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<document id="59C2A4316E268A0228375BE14EDDC8D7" ID-CLB-Dataset="300349" ID-DOI="10.3897/vz.74.e124752" ID-GBIF-Dataset="b637a981-80a3-4475-99c3-f213940e174d" ID-ZooBank="154CE236-EFA8-4411-834B-234A9B45A63F" ID-publisher-id="124752" URI-arpha="EA0088AE-2D74-569C-900B-1A82CA6D5784" URI-zoobank="http://zoobank.org/154CE236-EFA8-4411-834B-234A9B45A63F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1722264855187" checkinUser="pensoft" docAuthor="Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V. &amp; Das, Abhijit" docDate="2024" docId="CAD2A307ECEF5751BF30BB86E58CF3CE" docLanguage="en" docName="Vertebrate Zoology 74: 453-486" docOrigin="Vertebrate Zoology 74" docSource="https://vertebrate-zoology.arphahub.com/article/124752/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Cyrtodactylus kiphire Boruah, Narayanan, Deepak &amp; Das 2024, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="486" masterDocId="EA0088AE2D74569C900B1A82CA6D5784" masterDocTitle="Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India" masterLastPageNumber="486" masterPageNumber="453" pageNumber="453" updateTime="1722279389842" updateUser="ExternalLinkService">
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<mods:titleInfo id="F57A78F3CDC897AEC19FE21B809CD106">
<mods:title id="584B466A36FBAD3FA03D671D35E7E2D2">Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India</mods:title>
</mods:titleInfo>
<mods:name id="44FA0C094234BCC5D6DD0B220A2B9771" type="personal">
<mods:role id="2146F9FCFF01203B4A736ABD2CE7DD8F">
<mods:roleTerm id="D523D446F5775511EA194DA21A1553A9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36CB42462CD0A31DC3D2D69D94B2ACA3">Boruah, Bitupan</mods:namePart>
<mods:nameIdentifier id="00C42DF8C873CC23C0986B9C0E65268F" type="ORCID">0000-0001-8829-6069</mods:nameIdentifier>
<mods:affiliation id="61989B67EC77BB7E02A89AD2DDA3C8B8">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
</mods:name>
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<mods:namePart id="BDD6D462A545E9CC3A385795857C64EA">Narayanan, Surya</mods:namePart>
<mods:nameIdentifier id="E23904F738A9F3C499CDFC58803955FE" type="ORCID">0000-0001-9359-2815</mods:nameIdentifier>
<mods:affiliation id="D8F8AEC1AC0CEED2E8BD8B1322F75041">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India</mods:affiliation>
</mods:name>
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<mods:namePart id="D0799036D8F63874246B565F292AD180">Aravind, Neelavar Ananthram</mods:namePart>
<mods:nameIdentifier id="5314523F37805AFC863005A7A8F53AC6" type="ORCID">0000-0002-4515-8421</mods:nameIdentifier>
<mods:affiliation id="7D3E3AFD3CCDD60D00974872520B5F3C">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India &amp; Yenepoya Research Centre, Yenepoya University, Derlakatte, Mangalore, India</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="2B678D2263C291B7FDFFBCE5AF356166">Lalronunga, Samuel</mods:namePart>
<mods:nameIdentifier id="009AAEBBD7A756FC7257C81679CD08C9" type="ORCID">0000-0002-7542-4815</mods:nameIdentifier>
<mods:affiliation id="B0A698C5258818682A383B2310ED576C">Holy Child Society, Nalkata, Tripura 799263, India</mods:affiliation>
</mods:name>
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<mods:namePart id="ED8AEF50251AE060F94CBD071632A20E">Deepak, V.</mods:namePart>
<mods:nameIdentifier id="A3F2A70C8DC9FB87B973B7005EC1003C" type="ORCID">0000-0002-8826-9367</mods:nameIdentifier>
<mods:affiliation id="0D993FF04B7B91E4D7BBC95D41BE3CE6">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK &amp; School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
</mods:name>
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<mods:namePart id="B3E87DF33793F7C04046A7AC1C5E9131">Das, Abhijit</mods:namePart>
<mods:nameIdentifier id="DCAAA273D637FEA8A9283D35E89ABAB2" type="ORCID">0000-0002-5851-8457</mods:nameIdentifier>
<mods:affiliation id="EE9326D9B2D8E823EEFFED59DEE3E820">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
</mods:name>
<mods:typeOfResource id="4D7F16D3FBE4A8027BB702A3D994AE33">text</mods:typeOfResource>
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<mods:titleInfo id="203AF7E17C083E95DE344382F49B3905">
<mods:title id="0A9AD1D73C38568338B80D7D7C23E45C">Vertebrate Zoology</mods:title>
</mods:titleInfo>
<mods:part id="60D7988F62D5B9679152E9BBBF66F06D">
<mods:date id="DC2D770D0C0366BF4FAAA1D52C3A02B9">2024</mods:date>
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<mods:number id="C521024492B084985F6364C47C9D6122">2024-07-29</mods:number>
</mods:detail>
<mods:detail id="AF1A7B76C36F8CC0C9849473F7A47803" type="volume">
<mods:number id="A5F4134724ABFAF36EDF901E4FA8852B">74</mods:number>
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<mods:start id="144E4E4BF8993202608F4C3DD855CDEF">453</mods:start>
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<mods:classification id="2DD3405C2C9CBA07CE53EA1954C21182">journal article</mods:classification>
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<treatment id="CAD2A307ECEF5751BF30BB86E58CF3CE" ID-DOI="http://doi.org/10.5281/zenodo.13126867" ID-Zenodo-Dep="13126867" ID-arpha="CAD2A307-ECEF-5751-BF30-BB86E58CF3CE" ID-zoobank="https://zoobank.org/ECA0B2C2-D3C2-4C26-AA46-320CBD36AE06" LSID="urn:lsid:plazi:treatment:CAD2A307ECEF5751BF30BB86E58CF3CE" LSID-ZBK="urn:lsid:zoobank.org:act:ECA0B2C2-D3C2-4C26-AA46-320CBD36AE06" httpUri="http://treatment.plazi.org/id/CAD2A307ECEF5751BF30BB86E58CF3CE">
<subSubSection id="E8F1BA40F668079F10D89AEB7E2FA562" type="nomenclature">
<paragraph id="88804F2C137A12061DF2429B62536AAB">
<taxonomicName id="5DC2BC4BB3E5603218C05879E268ED33" authority="Boruah, Narayanan, Deepak &amp; Das" authorityName="Boruah, Narayanan, Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="6B4012FD4A8FC5A79FA0295604577F3D" italics="true">Cyrtodactylus kiphire</emphasis>
Boruah, Narayanan, Deepak &amp; Das
</taxonomicName>
<taxonomicNameLabel id="9A0F2EBB63A24E10D6B98BEC79B47179" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6942BE4B998A2636DECE47BA9D72AB2B" type="description">
<paragraph id="F0B732943FC6C40387484B4A87E85516">
<figureCitation id="765185CB1654C391B1EA949B063DDE4A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">Figure 4</figureCitation>
;
<tableCitation id="E8FA5885DBC629B868F0C921CE246C55" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">Tables 2, S 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOVDI" type="Holotype">
<paragraph id="5C5E2E60A3D53D8A678234BAAE141C4B">
<heading id="7AC59DBAEAC1F75523A46D56DAE20C4F" reason="title">
<typeStatus id="8C9D6601413CFCED5126D37D30A20D77" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="8532EB0165DCFABDF0E829B71B95A499">
<materialsCitation id="309CC2CE64088676A4C54689D1A2D5B6" collectedFrom="from a trail beside Forest Colony" collectingDate="2021-08-01" collectorName="Abhijit Das &amp; Bitupan Boruah" country="India" elevation="1300" latitude="25.8994" location="Kiphire Forest Division" longLatPrecision="6" longitude="94.7694" municipality="Kiphire District" specimenCode="WII-ADR 964" specimenCount="1" specimenCount-male="1" stateProvince="Nagaland" typeStatus="holotype">
<specimenCount id="7FDCC1ECB27028C7ED207787B20D333C" count="1" type="male">Adult male</specimenCount>
(
<specimenCode id="8AE08E5EB4A82E917B663BDD6B8CC984">WII-ADR 964</specimenCode>
; Fig.
<figureCitation id="D4AD9E555AE2E9F6B9BC46D11E88CACA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">4 A E</figureCitation>
),
<collectedFrom id="A2915236A16AEB97F87FD75389B88ECA">from a trail beside Forest Colony</collectedFrom>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2VDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[94.769400,25.899400]}">
<geoCoordinate id="D53F826442D4CE265727048678F1249E" degrees="25.8994" direction="north" orientation="latitude" precision="5" value="25.8994">25.8994 ° N</geoCoordinate>
;
<geoCoordinate id="3FA69315812F783952BDE24F08774B36" degrees="94.7694" direction="east" orientation="longitude" precision="5" value="94.7694">94.7694 ° E</geoCoordinate>
</named-content>
;
<elevation id="C30BAA20B829E9565424A94591EA0D05" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
elevation
<quantity id="285C206AEB1F32774894FF6378DE566F" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</quantity>
a. s. l.
</elevation>
; Fig.
<figureCitation id="83E1D4B3224A6E6052FED73706F76B55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
),
<location id="7BE4A1DB51A7B6BEC76D2A5E4816BF3A" LSID="urn:lsid:plazi:treatment:CAD2A307ECEF5751BF30BB86E58CF3CE:7BE4A1DB51A7B6BEC76D2A5E4816BF3A" country="India" latitude="25.8994" longLatPrecision="6" longitude="94.7694" municipality="Kiphire District" name="Kiphire Forest Division" stateProvince="Nagaland">Kiphire Forest Division</location>
,
<collectingMunicipality id="6FF1644153C380C00DD89FB2DE2417B5">Kiphire District</collectingMunicipality>
,
<collectingRegion id="0CF90506FB6E994271D00180E602EC21" country="India" name="Nagaland">Nagaland</collectingRegion>
,
<collectingCountry id="D7BA8ED0DAB73B5D853F6A6DBD7CEA67" name="India">India</collectingCountry>
; collected by
<collectorName id="8D0A51866B9AFE5D570C734228E40C3E">Abhijit Das</collectorName>
and
<collectorName id="4D09BEC4656FAA9780DF114A49AF886C">Bitupan Boruah</collectorName>
on
<date id="7FA9382C7482B5955F21AE5DC92B2296" value="2021-08-01">
<collectingDate id="D69B2CD56C0185A8683EB7398AC2B2ED" value="2021-08-01">1 August 2021</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="6B88B2D1CD1AEF31BACC5C0A360B9A1B" ID-DOI="10.3897/vz.74.e124752.figure4" ID-arpha="DABDFDAF-7299-516F-8A29-87172C0710E2" httpUri="https://binary.pensoft.net/fig/1102270" startId="F4">
<paragraph id="4EDE8C573D85E2BCB7E21132E1F81173">
<label id="415C02CA2C8E0862A800AB0F487483EB">Figure 4.</label>
</paragraph>
<paragraph id="B212032A0A129C90CE0B73CA4883CD75">
<taxonomicName id="946AB9E7A1669C8CE12E05360FCA00B6" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="F6FC7AC1F87EE5287579F45D572D62E0" italics="true">Cyrtodactylus kiphire</emphasis>
</taxonomicName>
<emphasis id="4145365D4FA0F5E65C9EF85D0BF95A39" bold="true">
<taxonomicNameLabel id="B332C00C6F05D543C18A72A83B6E93AE" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
<emphasis id="671A31BD0999EC7F424AE16FE2E91B3C" bold="true">E</emphasis>
holotype (
<specimenCode id="7460332CE31CAB5CDAB4F4CB4F98C5F0">WII-ADR 964</specimenCode>
),
<emphasis id="1CB5CDE07BD878F43321B2CCD09F853B" bold="true">A</emphasis>
dorsal view,
<emphasis id="A3803197A7227AB7247EF372609FB1AF" bold="true">B</emphasis>
ventral view,
<emphasis id="A8EBECF77AF40C420476A6460237DA73" bold="true">C</emphasis>
dorsal view of head,
<emphasis id="AB66D3E331A9D84DD90BC9FC3E68F981" bold="true">D</emphasis>
ventral view of head,
<emphasis id="AE3E2216532EB8F960D138414E1BEC04" bold="true">E</emphasis>
lateral view of head;
<emphasis id="16B6AD7F60FC7765572C415ADD43893C" bold="true">F</emphasis>
,
<emphasis id="C3826622407A71BEDAD4EFE394506CCF" bold="true">G</emphasis>
dorsal and ventral view of paratype (
<specimenCode id="90EA8A02033FFC40A148EDE0914772E1">WII-ADR 963</specimenCode>
);
<emphasis id="32F9310303BC469C3D21569F0EE0F51A" bold="true">I</emphasis>
ventral view of right manus and
<emphasis id="70C5CD71BA68615D01FD2A72C203C099" bold="true">J</emphasis>
right pes (
<specimenCode id="2670623CECCF651E68755E75AADA2ECF">WII-ADR 964</specimenCode>
);
<emphasis id="68B14F9F66EA7573A44695078BBCBCEB" bold="true">K</emphasis>
holotype in life. Scale bar: 10 mm.
<emphasis id="52EFF49FB98D23E8E4770FB97B6B1A1A" bold="true">J</emphasis>
not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EWXDI" type="Paratype">
<paragraph id="7DE0276CFD10B4EB1B1D825594CE4090">
<heading id="04792C8F8E6D98B92B65949F1924013B" reason="title">
<typeStatus id="AEA6440A4C3A6EA73EDD47FCDA895BA8" type="paratype">Paratype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="306FAFF52A4FBC9DC7F7E0C743EF4E91">
An adult male (
<specimenCode id="9F5318D0D3864E19283ABAFBAB32DE36">WII-ADR 963</specimenCode>
; Fig.
<figureCitation id="CD549BE059D39CFFFCE83EAD6C6A8961" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">4 F, G</figureCitation>
) collection locality details are same as the
<typeStatus id="6DC8E355E7A33B12C3C726A7BEEBCB9B" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAYDI" type="diagnosis">
<paragraph id="B5046B1797471F3C4F979DD9E22984C6">
<heading id="871B49C03CC4D5352A039382828BB9E9" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="A53C9EEC33C134BEDCBEB5FD282FC60B">
A medium-sized gecko (
<abbrev id="ABBRID0EGYDI" xlink_title="snout to vent length">SVL</abbrev>
up to at least
<quantity id="500D10E7197D9FEF17A1A8CEC75BC096" metricMagnitude="-2" metricUnit="m" metricValue="6.47" unit="mm" value="64.7">64.7 mm</quantity>
); 10 supralabials; 912 infralabials; 16 bluntly conical and feebly keeled dorsal tubercles; 2629 paravertebral tubercles; 35 or 36 midventral scales rows between the weak ventrolateral folds; no precloacal groove; six or seven precloacal pores in continuous series; 1215 total subdigital lamellae beneath digit IV of pes; dark brown irregular cross bars or reticulation on dorsum.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKYDI" type="description">
<paragraph id="47E0AC54D91A76A73931AF49501EAD5D">
<heading id="104516CA9DCB6705CC9281750190CFFA" reason="title">
Description of the
<typeStatus id="174456A277EAC414E5AA1F066905CC26" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="835DCB072BC230DA71F5AD0598E727CA">
<typeStatus id="96F18E6247D0EFCAB306646334D0315B" type="holotype">Holotype</typeStatus>
well preserved except single incision below left axilla ventrolaterally. Snout-vent length
<quantity id="7D77C098FFC4371856D1E2A8B910AFDC" metricMagnitude="-2" metricUnit="m" metricValue="6.39" unit="mm" value="63.9">63.9 mm</quantity>
. Head moderately large (
<abbrev id="ABBRID0EQYDI" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0EUYDI" xlink_title="snout to vent length">SVL</abbrev>
= 0.26), dorsoventrally depressed, longer than width (
<abbrev id="ABBRID0EYYDI" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0E3YDI" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.68), distinct from neck, broader at occipital region; snout tip rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat; a longitudinal furrow on dorsal surface of the snout; snout short (
<abbrev id="ABBRID0EAZDI" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0EEZDI" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.4), longer than orbit (
<abbrev id="ABBRID0EIZDI" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0EMZDI" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.7); nostril nearly rounded, opening directed posterolaterally; ear opening oval and oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous, supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at approximately anterior one third of it; rostral wide, a short groove at the middle on top; rostral connected with nasals, supranasals, internasal and first supralabials; two scale between the supranasals, larger than the rest of the granular snout scales; granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles starting from the level of posterior margin of the upper eyelids, size increases towards nape; supralabial 10 on both side, seven supralabials upto midorbit, size decreases towards angle of jaw; a series of narrow, enlarged scales above the supralabials between nostril and anterior orbital border; mental slightly wider than rostral (RW /
<collectionCode id="BF1CFBF3A1A6CD372D6F0E62C544DA45" country="Netherlands" name="Museum Wasmann">MW</collectionCode>
= 0.8), triangular, connected with first infralabials, inner postmentals; nine infralabials on both sides, size decreases towards angle of jaw; first infralabials connected with mental, second infralabial, inner and outer postmentals; inner pair of postmentals are larger than the outer postmentals; posterior margin of the inner postmentals bordered by eight granular scales of different size; two rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size decreases, elongated and narrow; rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards the throat where they become imbricate.
</paragraph>
<paragraph id="46012B2CE1FD77C5C3AC9A40F9F3FCA1">
Habitus slender (
<abbrev id="ABBRID0ESZDI" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0EWZDI" xlink_title="snout to vent length">SVL</abbrev>
= 0.2,
<abbrev id="ABBRID0E1ZDI" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0E5ZDI" xlink_title="snout to vent length">SVL</abbrev>
= 0.35), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, up to fourth segment of the tail, size increases towards posterior body and pronounced; 16 dorsal tubercles across mid dorsum; 26 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest on belly; 36 mid-ventral scales between ventrolateral fold; seven precloacal pores arranged in an inverted “
<collectionCode id="24B372FDE76E5802BE796622998EC79C" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
” shaped continuous series, followed by five unpored, large scales below it, largest at the apex.
</paragraph>
<paragraph id="0AA79FABF5A18751D2E20B81342C4F2B">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0EE1DI" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0EI1DI" xlink_title="snout to vent length">SVL</abbrev>
= 0.13,
<abbrev id="ABBRID0EM1DI" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0EQ1DI" xlink_title="snout to vent length">SVL</abbrev>
= 0.16); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 5 + 8 and 5 + 9 respectively; lamellae beneath digit IV of right and left pes (given as basal + distal) is 5 + 10 and 5 + 9 respectively; dorsal scales on forelimbs smooth and subimbricate, small and granular at elbow and proximal end of forearm; forearm scales intermixed with rounded large tubercles; dorsal scales of hindlimbs heterogeneous, intermixed with large, rounded and bluntly conical tubercles; horizontally upper half of the thigh scales are smooth, large and subimbricate, those on lower half small granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; scales on ventral side of hindlimbs smaller than those of belly, smooth, cycloid and subimbricate, but on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="0184FF468E2BDAD1150D5C34DEBD0929">
Tail regenerated (
<abbrev id="ABBRID0EW1DI" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="9C39D378FB5CE3C4864D5893AB95227D" metricMagnitude="-2" metricUnit="m" metricValue="6.8" unit="mm" value="68.0">68 mm</quantity>
), slender, gradually tapering towards tip, segments indistinct, dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; large feebly keeled scales upto fourth segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal and widest at the middle, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; two and three bluntly conical spurs on right and left side of the tail base respectively.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E11DI" type="Colouration in life">
<paragraph id="033F0219D647095F574FAF3618A441C5">
<heading id="D5F8E201F8827A4CDE8C351CAB18D48D" reason="title">Colouration in life.</heading>
</paragraph>
<paragraph id="0F044384C055053ED34F66A6F3F69E90">
Top of head intermixed with pale-yellow and pale-brown; dark-brown spots of irregular shape and size on posterior part of head; short dark-brown postorbital streak present; another streak along loreal region; supraciliary yellow; four dark-brown stripes on neck interspaced with cream colour; large spot at the middle on the anterior part of the nape; six dark-brown cross bars of irregular size and shape on dorsum between the level of axilla and groin (Fig.
<figureCitation id="5FF7F36F0F7AE4E2708F89FFE2AAF731" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">4 J</figureCitation>
); these cross bars intersected on mid dorsum and posteriorly bordered with cream coloured patches; limbs with slightly dark-brown indistinct reticulations intermixed with pale-yellow or cream coloured patches; scales on axillary region and base of forelimbs are with pinkish tinge; digits with alternating brown and pale-yellow bands; anterior half of the tail with alternating dark-brown and light-cream coloured cross bands of irregular shape and size; posterior half (regenerated part) of the tail pale-brown; ventrally head, trunk and limbs whitish; tail with brown mottling.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE2DI" type="Colouration in preservative">
<paragraph id="B968412DB2EE1679C20BCDA197A17C2A">
<heading id="AF9864EFCFE56F2EA8140CFEF8CF6CE5" reason="title">Colouration in preservative.</heading>
</paragraph>
<paragraph id="DA76161E7E1AF9F9CA1E033A0679C168">Top of head, back, limbs and tail pale-greyish-brown with dark-brown markings; upper eyelids grey; dorsal marking pattern as it was in life; and ventrally whitish.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ2DI" type="Morphological variation">
<paragraph id="4E56385D83D494FD5966B6B8C2A1736D">
<heading id="1F20E129B20877D4A63BB3AB015259C0" reason="title">Morphological variation.</heading>
</paragraph>
<paragraph id="983820DF28DC4CCAC87E389ADA0B9A89">
Detailed morphometric variations are given in Table S 2. Apart from those, the dorsal colour and marking pattern of the
<typeStatus id="21717126980873B2BB15A1E688889F63" type="paratype">paratype</typeStatus>
varied from that of
<typeStatus id="D106DB6D86377832CF441870A8E2FC60" type="holotype">holotype</typeStatus>
.
<typeStatus id="1CD925445846CC5D0DDFC3DCE0EACA18" type="paratype">Paratype</typeStatus>
has two post orbital streaks on each side, upper one shorter than the lower one; no other spots on occipital region as in the
<typeStatus id="1BEF2C625ADD472960E2B21F4048190E" type="holotype">holotype</typeStatus>
; dorsal cross bands somewhat reticulated (Fig.
<figureCitation id="49198000418951E600801ED5E07F5DEC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">4 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ET2DI" type="comparisons">
<paragraph id="D9661BFF7CD371A9469A1CD9D3A5C963">
<heading id="F2DC8F5D7E019F164A3A9694B83BB861" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="D43B67C1DB59215E0996E627D818DCA1">
<taxonomicName id="A9ED2B082567CBD60D77A782D87D5AC7" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="73340120157914E56D4AE01EC5F84F13" italics="true">Cyrtodactylus kiphire</emphasis>
</taxonomicName>
<emphasis id="9E897CE7108631970E02A2D72AFD611C" bold="true">
<taxonomicNameLabel id="65F0A5842EE375D39362B20BCFE0319C" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="42D36C484B0CBCAA593CB40F262E40F8" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aaronbaueri">
<emphasis id="830D9F2D057CFD62209ED90F92EFD5A0" italics="true">C. aaronbaueri</emphasis>
</taxonomicName>
by having larger body size in male,
<abbrev id="ABBRID0ER3DI" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="68385ED4FD2BC53CF38270D9CD5ECDC2" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0EV3DI" xlink_title="snout to vent length">
<collectionCode id="4F4368FA770BE34E4508298FEB2C8ED2">SVL</collectionCode>
</abbrev>
<quantity id="35781095592196AD824B57C0EC1875C5" metricMagnitude="-2" metricUnit="m" metricValue="5.81" metricValueMax="6.2" metricValueMin="5.42" unit="mm" value="58.1" valueMax="62.0" valueMin="54.2">54.262 mm</quantity>
in male), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EZ3DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0E43DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="429F5E63A38F288D86E5E154FF3B72C1">DTR</collectionCode>
</abbrev>
2228), fewer paravertebral tubercles, 2629 (vs.
<abbrev id="ABBRID0EB4DI" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="8D7AC1565ED447B0EF663050DD53F875">PVT</collectionCode>
</abbrev>
3639); differs from
<taxonomicName id="B6F5E5DC95AAFBE6586348E96689BD06" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aunglini">
<emphasis id="C21E0954E42CCD636B3579F23D142CF2" italics="true">C. aunglini</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EQ4DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EU4DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="37F6966BD7E55C8BE41E117FBC4C7615">DTR</collectionCode>
</abbrev>
2126), fewer precloacal pores,
<abbrev id="ABBRID0EY4DI" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0E34DI" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
1213), fewer mid-ventral scale rows,
<abbrev id="ABBRID0EA5DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="E6F66EEFAD126F98FB4D4D47B7C162FB">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EE5DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="D04C97438098684BD09BD443DC338797">MVSR</collectionCode>
</abbrev>
4149), fewer paravertebral tubercles,
<abbrev id="ABBRID0EI5DI" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3645); differs from
<taxonomicName id="548E4F02D9EA3BA377B8557BBA84EC70" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="bengkhuaiai">
<emphasis id="B679918FE006553C6AC0D6837513A3F9" italics="true">C. bengkhuaiai</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EX5DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0E25DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="024797FCC8F95CBBEE18FEB71894BB65">DTR</collectionCode>
</abbrev>
2226), fewer paravertebral tubercles, 2629 (vs.
<abbrev id="ABBRID0E65DI" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="7BF268A492949AE148969AB3EF45FE6E">PVT</collectionCode>
</abbrev>
3541), fewer mid-ventral scale rows,
<abbrev id="ABBRID0ED6DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="B6EDCA3DD9116075BCF70684E50F1219">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EH6DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="998672D351E6EDF4CB3201E4CF179F0D">MVSR</collectionCode>
</abbrev>
3742); differs from
<taxonomicName id="68E2350FA4A0FA9722BE55F84412793A" authorityName="Bauer" authorityYear="2002" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="brevidactylus">
<emphasis id="C64723241CF178ACC3950FA813524218" italics="true">C. brevidactylus</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EW6DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0E16DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="5AC732E91A16E1FDC1EFF613705CF15B">DTR</collectionCode>
</abbrev>
27), fewer mid-ventral scale rows,
<abbrev id="ABBRID0E56DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="75C7B710CC09C77EBB8CFAD473892B4F">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EDAAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="EB5E2154CDDF917C5E0B5587BDF04FE9">MVSR</collectionCode>
</abbrev>
45), fewer precloacal pores,
<abbrev id="ABBRID0EHAAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0ELAAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
8); differs from
<taxonomicName id="9161941C4496C993F85F69533F023CF2" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chrysopylos">
<emphasis id="DA38365F3D44E564CF9E5A49CBE7B5C5" italics="true">C. chrysopylos</emphasis>
</taxonomicName>
by fewer precloacal pores,
<abbrev id="ABBRID0E1AAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0E5AAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
813), mid ventral scales,
<abbrev id="ABBRID0ECBAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="5501FDC09ED8C70A65F67F64F0E6384F">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EGBAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="79DA4C6F441E83D04D8F774749596D4A">MVSR</collectionCode>
</abbrev>
3755), dorsal tubercle rows,
<abbrev id="ABBRID0EKBAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
2629 (vs.
<abbrev id="ABBRID0EOBAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="5000F93A3AECB76A67A961BF16F4C09C">DTR</collectionCode>
</abbrev>
3035); differs from
<taxonomicName id="92404894FB68DC8A56155096D86F49DF" authorityName="Liu &amp; Rao" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="dianxiensis">
<emphasis id="93E6B41D6E5D0E74B08AAEABE13F0CC5" italics="true">C. dianxiensis</emphasis>
</taxonomicName>
by smaller body size,
<abbrev id="ABBRID0E4BAK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="B1DE3820F1B9218CED1AB0D0C0BE0553" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0EBCAK" xlink_title="snout to vent length">
<collectionCode id="F7E51A5ABFCF905FF72B2FC0E70E43EB">SVL</collectionCode>
</abbrev>
<quantity id="1038C09EB9A5B2B0DA22DEC6CA3F1327" metricMagnitude="-2" metricUnit="m" metricValue="7.685" metricValueMax="7.99" metricValueMin="7.38" unit="mm" value="76.85" valueMax="79.9" valueMin="73.8">73.879.9 mm</quantity>
), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EFCAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EJCAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="3C08742B58B11269EEAC323C07CAD33A">DTR</collectionCode>
</abbrev>
18 or 19), 35 or 36 mid ventral scales (vs.
<abbrev id="ABBRID0ENCAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="214E497013A09441C2E21114ADD15CDA">MVSR</collectionCode>
</abbrev>
3741), 26 or 29 paravertebral tubercles (vs.
<abbrev id="ABBRID0ERCAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="173C97DAAD89C90C552B40CFCC05693A">PVT</collectionCode>
</abbrev>
31 or 32); differs from
<taxonomicName id="1873EFE2845B0FAAC9CE9DC05DD1F406" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="gansi">
<emphasis id="0844A91513E4A4C4B0E28AD3678654AA" italics="true">C. gansi</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EADAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EEDAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="D7830C7F101D3A262172DC38B303F931">DTR</collectionCode>
</abbrev>
2025), having fewer precloacal pores,
<abbrev id="ABBRID0EIDAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EMDAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
1629); differs from
<taxonomicName id="F4ABB2165D18AA56A1EFA95CB3FF99ED" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="jaintiaensis">
<emphasis id="C8EB78F97D42605832E655F534B4FEF2" italics="true">C. jaintiaensis</emphasis>
</taxonomicName>
by smaller body size in male,
<abbrev id="ABBRID0E2DAK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="CA9CD25A3ABB4E0E3B6E1EB85F7BC56E" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0E6DAK" xlink_title="snout to vent length">
<collectionCode id="B64090FBE7C39B7D0F957878EB8CB6DA">SVL</collectionCode>
</abbrev>
<quantity id="F6E830D78CC87640E94D37ABD2BF76FC" metricMagnitude="-2" metricUnit="m" metricValue="8.765" metricValueMax="8.83" metricValueMin="8.7" unit="mm" value="87.65" valueMax="88.3" valueMin="87.0">8788.3 mm</quantity>
), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EDEAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EHEAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="B3C129AB4F1CDAA5A26EF1B7B9A41CE7">DTR</collectionCode>
</abbrev>
1920), fewer mid-ventral scale rows,
<abbrev id="ABBRID0ELEAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="5D85E15E4FA531D3CA87631BC9842E10">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EPEAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="4A69F92A686DD119E4343E3F366A4048">MVSR</collectionCode>
</abbrev>
4042), fewer precloacal pores,
<abbrev id="ABBRID0ETEAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EXEAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
1112); differs from
<taxonomicName id="94D182FF5864FC62F56BC3C73A72459F" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="lungleiensis">
<emphasis id="6D846123DC1B7EC197E5F1D630E5697F" italics="true">C. lungleiensis</emphasis>
</taxonomicName>
by smaller body size in male,
<abbrev id="ABBRID0EGFAK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="02601F2866564E0EA3B313C09BA19325" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0EKFAK" xlink_title="snout to vent length">
<collectionCode id="3D94CD6B8D4B99EF8610ED7EBAE546B4">SVL</collectionCode>
</abbrev>
<quantity id="7421E4649754BF1A7C650DA8C8295817" metricMagnitude="-2" metricUnit="m" metricValue="6.654999999999999" metricValueMax="6.81" metricValueMin="6.5" unit="mm" value="66.55" valueMax="68.1" valueMin="65.0">6568.1 mm</quantity>
), higher precloacal pores,
<abbrev id="ABBRID0EOFAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0ESFAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
35), fewer mid-ventral scale rows,
<abbrev id="ABBRID0EWFAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="CF0E0358701FA3E2CC99CBFEE5E45D10">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0E1FAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="C71E1D5B14DF1187A21FE42FD10D7798">MVSR</collectionCode>
</abbrev>
3743), fewer dorsal tubercle rows,
<abbrev id="ABBRID0E5FAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs. 2428), fewer paravertebral scales,
<abbrev id="ABBRID0ECGAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3240), fewer subdigital lamellae beneath toe IV,
<abbrev id="ABBRID0EGGAK" xlink_title="count for the apical series, comprising lamellae distal to the digital inflection but not including the ventral claw sheath and the small non lamellar scales between the basal and apical lamellae series, abbreviations for lamellae are as follows: on the pes">TIVLam</abbrev>
1214 (vs.
<abbrev id="ABBRID0ELGAK" xlink_title="count for the apical series, comprising lamellae distal to the digital inflection but not including the ventral claw sheath and the small non lamellar scales between the basal and apical lamellae series, abbreviations for lamellae are as follows: on the pes">TIVLam</abbrev>
1618); differs from
<taxonomicName id="13DA08BF8E9DFD4FD126CAAB68B4C26A" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="montanus">
<emphasis id="265D7E4702428457DE671AB29F227BDE" italics="true">C. montanus</emphasis>
</taxonomicName>
by larger body size in male,
<abbrev id="ABBRID0E2GAK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="3E7383B11425DD0D4A6CBFD1717BD39E" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0E6GAK" xlink_title="snout to vent length">
<collectionCode id="FD8FC39311734741BFB7FE268047B011">SVL</collectionCode>
</abbrev>
<quantity id="EAC8EA2B54C6FDC93D6014FE548969B8" metricMagnitude="-2" metricUnit="m" metricValue="5.43" metricValueMax="5.5" metricValueMin="5.36" unit="mm" value="54.3" valueMax="55.0" valueMin="53.6">53.655 mm</quantity>
), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EDHAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EHHAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="6C6BB8BC88B80862944785D142330047">DTR</collectionCode>
</abbrev>
2123), fewer precloacal pores,
<abbrev id="ABBRID0ELHAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EPHAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
810); differs from
<taxonomicName id="E24F666814C7214B113447567990D854" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="myaleiktaung">
<emphasis id="23124DDA4648AE5273EA915A1B786EC7" italics="true">C. myaleiktaung</emphasis>
</taxonomicName>
by fewer mid-ventral scale rows,
<abbrev id="ABBRID0E5HAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="2557C43D7F6C169208CCA135A7812409">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0ECIAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="873686AF51BF94FECE71DC9964424FAF">MVSR</collectionCode>
</abbrev>
57); differs from
<taxonomicName id="A5F378F3E95D1EB37EEFBC3218809EE7" authorityName="Mahony &amp; Kamei" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namtiram">
<emphasis id="4FADDC8A8F4511001B8DF680DC0DB4A0" italics="true">C. namtiram</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ERIAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EVIAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="5428C2C95167BE524094031A01233C3D">DTR</collectionCode>
</abbrev>
21), fewer number of paravertebral scales,
<abbrev id="ABBRID0EZIAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs.
<abbrev id="ABBRID0E4IAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="12818F958C3BCC912BA79C9AFE04F8CB">PVT</collectionCode>
</abbrev>
33), fewer precloacal pores,
<abbrev id="ABBRID0EBJAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EFJAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
12); differs from
<taxonomicName id="C0BB5068042CF66443927199D45495A6" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngopensis">
<emphasis id="C48BD89EE472DD2DD14A845C6714207A" italics="true">C. ngopensis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EUJAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs. 1920), fewer paravertebral tubercles,
<abbrev id="ABBRID0EYJAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3236); differs from
<taxonomicName id="DC59B8CC6952F90B9C1DDD5E5F830A4A" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siahaensis">
<emphasis id="A21A0AA325BB937D425B2AFE2C4F2510" italics="true">C. siahaensis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EHKAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0ELKAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="02B3862A1AFE6AA75D6C45C7CAB563DC">DTR</collectionCode>
</abbrev>
2224), fewer paravertebral tubercles, 2629 (
<abbrev id="ABBRID0EPKAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="B4FD861B8270962FE02ACB9DA2EE90C1">PVT</collectionCode>
</abbrev>
3639); differs from
<taxonomicName id="20285FB448434EEFB1536D8C344EA8B9" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="vairengtensis">
<emphasis id="7A828F534FE7867BA35C5895456ACFCF" italics="true">C. vairengtensis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0E5KAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs. 2223), fewer paravertebral tubercles,
<abbrev id="ABBRID0ECLAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3439), fewer precloacal pores in male,
<abbrev id="ABBRID0EGLAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EKLAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
911). The new species is morphologically close to
<taxonomicName id="C009C4A6459C6C7CD534EFFA47E288E2" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="9D2944D6A6B129E15FDABCBD0EAC25F5" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
; however, the new species differs from it by fewer paravertebral tubercles,
<abbrev id="ABBRID0EZLAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3537). We could not compare with male specimens as data for
<taxonomicName id="7C3115DFAE2094D186DC431946823B54" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="555283909F04F22D8469249D1AF9C3D0" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
is not available. Morphological differences with other members of
<taxonomicName id="E76E555048D38A4BF8FFA52D0EF42141" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="AB826410026D8845148E391049619862" italics="true">khasiensis</emphasis>
</taxonomicName>
group is presented in Table
<tableCitation id="65D36B8A1573A4A7E1DF28E6849D6876" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWMAK" type="Sequence divergence">
<paragraph id="00D27784C02B5DBD4640554A5340E1B5">
<heading id="50B96D17333767F0EEBBAB991C145818" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="3DAE78EEC9C60D3588C9C78880764ED5">
<taxonomicName id="0145A8271EEDA2740AE1F61A152A733A" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="8DB3010276D460D35CF3DB37E1429240" italics="true">Cyrtodactylus kiphire</emphasis>
</taxonomicName>
<emphasis id="C225477C71BD1D3DAB2543C3B0B03640" bold="true">
<taxonomicNameLabel id="EF42E0BFCC3A9E7B27A66FC520E16001" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a high genetic divergence of 12 % from its closely related
<taxonomicName id="F4336A89CCC1B7D5EA647EFD868118EB" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="7AEE5DFE91D749032B0F98E46F7678D4" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
, 16.3 % from
<taxonomicName id="946E5542B83C957C7B500704F20CE095" authorityName="Liu &amp; Rao" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="dianxiensis">
<emphasis id="F30A858F5375B637E1469BD48BE821D1" italics="true">C. dianxiensis</emphasis>
</taxonomicName>
, 16.9 % from
<taxonomicName id="3552A44864D633235EAA44B5960E1004" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="gansi">
<emphasis id="2A53D95B121DF67D4A2DB7C9C317ECC2" italics="true">C. gansi</emphasis>
</taxonomicName>
and 17.9 % from
<taxonomicName id="B7B217A50470968FEE9AD27845ADCF29" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="jaintiaensis">
<emphasis id="942A55421EED5AFEB2CCA950C7836404" italics="true">C. jaintiaensis</emphasis>
</taxonomicName>
. With other members of the clade,
<taxonomicName id="843A4541148A462981061EFA886207B5" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="B96878DC57BFB2FF18C14194F8C2C0FD" italics="true">C. kiphire</emphasis>
</taxonomicName>
<emphasis id="A422B104D0B147FF893F8EDDE3C86D5E" bold="true">
<taxonomicNameLabel id="E02E684DE170BE19661400FE32A79188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a genetic divergence of 16.4 % and 22.5 % in the ND 2 gene (Table S 3).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECPAK" type="etymology">
<paragraph id="6440FBEF1773FC21C532113FF3397EC3">
<heading id="AC90CCA57EDF93F4B04A24DC7B4414D7" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="BBD8AA84E78FC59E9E9EB53AAFA48068">
The specific epithet is derived from Kiphire, a District of
<collectingRegion id="29DB84EBDCAE1128B03F310817038E58" country="India" name="Nagaland">Nagaland</collectingRegion>
from where the type series of this species were collected.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHPAK" type="Suggested common name">
<paragraph id="1C913A888AD0ED31825423228A16D270">
<heading id="DFB9630D18077DBA15AFD3BB46546018" reason="title">Suggested common name.</heading>
</paragraph>
<paragraph id="59934BC653AC2E24692DC90BE886FC1B">Kiphire bent-toed gecko.</paragraph>
</subSubSection>
<subSubSection id="SECID0EMPAK" type="distribution">
<paragraph id="E7A508F09E1F52E41B9CEAC706C52802">
<heading id="467689291B31F11CF6474D0DB24C9688" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="DF4706F50A302C0C7F4AE22AF8B96C48">
We recorded
<taxonomicName id="325E165EBFD0FC824983E41BC4A2400A" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="300A91A478B7F1FE1E3AA9A8CD22AB19" italics="true">Cyrtodactylus kiphire</emphasis>
</taxonomicName>
<emphasis id="0701829309F9FB9E9C5FD2A447246C2A" bold="true">
<taxonomicNameLabel id="A1309266CAB55C67D3D47535B6C784C9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Kiphire forest colony, Kiphire District,
<collectingRegion id="1A731610357C0B82ACAE4CF190D61671" country="India" name="Nagaland">Nagaland</collectingRegion>
,
<collectingCountry id="4089BF9DF2F4ABABD1AF4DCE69C038EE" name="India">India</collectingCountry>
. The area is characterised as subtropical forest with regenerating jhum forest dominated by
<taxonomicName id="B4671D4638210AFE089E0EBD0B371B9F" authority="(Roxb. ex Lindl.) A. DC." authorityName="A. DC." baseAuthorityName="Lindl." class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="8C0717B0D2AB95845AF66A4BF87A828F" italics="true">Castanopsis indica</emphasis>
(Roxb. ex Lindl.) A. DC.
</taxonomicName>
,
<taxonomicName id="338DF10302A2896AE567D4BAE22FF9EB" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="F19DA128BB4D8C3E4E526698ABA96C9D" italics="true">Quercus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="C8E9C5A714998638011E0EB16503A3B1" authority="Wall. ex Roxb." authorityName="Roxb." class="Magnoliopsida" family="Iteaceae" genus="Itea" kingdom="Plantae" order="Saxifragales" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis id="AEBA572ED33B69487D97D8211A139E6E" italics="true">Itea macrophylla</emphasis>
Wall. ex Roxb.
</taxonomicName>
,
<taxonomicName id="ECB461268F53803A7C372CC05CAF258B" authority="(Osbeck) Merr" authorityName="Merr" baseAuthorityName="Osbeck" class="Magnoliopsida" family="Fabaceae" genus="Albizia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis id="57DC8F1543B2DFA132A4A202844401E3" italics="true">Albizia chinensis</emphasis>
(Osbeck) Merr
</taxonomicName>
.. The
<typeStatus id="496B55122EA9BC41F0F20B5A364D0DE7" type="holotype">holotype</typeStatus>
was collected from on a forest trail shrub (approximately 2.0 m from the ground) at approximately 23: 00 hrs. The
<typeStatus id="DEB7DC6D6974F3F8A01F48A53DFD7342" type="paratype">paratype</typeStatus>
was collected from a rocky slope along the edge of a stream.
</paragraph>
</subSubSection>
</treatment>
</document>