holotype">holotype</typeStatus>
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the left tooth of clypeus is broadened and not acute.
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</paragraph>
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<paragraph id="4502E25FFFE1053EEFD2FE1EEE5BFDF3" blockId="12.[178,1085,472,552]" pageId="12" pageNumber="209">
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Additional material: Some specimens are not as black as the
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<typeStatus id="9A065CFDFFE1053EEC53FE1EECC2FE35" box="[819,910,472,494]" pageId="12" pageNumber="209" type="holotype">holotype</typeStatus>
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, thought slightly more piceous. Variation was also noted for the frons of the head within the irregularity of the rugae.
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</paragraph>
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<paragraph id="4502E25FFFE1053EEFD2FDFEEC18FD7E" blockId="12.[178,1085,568,678]" pageId="12" pageNumber="209">
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D i s t r i b u t i o n: According to the labels, the original description (in Latin) and the rewriting (in French) of
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<bibRefCitation id="212C9FAEFFE1053EEEDBFD90ED09FDB0" author="LESNE P" box="[443,581,597,619]" pageId="12" pageNumber="209" pagination="238 - 259" refId="ref9588" refString="LESNE P. (1896): Cicindelides et Carabides Indo-Chinois recuellis par M. Pavie. Diagnoses des especes nouvelles et d'un genre nouveau. - Bulletin du Muse um d'Histoire Naturelle de Paris 1896: 238 - 259." type="journal article" year="1896">LESNE (1896</bibRefCitation>
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,
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<bibRefCitation id="212C9FAEFFE1053EED34FD93EDC6FDB0" author="LESNE P" box="[596,650,597,619]" pageId="12" pageNumber="209" pagination="1879 - 1895" refId="ref9629" refString="LESNE P. (1904): Famille des Carabides. In: Pavie A. Mission Pavie Indo-Chine 1879 - 1895. Etudes diverses, III Recherches sur l'histoire Naturelle de Indo-Chine Orientale. - Paris: Ernst Leroux, pp. 549." type="journal article" year="1904">1904</bibRefCitation>
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), the specimens were collected in "Conchinchine" and "
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<collectingCountry id="3DAAA2CFFFE1053EEE3FFDB4EE9DFD53" box="[351,465,626,648]" name="Cambodia" pageId="12" pageNumber="209">Cambodge</collectingCountry>
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|
" without further specification. These are old terms for the South of
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|
<collectingCountry id="3DAAA2CFFFE1053EEE6FFD56EE2AFD7D" box="[271,358,656,678]" name="Vietnam" pageId="12" pageNumber="209">Vietnam</collectingCountry>
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|
(geographically) and parts of East of
|
|
<collectingCountry id="3DAAA2CFFFE1053EED86FD56EC02FD7D" box="[742,846,656,678]" name="Cambodia" pageId="12" pageNumber="209">Cambodia</collectingCountry>
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.
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</paragraph>
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<paragraph id="4502E25FFFE1053EEFD2FD70EE14FCDD" blockId="12.[178,1085,693,774]" pageId="12" pageNumber="209">
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|
R e l a t i o n s h i p s: The species
|
|
<taxonomicName id="82BD99DCFFE1053EED7FFD70EDDBFD11" box="[543,663,693,715]" class="Insecta" family="Carabidae" genus="Rugiluclivina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="alutacea">
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<emphasis id="77C93E4DFFE1053EED7FFD70EDDBFD11" box="[543,663,693,715]" italics="true" pageId="12" pageNumber="209">R. alutacea</emphasis>
|
|
</taxonomicName>
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|
and
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|
<taxonomicName id="82BD99DCFFE1053EEDB1FD70EC19FD11" authorityName="Balkenohl" authorityYear="1996" box="[721,853,693,715]" class="Insecta" family="Carabidae" genus="Rugiluclivina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="reticulata">
|
|
<emphasis id="77C93E4DFFE1053EEDB1FD70EC19FD11" box="[721,853,693,715]" italics="true" pageId="12" pageNumber="209">R. reticulata</emphasis>
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|
</taxonomicName>
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|
seem to be vicarious species, one occurring at the border to the Palearctic region and the other in the middle of the tropical belt.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |