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<document id="CC9599EEE96D3DDA7F1E4882F5DCABEE" ID-CLB-Dataset="51211" ID-DOI="10.11646/zootaxa.4061.4.5" ID-GBIF-Dataset="a54cdc47-68ee-4c8c-96eb-c146baf054ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="256219" ID-ZooBank="3B7CB094-DFA9-4C1B-A54F-3C9E4BDD40A3" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459679212016" checkinUser="plazi" docAuthor="Ito, Tomiko" docDate="2016" docId="C213FF65FFF5F647519FFE07FD60FADC" docLanguage="en" docName="zootaxa.4061.4.5.pdf" docOrigin="Zootaxa 4061 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidostoma coreanum Kumanski &amp; Weaver 1992" docType="treatment" docVersion="9" lastPageNumber="403" masterDocId="3E2A871DFFF4F6415108FFB0FFD9FF89" masterDocTitle="The Lepidostoma coreanum Species Complex (Trichoptera, Lepidostomatidae) in the Asian Far East" masterLastPageNumber="417" masterPageNumber="397" pageNumber="398" updateTime="1698206791494" updateUser="ExternalLinkService">
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<mods:title id="A08D4F473F78DC6C5172F3D98EE6731B">The Lepidostoma coreanum Species Complex (Trichoptera, Lepidostomatidae) in the Asian Far East</mods:title>
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<mods:namePart id="459566D764CCBC4167609F51A79C8E46">Ito, Tomiko</mods:namePart>
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<treatment id="C213FF65FFF5F647519FFE07FD60FADC" ID-DOI="http://doi.org/10.5281/zenodo.6055839" ID-GBIF-Taxon="119332923" ID-Zenodo-Dep="6055839" LSID="urn:lsid:plazi:treatment:C213FF65FFF5F647519FFE07FD60FADC" httpUri="http://treatment.plazi.org/id/C213FF65FFF5F647519FFE07FD60FADC" lastPageId="6" lastPageNumber="403" pageId="1" pageNumber="398">
<subSubSection id="02A01DF8FFF5F640519FFE07FEE2FE7A" pageId="1" pageNumber="398" type="nomenclature">
<paragraph id="4A054E73FFF5F640519FFE07FCC0FE5B" blockId="1.[151,793,439,499]" box="[151,793,439,466]" pageId="1" pageNumber="398">
<heading id="114DF91FFFF5F640519FFE07FCC0FE5B" bold="true" box="[151,793,439,466]" fontSize="11" level="1" pageId="1" pageNumber="398" reason="1">
<taxonomicName id="8DBA35F0FFF5F640519FFE07FCC0FE5B" authority="Kumanski &amp; Weaver 1992" authorityName="Kumanski &amp; Weaver" authorityYear="1992" box="[151,793,439,466]" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="coreanum">
<emphasis id="78CE9261FFF5F640519FFE07FCC0FE5B" bold="true" box="[151,793,439,466]" pageId="1" pageNumber="398">
<emphasis id="78CE9261FFF5F640519FFE07FE6AFE58" bold="true" box="[151,435,439,465]" italics="true" pageId="1" pageNumber="398">Lepidostoma coreanum</emphasis>
(
<bibRefCitation id="2E2B3382FFF5F64050CAFE08FCC8FE5B" author="Kumanski" box="[450,785,440,466]" pageId="1" pageNumber="417" refString="Kumanski, K. &amp; Weaver, J. S. III (1992) Studies on the fauna of Trichoptera (Insecta) of Korea. IV. The family Lepidostomatidae. Aquatic Insects, 14, 153 - 168. http: // dx. doi. org / 10.1080 / 01650429209361478" type="journal article" year="1992">Kumanski &amp; Weaver 1992</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4A054E73FFF5F640519FFE6AFEE2FE7A" blockId="1.[151,793,439,499]" box="[151,315,474,499]" pageId="1" pageNumber="398">
(
<figureCitation id="D28152F6FFF5F6405197FE6AFED2FE7A" box="[159,267,474,499]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2, 3" captionStart-2="FIGURES 4, 5" captionStart-3="FIGURE 6" captionStartId-0="2.[151,250,1819,1841]" captionStartId-1="3.[151,264,1749,1772]" captionStartId-2="4.[151,264,1822,1845]" captionStartId-3="5.[151,250,1877,1899]" captionTargetBox-0="[176,1409,216,1782]" captionTargetBox-1="[175,1400,213,1704]" captionTargetBox-2="[259,1417,226,1777]" captionTargetBox-3="[234,1370,195,1876]" captionTargetId-0="figure@2.[151,1436,193,1812]" captionTargetId-1="figure@3.[151,1436,193,1728]" captionTargetId-2="figure@4.[151,1436,193,1801]" captionTargetId-3="figure@5.[232,1370,193,1907]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Lepidostoma coreanum, male, specimens from Primorye. A, head, right scape and maxillary palp, right lateral; B, right scape, ventral; C, right forewing, dorsal; D, right hind wing, dorsal; E, genitalia, left lateral; F, same, dorsal; G, same, ventral. Abbreviations. Body: IX = abdominal segment IX, X = abdominal segment X; ma, la and va = mesal, lateral and ventral arms of segment X, respectively; ia = inferior appendage; ih, dh and maia = inner hook, dorsal hook and main article of an inferior appendage, respectively; ph = phallic apparatus. Wings: 1 – 2 = apical forks I and II; d = discoidal cell; th = thyridial cell; ar = arculus; C = costa; Sc = subcosta; R = radius; Ma = anterior media; Mp = posterior media; Cu = cubitus; A = anal vein." captionText-1="FIGURES 2, 3. Lepidostoma coreanum, male, variation. 2, specimen from Shizuoka, Amagi-yugashima-cho; 3, specimen from Wakayama. A, head, left scape and maxillary palp, left lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral; F, apex of main article of right inferior appendage, ventral, enlarged." captionText-2="FIGURES 4, 5. Lepidostoma coreanum, male, variation. 4, specimen from Ehime; 5, specimen from Kochi. A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral." captionText-3="FIGURE 6. Lepidostoma coreanum, female, larva, and case, specimens from Primorye. A – F, Female: A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, abdominal segments VII – X, left lateral; E, same, ventral, subgenital plate omitted; F, subgenital plate, ventral, posterior edge up. G – I, Larva: G, habitus, left lateral; H, right half of thorax, dorsal. I, Case: case of larva, left lateral. G – H, from Ito (1998). Abbreviations: VII, VIII, IX, and X = abdominal segments VII – X, respectively; lp = lateral plate; sp = subgenital plate; wing labels, as in Fig. 1." httpUri-0="https://zenodo.org/record/256220/files/figure.png" httpUri-1="https://zenodo.org/record/256221/files/figure.png" httpUri-2="https://zenodo.org/record/256222/files/figure.png" httpUri-3="https://zenodo.org/record/256223/files/figure.png" pageId="1" pageNumber="398">Figs. 1–6</figureCitation>
,
<figureCitation id="D28152F6FFF5F640501EFE6BFEEAFE7A" box="[278,307,475,499]" captionStart="FIGURE 21" captionStartId="19.[151,250,1367,1389]" captionTargetBox="[152,1414,187,1366]" captionTargetId="figure@19.[151,1436,171,1368]" captionTargetPageId="19" captionText="FIGURE 21. Distribution of the 5 species of the Lepidostoma coreanum Species Complex of the genus Lepidostoma. The plot in North Korea was referred from Kumanski &amp; Weaver (1992)." httpUri="https://zenodo.org/record/256233/files/figure.png" pageId="1" pageNumber="398">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="02A01DF8FFF5F640519FFD92FC11FD1D" pageId="1" pageNumber="398" type="reference_group">
<paragraph id="4A054E73FFF5F640519FFD92FC66FDFF" blockId="1.[151,1435,546,661]" pageId="1" pageNumber="398">
<taxonomicName id="8DBA35F0FFF5F640519FFD92FD40FDB1" ID-CoL="7VJGN" authority="Kumanski &amp; Weaver 1992" authorityName="Kumanski &amp; Weaver" authorityYear="1992" box="[151,665,546,568]" class="Insecta" family="Lepidostomatidae" genus="Indocrunoecia" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="coreana">
<emphasis id="78CE9261FFF5F640519FFD92FE5AFDB1" box="[151,387,546,568]" italics="true" pageId="1" pageNumber="398">Indocrunoecia coreana</emphasis>
<bibRefCitation id="2E2B3382FFF5F6405081FD92FD40FDB1" author="Kumanski" box="[393,665,546,568]" pageId="1" pageNumber="417" refString="Kumanski, K. &amp; Weaver, J. S. III (1992) Studies on the fauna of Trichoptera (Insecta) of Korea. IV. The family Lepidostomatidae. Aquatic Insects, 14, 153 - 168. http: // dx. doi. org / 10.1080 / 01650429209361478" type="journal article" year="1992">Kumanski &amp; Weaver 1992</bibRefCitation>
</taxonomicName>
, 163–165, male, female,
<collectingCountry id="32AD0EE3FFF5F6405290FD92FBCEFDB1" box="[920,1047,546,568]" name="North Korea" pageId="1" pageNumber="398">North Korea</collectingCountry>
(Province Pyongan Pukudo).
<taxonomicName id="8DBA35F0FFF5F640519FFDF1FE39FDDE" ID-CoL="7T7BW" authority="Ito 1998" authorityName="Ito" authorityYear="1998" box="[151,480,577,599]" class="Insecta" family="Lepidostomatidae" genus="Dinarthrum" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="coreanum">
<emphasis id="78CE9261FFF5F640519FFDF1FEA3FDDE" box="[151,378,577,599]" italics="true" pageId="1" pageNumber="398">Dinarthrum coreanum</emphasis>
:
<bibRefCitation id="2E2B3382FFF5F6405080FDF2FE39FDDE" author="Ito" box="[392,480,577,599]" pageId="1" pageNumber="417" refString="Ito, T. (1998) Description of a Far Eastern lepidostomatid caddisfly, Dinarthrum coreanum (Kumanski et Weaver, 1992) (Trichoptera). Entomological Science, 1, 585 - 588." type="journal article" year="1998">Ito 1998</bibRefCitation>
</taxonomicName>
, male, female, larva, case, Russian Far East (Primorye, Khasanski),
<collectingCountry id="32AD0EE3FFF5F6405590FDF2FB0BFDDE" box="[1176,1234,578,599]" name="Japan" pageId="1" pageNumber="398">Japan</collectingCountry>
(Honshu: Shizuoka (Amagi-yugashima-cho), Wakayama);
<bibRefCitation id="2E2B3382FFF5F640535AFDD1FD3BFDFF" author="Ito" box="[594,738,608,630]" pageId="1" pageNumber="417" refString="Ito, T., Yamamoto, E., Doi, M. &amp; Ohkawa, A. (2002) The family Lepidostomatidae and the genus Palaeagapetus of the family Hydroptilidae in Shikoku, western Japan (Trichoptera). Hyogo Freshwater Biology, 54, 21 - 40. [in Japanese with English abstract]" type="journal article" year="2002">
Ito
<emphasis id="78CE9261FFF5F640537CFDD1FD7EFDFF" box="[628,679,608,630]" italics="true" pageId="1" pageNumber="398">et al.</emphasis>
2002
</bibRefCitation>
, 23,
<collectingCountry id="32AD0EE3FFF5F640521BFDD1FC94FDFF" box="[787,845,609,630]" name="Japan" pageId="1" pageNumber="398">Japan</collectingCountry>
(Shikoku).
</paragraph>
<paragraph id="4A054E73FFF5F640519FFDCEFC11FD1D" blockId="1.[151,1435,546,661]" box="[151,968,638,661]" pageId="1" pageNumber="398">
<taxonomicName id="8DBA35F0FFF5F640519FFDCEFDC0FD1D" authority="Weaver 2002" authorityName="Weaver" authorityYear="2002" box="[151,537,638,661]" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="coreanum">
<emphasis id="78CE9261FFF5F640519FFDCEFE5DFD1C" box="[151,388,638,661]" italics="true" pageId="1" pageNumber="398">Lepidostoma coreanum</emphasis>
:
<bibRefCitation id="2E2B3382FFF5F640509AFDCFFDC0FD1D" author="Weaver" box="[402,537,638,660]" pageId="1" pageNumber="417" refString="Weaver, J. S. III (2002) A synonymy of the caddisfly genus Lepidostoma Rumbur (Trichoptera: Lepidostomatidae), including a species checklist. Tijdschrift voor Entomologie, 145, 173 - 192." type="journal article" year="2002">Weaver 2002</bibRefCitation>
</taxonomicName>
, 180;
<bibRefCitation id="2E2B3382FFF5F640535FFDCFFD76FD1D" author="Ito" box="[599,687,638,660]" pageId="1" pageNumber="417" refString="Ito, T. (2005) Lepidostomatidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations. Tokai University Press, Kanagawa, pp. 504 - 516. [in Japanese]" type="book chapter" year="2005">Ito 2005</bibRefCitation>
, 504, 507, 512, male, larva.
</paragraph>
</subSubSection>
<subSubSection id="02A01DF8FFF5F647519FFD71FBA0FD66" lastPageId="6" lastPageNumber="403" pageId="1" pageNumber="398" type="description">
<paragraph id="4A054E73FFF5F640519FFD71FE81FCE3" blockId="1.[151,1437,705,1990]" pageId="1" pageNumber="398">
<emphasis id="78CE9261FFF5F640519FFD71FF0DFD53" bold="true" box="[151,212,705,730]" pageId="1" pageNumber="398">Male</emphasis>
(
<figureCitation id="D28152F6FFF5F64051ECFD71FE88FD53" box="[228,337,705,730]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2, 3" captionStart-2="FIGURES 4, 5" captionStartId-0="2.[151,250,1819,1841]" captionStartId-1="3.[151,264,1749,1772]" captionStartId-2="4.[151,264,1822,1845]" captionTargetBox-0="[176,1409,216,1782]" captionTargetBox-1="[175,1400,213,1704]" captionTargetBox-2="[259,1417,226,1777]" captionTargetId-0="figure@2.[151,1436,193,1812]" captionTargetId-1="figure@3.[151,1436,193,1728]" captionTargetId-2="figure@4.[151,1436,193,1801]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Lepidostoma coreanum, male, specimens from Primorye. A, head, right scape and maxillary palp, right lateral; B, right scape, ventral; C, right forewing, dorsal; D, right hind wing, dorsal; E, genitalia, left lateral; F, same, dorsal; G, same, ventral. Abbreviations. Body: IX = abdominal segment IX, X = abdominal segment X; ma, la and va = mesal, lateral and ventral arms of segment X, respectively; ia = inferior appendage; ih, dh and maia = inner hook, dorsal hook and main article of an inferior appendage, respectively; ph = phallic apparatus. Wings: 1 – 2 = apical forks I and II; d = discoidal cell; th = thyridial cell; ar = arculus; C = costa; Sc = subcosta; R = radius; Ma = anterior media; Mp = posterior media; Cu = cubitus; A = anal vein." captionText-1="FIGURES 2, 3. Lepidostoma coreanum, male, variation. 2, specimen from Shizuoka, Amagi-yugashima-cho; 3, specimen from Wakayama. A, head, left scape and maxillary palp, left lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral; F, apex of main article of right inferior appendage, ventral, enlarged." captionText-2="FIGURES 4, 5. Lepidostoma coreanum, male, variation. 4, specimen from Ehime; 5, specimen from Kochi. A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral." httpUri-0="https://zenodo.org/record/256220/files/figure.png" httpUri-1="https://zenodo.org/record/256221/files/figure.png" httpUri-2="https://zenodo.org/record/256222/files/figure.png" pageId="1" pageNumber="398">Figs. 1–5</figureCitation>
). Specimens from Primorye (
<figureCitation id="D28152F6FFF5F64053A9FD71FD3AFD53" box="[673,739,705,730]" captionStart="FIGURE 1" captionStartId="2.[151,250,1819,1841]" captionTargetBox="[176,1409,216,1782]" captionTargetId="figure@2.[151,1436,193,1812]" captionTargetPageId="2" captionText="FIGURE 1. Lepidostoma coreanum, male, specimens from Primorye. A, head, right scape and maxillary palp, right lateral; B, right scape, ventral; C, right forewing, dorsal; D, right hind wing, dorsal; E, genitalia, left lateral; F, same, dorsal; G, same, ventral. Abbreviations. Body: IX = abdominal segment IX, X = abdominal segment X; ma, la and va = mesal, lateral and ventral arms of segment X, respectively; ia = inferior appendage; ih, dh and maia = inner hook, dorsal hook and main article of an inferior appendage, respectively; ph = phallic apparatus. Wings: 1 – 2 = apical forks I and II; d = discoidal cell; th = thyridial cell; ar = arculus; C = costa; Sc = subcosta; R = radius; Ma = anterior media; Mp = posterior media; Cu = cubitus; A = anal vein." httpUri="https://zenodo.org/record/256220/files/figure.png" pageId="1" pageNumber="398">Fig. 1</figureCitation>
). Antennae about
<quantity id="8D42E396FFF5F64052BFFD71FBD6FD50" box="[951,1039,705,730]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="398" unit="mm" value="5.5">5.5 mm</quantity>
long, scape setose, thick and long, 1.5 times as long as head, short mesal process near middle of each scape. Maxillary palps extremely setose, each 2- segmented,
<quantity id="8D42E396FFF5F6405017FCBAFEA1FCA8" box="[287,376,778,802]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="398" unit="mm" value="0.8">0.8 mm</quantity>
long in total, basal segment curved anteriorly at half its length with tuft of very long setae apically, long apical segment emerging near middle of basal segment. Labial palps each 3-segmented, slender, 1.0 mm long in total.
</paragraph>
<paragraph id="4A054E73FFF5F64051CFFCC5FEB2FB97" blockId="1.[151,1437,705,1990]" pageId="1" pageNumber="398">
Wings covered with fine setae, short scales, and spindle-like setae; spindle-like setae lined and circled in coastal area; forewings each with discoidal cell, thyridial cell, and apical forks 1 and 2, Cu2 indistinct at basal half and fused to Cu1 and A1+2+3 apically, arculus large; hind wings each with fork 1, Cu2 fused to Cu1 apically, A1+2+3 also fused to Cu1 apically; forewings and hind wings each 5.0–
<quantity id="8D42E396FFF5F64052FDFC51FB89FC70" box="[1013,1104,993,1018]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="398" unit="mm" value="5.2">5.2 mm</quantity>
long and
<quantity id="8D42E396FFF5F64055C4FC52FA81FC70" box="[1228,1368,994,1018]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.7" pageId="1" pageNumber="398" unit="mm" value="4.75" valueMax="4.8" valueMin="4.7">4.7–4.8 mm</quantity>
long, respectively (n=3).
</paragraph>
<paragraph id="4A054E73FFF5F64051CFFB99FF3FFAEB" blockId="1.[151,1437,705,1990]" pageId="1" pageNumber="398">
<emphasis id="78CE9261FFF5F64051CFFB99FEE8FBC8" box="[199,305,1065,1089]" italics="true" pageId="1" pageNumber="398">Genitalia</emphasis>
. Abdominal segment IX (IX) short, dorsum longer than venter. Segment X (X) composed of slender mesal (ma), lateral (la), and ventral arms (va); mesal arms longest and directed caudad, lateral arms directed caudad and distinctly shorter than mesal arms, ventral arms curved slightly dorsocaudad with length subequal to lateral arms; 3 arms subacute apically. Inferior appendages (ia) setose, each composed of main article (maia), dorsal hook (dh), and inner hook (ih); basal half of main article broad in lateral view, semicircular with serrate posterior margin in ventral view; apical half of main article bar-like, curved mesad, mesocaudal apex rather acute; dorsal hook directed laterocaudad, length subequal to main article; inner hook much shorter than dorsal hook, bar-like and directed laterocaudad. Phallic apparatus thick, apical half membranous with inner U-shaped sclerite, paramere absent.
</paragraph>
<paragraph id="4A054E73FFF5F64051CFFADEFC78F9D7" blockId="1.[151,1437,705,1990]" pageId="1" pageNumber="398">
<emphasis id="78CE9261FFF5F64051CFFADEFEF6FA0C" box="[199,303,1390,1413]" italics="true" pageId="1" pageNumber="398">Variation</emphasis>
(
<figureCitation id="D28152F6FFF5F6405048FADDFE69FA0F" box="[320,432,1389,1414]" captionStart-0="FIGURES 2, 3" captionStart-1="FIGURES 4, 5" captionStartId-0="3.[151,264,1749,1772]" captionStartId-1="4.[151,264,1822,1845]" captionTargetBox-0="[175,1400,213,1704]" captionTargetBox-1="[259,1417,226,1777]" captionTargetId-0="figure@3.[151,1436,193,1728]" captionTargetId-1="figure@4.[151,1436,193,1801]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 2, 3. Lepidostoma coreanum, male, variation. 2, specimen from Shizuoka, Amagi-yugashima-cho; 3, specimen from Wakayama. A, head, left scape and maxillary palp, left lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral; F, apex of main article of right inferior appendage, ventral, enlarged." captionText-1="FIGURES 4, 5. Lepidostoma coreanum, male, variation. 4, specimen from Ehime; 5, specimen from Kochi. A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral." httpUri-0="https://zenodo.org/record/256221/files/figure.png" httpUri-1="https://zenodo.org/record/256222/files/figure.png" pageId="1" pageNumber="398">Figs. 2–5</figureCitation>
). Specimens of all 4 localities in
<collectingCountry id="32AD0EE3FFF5F6405238FADEFCA9FA0C" box="[816,880,1390,1413]" name="Japan" pageId="1" pageNumber="398">Japan</collectingCountry>
exhibit some variations. In a male from Amagiyugashima-cho, Shizuoka (
<figureCitation id="D28152F6FFF5F64050C6FA21FDCCFA23" box="[462,533,1425,1450]" captionStart="FIGURES 2, 3" captionStartId="3.[151,264,1749,1772]" captionTargetBox="[175,1400,213,1704]" captionTargetId="figure@3.[151,1436,193,1728]" captionTargetPageId="3" captionText="FIGURES 2, 3. Lepidostoma coreanum, male, variation. 2, specimen from Shizuoka, Amagi-yugashima-cho; 3, specimen from Wakayama. A, head, left scape and maxillary palp, left lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral; F, apex of main article of right inferior appendage, ventral, enlarged." httpUri="https://zenodo.org/record/256221/files/figure.png" pageId="1" pageNumber="398">Fig. 2</figureCitation>
), the genitalia are almost identical to those of males from the
<typeStatus id="9501F0D1FFF5F64055E6FA22FAC7FA23" box="[1262,1310,1426,1450]" pageId="1" pageNumber="398">type</typeStatus>
locality in
<collectingCountry id="32AD0EE3FFF5F640519FFA05FEFCFA44" box="[151,293,1461,1486]" name="North Korea" pageId="1" pageNumber="398">North Korea</collectingCountry>
(
<bibRefCitation id="2E2B3382FFF5F640503DFA05FDBEFA47" author="Kumanski" box="[309,615,1461,1486]" pageId="1" pageNumber="417" refString="Kumanski, K. &amp; Weaver, J. S. III (1992) Studies on the fauna of Trichoptera (Insecta) of Korea. IV. The family Lepidostomatidae. Aquatic Insects, 14, 153 - 168. http: // dx. doi. org / 10.1080 / 01650429209361478" type="journal article" year="1992">Kumanski &amp; Weaver 1992</bibRefCitation>
) and from Primorye (
<figureCitation id="D28152F6FFF5F6405268FA05FC7AFA47" box="[864,931,1461,1486]" captionStart="FIGURE 1" captionStartId="2.[151,250,1819,1841]" captionTargetBox="[176,1409,216,1782]" captionTargetId="figure@2.[151,1436,193,1812]" captionTargetPageId="2" captionText="FIGURE 1. Lepidostoma coreanum, male, specimens from Primorye. A, head, right scape and maxillary palp, right lateral; B, right scape, ventral; C, right forewing, dorsal; D, right hind wing, dorsal; E, genitalia, left lateral; F, same, dorsal; G, same, ventral. Abbreviations. Body: IX = abdominal segment IX, X = abdominal segment X; ma, la and va = mesal, lateral and ventral arms of segment X, respectively; ia = inferior appendage; ih, dh and maia = inner hook, dorsal hook and main article of an inferior appendage, respectively; ph = phallic apparatus. Wings: 1 – 2 = apical forks I and II; d = discoidal cell; th = thyridial cell; ar = arculus; C = costa; Sc = subcosta; R = radius; Ma = anterior media; Mp = posterior media; Cu = cubitus; A = anal vein." httpUri="https://zenodo.org/record/256220/files/figure.png" pageId="1" pageNumber="398">Fig. 1</figureCitation>
), but the wing venation and maxillary palps are a little different from those of the later; in the Amagi-yugashima-cho specimens, Ma and Mp are stalked, crossvein
<emphasis id="78CE9261FFF5F6405001FA4EFE98F99C" box="[265,321,1534,1557]" italics="true" pageId="1" pageNumber="398">m-cu</emphasis>
and the thyridial cell are absent in each forewing; crossvein
<emphasis id="78CE9261FFF5F64052FFFA4EFBF6F99C" box="[1015,1071,1534,1557]" italics="true" pageId="1" pageNumber="398">m-cu</emphasis>
and the apical parts of Cu2 and A1+A2+A3 are lacking in each hind wing; the basal segment of each maxillary palp is thickened apically and the long apical segment emerges from the basal 2/3 of the basal segment.
</paragraph>
<paragraph id="4A054E73FFF5F64051CFF9DAFE17F92F" blockId="1.[151,1437,705,1990]" pageId="1" pageNumber="398">
In a male from Wakayama (
<figureCitation id="D28152F6FFF5F64050F6F9D9FD9BF90B" box="[510,578,1641,1666]" captionStart="FIGURES 2, 3" captionStartId="3.[151,264,1749,1772]" captionTargetBox="[175,1400,213,1704]" captionTargetId="figure@3.[151,1436,193,1728]" captionTargetPageId="3" captionText="FIGURES 2, 3. Lepidostoma coreanum, male, variation. 2, specimen from Shizuoka, Amagi-yugashima-cho; 3, specimen from Wakayama. A, head, left scape and maxillary palp, left lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral; F, apex of main article of right inferior appendage, ventral, enlarged." httpUri="https://zenodo.org/record/256221/files/figure.png" pageId="1" pageNumber="398">Fig. 3</figureCitation>
), the lateral arms of segment X are reduced, hump-like; the basal 2/3 of Cu2 is lacking in each forewing.
</paragraph>
<paragraph id="4A054E73FFF5F64051CFF902FDDBF8D3" blockId="1.[151,1437,705,1990]" pageId="1" pageNumber="398">
In the males from Ehime (
<figureCitation id="D28152F6FFF5F64050E4F901FDF6F943" box="[492,559,1713,1738]" captionStart="FIGURES 4, 5" captionStartId="4.[151,264,1822,1845]" captionTargetBox="[259,1417,226,1777]" captionTargetId="figure@4.[151,1436,193,1801]" captionTargetPageId="4" captionText="FIGURES 4, 5. Lepidostoma coreanum, male, variation. 4, specimen from Ehime; 5, specimen from Kochi. A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral." httpUri="https://zenodo.org/record/256222/files/figure.png" pageId="1" pageNumber="398">Fig. 4</figureCitation>
) and
<collectingCountry id="32AD0EE3FFF5F6405366F901FD6AF943" box="[622,691,1713,1738]" name="India" pageId="1" pageNumber="398">Kochi</collectingCountry>
(
<figureCitation id="D28152F6FFF5F64053C9F901FCDCF943" box="[705,773,1713,1738]" captionStart="FIGURES 4, 5" captionStartId="4.[151,264,1822,1845]" captionTargetBox="[259,1417,226,1777]" captionTargetId="figure@4.[151,1436,193,1801]" captionTargetPageId="4" captionText="FIGURES 4, 5. Lepidostoma coreanum, male, variation. 4, specimen from Ehime; 5, specimen from Kochi. A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral." httpUri="https://zenodo.org/record/256222/files/figure.png" pageId="1" pageNumber="398">Fig. 5</figureCitation>
), the lateral arms of segment X are reduced to a low hump, the main article of each inferior appendage has a round convex posteromesal margin in ventral view. The maxillary palps are setose, short, and unsegmented. Cubital and anal veins of forewings and hind wings are not fused apically and an arculus is absent in cubital area and some crossveins are lacking. Cu1 has a short apical fork
<quantity id="8D42E396FFF5F64055F5F8ADFAF0F8BF" box="[1277,1321,1821,1846]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="398" unit="in" value="5.0">5 in</quantity>
each hind wing of a specimen from
<collectingCountry id="32AD0EE3FFF5F64050BEF8F1FE27F8D3" box="[438,510,1857,1882]" name="India" pageId="1" pageNumber="398">Kochi</collectingCountry>
.
</paragraph>
<paragraph id="4A054E73FFF5F64051CFF8D5FD29F84F" blockId="1.[151,1437,705,1990]" pageId="1" pageNumber="398">
<emphasis id="78CE9261FFF5F64051CFF8D5FEC7F8F7" bold="true" box="[199,286,1893,1918]" pageId="1" pageNumber="398">Female</emphasis>
(
<figureCitation id="D28152F6FFF5F6405026F8D5FEA8F8F7" box="[302,369,1893,1918]" captionStart="FIGURE 6" captionStartId="5.[151,250,1877,1899]" captionTargetBox="[234,1370,195,1876]" captionTargetId="figure@5.[232,1370,193,1907]" captionTargetPageId="5" captionText="FIGURE 6. Lepidostoma coreanum, female, larva, and case, specimens from Primorye. A – F, Female: A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, abdominal segments VII – X, left lateral; E, same, ventral, subgenital plate omitted; F, subgenital plate, ventral, posterior edge up. G – I, Larva: G, habitus, left lateral; H, right half of thorax, dorsal. I, Case: case of larva, left lateral. G – H, from Ito (1998). Abbreviations: VII, VIII, IX, and X = abdominal segments VII – X, respectively; lp = lateral plate; sp = subgenital plate; wing labels, as in Fig. 1." httpUri="https://zenodo.org/record/256223/files/figure.png" pageId="1" pageNumber="398">Fig. 6</figureCitation>
A–F). Specimens from Primorye. Antennae about
<quantity id="8D42E396FFF5F64052A0F8D5FC27F8F4" box="[936,1022,1893,1918]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="398" unit="mm" value="5.5">5.5 mm</quantity>
long, scape setose, thick, long and 2 times as long as head, mesal processes absent. Maxillary palps each 5-segmented and about
<quantity id="8D42E396FFF5F640559FF83AFB01F828" box="[1175,1240,1930,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="398" unit="mm" value="1.0">1 mm</quantity>
long, labial palps each 3-segmented and
<quantity id="8D42E396FFF5F640509FF81DFE34F84C" box="[407,493,1965,1990]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="398" unit="mm" value="0.6">0.6 mm</quantity>
long; all palps slender.
</paragraph>
<caption id="1EC51EFBFFF6F643519FF8ABFF13F860" httpUri="https://zenodo.org/record/256220/files/figure.png" pageId="2" pageNumber="399" targetBox="[176,1409,216,1782]" targetPageId="2">
<paragraph id="4A054E73FFF6F643519FF8ABFF13F860" blockId="2.[151,1436,1819,2025]" pageId="2" pageNumber="399">
<emphasis id="78CE9261FFF6F643519FF8ABFED5F8B9" bold="true" box="[151,268,1819,1841]" pageId="2" pageNumber="399">FIGURE 1</emphasis>
.
<taxonomicName id="8DBA35F0FFF6F6435010F8ABFDD4F8B8" box="[280,525,1819,1841]" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="coreanum">
<emphasis id="78CE9261FFF6F6435010F8ABFDD4F8B8" bold="true" box="[280,525,1819,1841]" italics="true" pageId="2" pageNumber="399">Lepidostoma coreanum</emphasis>
</taxonomicName>
, male, specimens from Primorye. A, head, right scape and maxillary palp, right lateral; B, right scape, ventral; C, right forewing, dorsal; D, right hind wing, dorsal; E, genitalia, left lateral; F, same, dorsal; G, same, ventral. Abbreviations. Body: IX = abdominal segment IX, X = abdominal segment X; ma, la and va = mesal, lateral and ventral arms of segment X, respectively; ia = inferior appendage; ih, dh and maia = inner hook, dorsal hook and main article of an inferior appendage, respectively; ph = phallic apparatus. Wings: 1–2 = apical forks I and II; d = discoidal cell; th = thyridial cell; ar = arculus; C = costa; Sc = subcosta; R = radius; Ma = anterior media; Mp = posterior media; Cu = cubitus; A = anal vein.
</paragraph>
</caption>
<caption id="1EC51EFBFFF7F642519FF965FB0FF8A0" httpUri="https://zenodo.org/record/256221/files/figure.png" pageId="3" pageNumber="400" targetBox="[175,1400,213,1704]" targetPageId="3">
<paragraph id="4A054E73FFF7F642519FF965FB0FF8A0" blockId="3.[151,1436,1749,1833]" pageId="3" pageNumber="400">
<emphasis id="78CE9261FFF7F642519FF965FEE3F962" bold="true" box="[151,314,1749,1772]" pageId="3" pageNumber="400">FIGURES 2, 3</emphasis>
.
<taxonomicName id="8DBA35F0FFF7F6425041F965FD98F962" box="[329,577,1749,1771]" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="coreanum">
<emphasis id="78CE9261FFF7F6425041F965FD98F962" bold="true" box="[329,577,1749,1771]" italics="true" pageId="3" pageNumber="400">Lepidostoma coreanum</emphasis>
</taxonomicName>
, male, variation. 2, specimen from Shizuoka, Amagi-yugashima-cho; 3, specimen from Wakayama. A, head, left scape and maxillary palp, left lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral; F, apex of main article of right inferior appendage, ventral, enlarged.
</paragraph>
</caption>
<caption id="1EC51EFBFFF0F645519FF8AEFF3CF8FB" httpUri="https://zenodo.org/record/256222/files/figure.png" pageId="4" pageNumber="401" targetBox="[259,1417,226,1777]" targetPageId="4">
<paragraph id="4A054E73FFF0F645519FF8AEFF3CF8FB" blockId="4.[151,1436,1822,1906]" pageId="4" pageNumber="401">
<emphasis id="78CE9261FFF0F645519FF8AEFEE1F8BD" bold="true" box="[151,312,1822,1845]" pageId="4" pageNumber="401">FIGURES 4, 5</emphasis>
.
<taxonomicName id="8DBA35F0FFF0F645504EF8AEFDE7F8BD" box="[326,574,1822,1844]" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="coreanum">
<emphasis id="78CE9261FFF0F645504EF8AEFDE7F8BD" bold="true" box="[326,574,1822,1844]" italics="true" pageId="4" pageNumber="401">Lepidostoma coreanum</emphasis>
</taxonomicName>
, male, variation. 4, specimen from Ehime; 5, specimen from Kochi. A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, genitalia, left lateral; E, same, ventral.
</paragraph>
</caption>
<caption id="1EC51EFBFFF1F644519FF8E5FB21F86F" httpUri="https://zenodo.org/record/256223/files/figure.png" pageId="5" pageNumber="402" targetBox="[234,1370,195,1876]" targetPageId="5">
<paragraph id="4A054E73FFF1F644519FF8E5FB21F86F" blockId="5.[151,1436,1877,2022]" pageId="5" pageNumber="402">
<emphasis id="78CE9261FFF1F644519FF8E5FED7F8E2" bold="true" box="[151,270,1877,1899]" pageId="5" pageNumber="402">FIGURE 6</emphasis>
.
<taxonomicName id="8DBA35F0FFF1F6445014F8E5FDCDF8E2" box="[284,532,1877,1899]" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="402" phylum="Arthropoda" rank="species" species="coreanum">
<emphasis id="78CE9261FFF1F6445014F8E5FDCDF8E2" bold="true" box="[284,532,1877,1899]" italics="true" pageId="5" pageNumber="402">Lepidostoma coreanum</emphasis>
</taxonomicName>
, female, larva, and case, specimens from Primorye. A–F,
<emphasis id="78CE9261FFF1F6445567F8E6FB67F8E2" bold="true" box="[1135,1214,1878,1899]" pageId="5" pageNumber="402">Female</emphasis>
: A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, abdominal segments VII–X, left lateral; E, same, ventral, subgenital plate omitted; F, subgenital plate, ventral, posterior edge up. G–I,
<emphasis id="78CE9261FFF1F64455BCF823FB2FF821" bold="true" box="[1204,1270,1939,1960]" pageId="5" pageNumber="402">Larva</emphasis>
: G, habitus, left lateral; H, right half of thorax, dorsal. I,
<emphasis id="78CE9261FFF1F6445322F801FD87F84E" bold="true" box="[554,606,1969,1991]" pageId="5" pageNumber="402">Case</emphasis>
: case of larva, left lateral. G–H, from Ito (1998). Abbreviations: VII, VIII, IX, and X = abdominal segments VII–X, respectively; lp = lateral plate; sp = subgenital plate; wing labels, as in Fig. 1.
</paragraph>
</caption>
<paragraph id="4A054E73FFF2F64751CFFF27FC41FF71" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="403">Wings covered with short fine setae, typical venation in females of the family; forewings with apical forks 1, 2, 3, 5, discoidal and thyridial cells, and arculus; hind wings with forks 1, 2 and 5. Lengths of forewings and hind wings on each side 5.1–6.0 mm and 4.0–5.0 mm, respectively (n=5).</paragraph>
<paragraph id="4A054E73FFF2F64751CFFEB4FE03FEEC" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="403">Lateral ends of tergite VIII broad. Lateral plates weakly sclerotized, trianglular. Subgenital plate sclerotized, subquadrate with slightly concave posterior margin and middle depression on anterior margin. Vaginal apparatus oval and slightly sclerotized.</paragraph>
<paragraph id="4A054E73FFF2F64751CFFEDFFD89FD21" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="403">
<emphasis id="78CE9261FFF2F64751CFFEDFFEC9FE01" bold="true" box="[199,272,367,392]" pageId="6" pageNumber="403">Larva</emphasis>
(
<figureCitation id="D28152F6FFF2F6475016FEDFFEB2FE01" box="[286,363,367,392]" captionStart="FIGURE 6" captionStartId="5.[151,250,1877,1899]" captionTargetBox="[234,1370,195,1876]" captionTargetId="figure@5.[232,1370,193,1907]" captionTargetPageId="5" captionText="FIGURE 6. Lepidostoma coreanum, female, larva, and case, specimens from Primorye. A – F, Female: A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, abdominal segments VII – X, left lateral; E, same, ventral, subgenital plate omitted; F, subgenital plate, ventral, posterior edge up. G – I, Larva: G, habitus, left lateral; H, right half of thorax, dorsal. I, Case: case of larva, left lateral. G – H, from Ito (1998). Abbreviations: VII, VIII, IX, and X = abdominal segments VII – X, respectively; lp = lateral plate; sp = subgenital plate; wing labels, as in Fig. 1." httpUri="https://zenodo.org/record/256223/files/figure.png" pageId="6" pageNumber="403">Figs. 6</figureCitation>
G, H). Specimens from Primorye. Head width and body length of final instar larva up to
<quantity id="8D42E396FFF2F647544FFEC0FA45FE0E" box="[1351,1436,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="403" unit="mm" value="0.7">0.7 mm</quantity>
and
<quantity id="8D42E396FFF2F64751C4FE25FECDFE25" box="[204,276,405,428]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="403" unit="mm" value="7.0">7 mm</quantity>
, respectively. Head dark brown, with numerous short secondary setae on dorsum in early instars. Pronotum dark brown. Mesonotal plate brown with some light colored dots on anterior half and light brown on posterior half. Chaetotaxy of thorax: 35–40 setae on pronotum; 7, about 12 and about 12 setae on setal areas (
<emphasis id="78CE9261FFF2F6475456FE6DFAAEFE7D" box="[1374,1399,477,500]" italics="true" pageId="6" pageNumber="403">sa</emphasis>
) 1,
<emphasis id="78CE9261FFF2F647519FFDB1FF69FD91" box="[151,176,513,536]" italics="true" pageId="6" pageNumber="403">sa</emphasis>
2 and
<emphasis id="78CE9261FFF2F647500AFDB1FEC2FD91" box="[258,283,513,536]" italics="true" pageId="6" pageNumber="403">sa</emphasis>
3, respectively, of mesonotum; and 1, about 5, and about 12 on
<emphasis id="78CE9261FFF2F647550DFDB1FBC7FD91" box="[1029,1054,513,536]" italics="true" pageId="6" pageNumber="403">sa</emphasis>
1,
<emphasis id="78CE9261FFF2F6475534FDB1FB8CFD91" box="[1084,1109,513,536]" italics="true" pageId="6" pageNumber="403">sa</emphasis>
2, and
<emphasis id="78CE9261FFF2F64755A6FDB1FB1EFD91" box="[1198,1223,513,536]" italics="true" pageId="6" pageNumber="403">sa</emphasis>
3, respectively, of metonotum. Single tracheal gills present in subdorsal rows near posterior margins of segments II–VI, in subventral rows near posterior margins of segments II–VII, and in subdorsal rows near anterior margins of segments II–III (or II–IV or II–V). Single subventral gill present near anterior margin of segment II in some specimens. Other characters as in
<taxonomicName id="8DBA35F0FFF2F6475042FD21FD92FD21" authority="Ito 1984" authorityName="Ito" authorityYear="1984" box="[330,587,656,680]" class="Insecta" family="Lepidostomatidae" genus="Lepidostoma" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="78CE9261FFF2F6475042FD21FE08FD21" box="[330,465,656,680]" italics="true" pageId="6" pageNumber="403">L. robustum</emphasis>
(
<bibRefCitation id="2E2B3382FFF2F64750E8FD20FD9AFD21" author="Ito" box="[480,579,656,680]" pageId="6" pageNumber="417" refString="Ito, T. (1984) Three new species of Dinarthrum (Trichoptera, Lepidostomatidae). Kontyu, Tokyo, 52, 1 - 20." type="journal article" year="1984">Ito 1984</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="4A054E73FFF2F64751CFFD04FBA0FD66" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="403">
<emphasis id="78CE9261FFF2F64751CFFD04FEAEFD44" bold="true" box="[199,375,692,717]" pageId="6" pageNumber="403">Case of larva</emphasis>
(
<figureCitation id="D28152F6FFF2F6475087FD04FE04FD44" box="[399,477,692,717]" captionStart="FIGURE 6" captionStartId="5.[151,250,1877,1899]" captionTargetBox="[234,1370,195,1876]" captionTargetId="figure@5.[232,1370,193,1907]" captionTargetPageId="5" captionText="FIGURE 6. Lepidostoma coreanum, female, larva, and case, specimens from Primorye. A – F, Female: A, head, right scape and maxillary palp, right lateral; B, right forewing, dorsal; C, right hind wing, dorsal; D, abdominal segments VII – X, left lateral; E, same, ventral, subgenital plate omitted; F, subgenital plate, ventral, posterior edge up. G – I, Larva: G, habitus, left lateral; H, right half of thorax, dorsal. I, Case: case of larva, left lateral. G – H, from Ito (1998). Abbreviations: VII, VIII, IX, and X = abdominal segments VII – X, respectively; lp = lateral plate; sp = subgenital plate; wing labels, as in Fig. 1." httpUri="https://zenodo.org/record/256223/files/figure.png" pageId="6" pageNumber="403">
Fig.
<date id="3E0468B3FFF2F64750C5FD04FE04FD44" box="[461,477,692,717]" pageId="6" pageNumber="403">6</date>
</figureCitation>
I). Specimens from Primorye. Cylindrical, slightly curved and slightly tapered posteriorly, composed of sand grains with round posterior opening. Length up to
<quantity id="8D42E396FFF2F647552BFD68FBACFD66" box="[1059,1141,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="403" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="02A01DF8FFF2F64751CFFD4CFB83FB4C" pageId="6" pageNumber="403" type="discussion">
<paragraph id="4A054E73FFF2F64751CFFD4CFD30FC09" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="403">
<emphasis id="78CE9261FFF2F64751CFFD4CFEEDFC9C" bold="true" box="[199,308,764,789]" pageId="6" pageNumber="403">Remarks</emphasis>
. The males of this species have some variations locally but they are clearly distinguished from those of other related species by their elongated ventral arms of segment X. Records of this species from Mie, Honshu,
<collectingCountry id="32AD0EE3FFF2F647519FFCF5FF03FCD5" box="[151,218,837,860]" name="Japan" pageId="6" pageNumber="403">Japan</collectingCountry>
, by
<bibRefCitation id="2E2B3382FFF2F647500FFCF5FDF4FCD5" author="Kawase" box="[263,557,836,861]" pageId="6" pageNumber="417" refString="Kawase, H. &amp; Morita, H. (2001) 2000 nen ni saishushita tobikera rui [Caddisflies collected in 2000]. Tokkuribachi, 68, 8 - 9. [in Japanese]" type="journal article" year="2001">Kawase and Morita (2001</bibRefCitation>
, 2010) and
<bibRefCitation id="2E2B3382FFF2F64753A7FCF4FC9FFCD5" author="Morita" box="[687,838,836,861]" pageId="6" pageNumber="417" refString="Morita, H. (2008) Minami-ise-cho no tobikera so [Caddisfly fauna of Minami-ise-cho]. Hirakura, 52, 91 - 93. [in Japanese]" type="journal article" year="2008">Morita (2008</bibRefCitation>
,
<bibRefCitation id="2E2B3382FFF2F6475259FCF4FC53FCD5" author="Morita" box="[849,906,836,860]" pageId="6" pageNumber="417" refString="Morita, H. (2011) Shima-hanto no tobikera-rui [Trichoptera of Shima Peninsula]. In: Narukawa, N. (Ed.), Shima-hanto no kontyu [Insects of Shima Peninsula]. Mie Kontyu Danwakai, Suzuka, pp. 212 - 218. [in Japanese]" type="book chapter" year="2011">2011</bibRefCitation>
) are uncertain because the specimens could not be examined with our present taxonomic knowledge.
</paragraph>
<paragraph id="4A054E73FFF2F64751CFFC3CFB83FB4C" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="403">
<emphasis id="78CE9261FFF2F64751CFFC3CFE1CFC2C" bold="true" box="[199,453,908,933]" pageId="6" pageNumber="403">Specimens examined</emphasis>
.
<emphasis id="78CE9261FFF2F64750D1FC3CFCEEFC2C" bold="true" box="[473,823,908,933]" pageId="6" pageNumber="403">Russian Far East, Primorye</emphasis>
,
<emphasis id="78CE9261FFF2F6475243FC3CFC10FC2C" bold="true" box="[843,969,908,933]" pageId="6" pageNumber="403">Khasanski</emphasis>
:
<specimenCount id="5CBC85FAFFF2F64752D5FC3CFBE5FC2C" box="[989,1084,908,933]" pageId="6" pageNumber="403" type="male">4 males</specimenCount>
, small water flow near Vitya Village,
<date id="3E0468B3FFF2F64751FDFC1FFEB6FC41" box="[245,367,943,968]" pageId="6" pageNumber="403" value="1988-06-30">30.vi.1988</date>
, T.S. Vshivkova;
<specimenCount id="5CBC85FAFFF2F647533EFC1FFD48FC41" box="[566,657,943,968]" pageId="6" pageNumber="403" type="larva">5 larvae</specimenCount>
, same locality,
<date id="3E0468B3FFF2F6475234FC1FFC72FC41" box="[828,939,943,968]" pageId="6" pageNumber="403" value="1996-05-11">11.v.1996</date>
, T.S. Vshivkova &amp; T. Ito;
<specimenCount id="5CBC85FAFFF2F64755D9FC00FAC7FC41" box="[1233,1310,943,968]" pageId="6" pageNumber="403" type="male">1 male</specimenCount>
,
<specimenCount id="5CBC85FAFFF2F6475422FC1FFA41FC41" box="[1322,1432,943,968]" pageId="6" pageNumber="403" type="female">6 females</specimenCount>
,
<specimenCount id="5CBC85FAFFF2F647519FFC64FF2BFC65" box="[151,242,980,1004]" pageId="6" pageNumber="403" type="pupa">2 pupae</specimenCount>
(
<specimenCount id="5CBC85FAFFF2F6475000FC64FE81FC64" box="[264,344,980,1005]" pageId="6" pageNumber="403" type="male">1 male</specimenCount>
,
<specimenCount id="5CBC85FAFFF2F647506FFC64FE08FC64" box="[359,465,980,1005]" pageId="6" pageNumber="403" type="female">1 female</specimenCount>
), same locality,
<date id="3E0468B3FFF2F6475387FC64FD25FC64" box="[655,764,980,1005]" pageId="6" pageNumber="403" value="1996-05-11">11.v.1996</date>
(larvae), reared and preserved in alcohol in
<date id="3E0468B3FFF2F6475401FC64FAB9FC64" box="[1289,1376,980,1005]" pageId="6" pageNumber="403" value="1996-06">vi.1996</date>
, T.S. Vshivkova &amp; T. Ito.
<emphasis id="78CE9261FFF2F6475089FC47FDEFFB99" bold="true" box="[385,566,1015,1040]" pageId="6" pageNumber="403">
<collectingCountry id="32AD0EE3FFF2F6475089FC47FE15FB99" box="[385,460,1015,1040]" name="Japan" pageId="6" pageNumber="403">Japan</collectingCountry>
, Honshu
</emphasis>
,
<emphasis id="78CE9261FFF2F647534DFC47FD6AFB99" bold="true" box="[581,691,1015,1040]" pageId="6" pageNumber="403">Shizuoka</emphasis>
:
<specimenCount id="5CBC85FAFFF2F64753CBFC48FCC8FB99" box="[707,785,1015,1040]" pageId="6" pageNumber="403" type="male">1 male</specimenCount>
,
<specimenCount id="5CBC85FAFFF2F6475215FC48FC5BFB99" box="[797,898,1015,1040]" pageId="6" pageNumber="403" type="female">1 female</specimenCount>
,
<specimenCount id="5CBC85FAFFF2F6475286FC48FC3EFB86" box="[910,999,1016,1040]" pageId="6" pageNumber="403" type="pupa">2 pupae</specimenCount>
(females), Izu-shi, Amagi-yugashimacho, Kami-funabara, Funabara-gawa,
<quantity id="8D42E396FFF2F6475348FBACFD51FBBD" box="[576,648,1052,1077]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="403" unit="m" value="550.0">550 m</quantity>
,
<date id="3E0468B3FFF2F647539CFBACFCD2FBBC" box="[660,779,1052,1077]" pageId="6" pageNumber="403" value="1996-04-21">21.iv.1996</date>
(larvae), reared and preserved in alcohol on
<date id="3E0468B3FFF2F6475408FBACFAA2FBBC" box="[1280,1403,1052,1077]" pageId="6" pageNumber="403" value="1996-06-24">24.vi.1996</date>
, T. Hattori &amp; T. Ito.
<emphasis id="78CE9261FFF2F647505FFB8FFE07FBD1" bold="true" box="[343,478,1087,1112]" pageId="6" pageNumber="403">Wakayama</emphasis>
:
<specimenCount id="5CBC85FAFFF2F64750E6FBF0FDE4FBD1" box="[494,573,1087,1112]" pageId="6" pageNumber="403" type="male">1 male</specimenCount>
, Kozagawa-cho, Hirai,
<date id="3E0468B3FFF2F6475240FBF0FC61FBD1" box="[840,952,1088,1112]" pageId="6" pageNumber="403" value="1993-05-20">20.v.1993</date>
, N. Kuhara, light trap.
<emphasis id="78CE9261FFF2F64755B6FB8FFAA6FBD1" bold="true" box="[1214,1407,1087,1112]" pageId="6" pageNumber="403">Shikoku, Ehime</emphasis>
:
<specimenCount id="5CBC85FAFFF2F6475486FBF0FF02FBF4" pageId="6" pageNumber="403" type="male">3 males</specimenCount>
,
<specimenCount id="5CBC85FAFFF2F64751FAFBD4FEBBFBF4" box="[242,354,1124,1149]" pageId="6" pageNumber="403" type="female">1 female</specimenCount>
, Uchiko-machi, Odamiyama, Miyamaso-mukai, small water flow,
<date id="3E0468B3FFF2F64755AAFBD4FAB4FBF4" box="[1186,1389,1124,1149]" pageId="6" pageNumber="403" value="2000-07-24" valueMax="2000-08-08" valueMin="2000-07-24">24.vii–8.viii.2000</date>
, E. Yamamoto, Malaise trap;
<specimenCount id="5CBC85FAFFF2F64750B1FB38FDF1FB29" box="[441,552,1159,1184]" pageId="6" pageNumber="403" type="female">3 females</specimenCount>
, Kuma-kogen-cho, Yurano-no-mori,
<date id="3E0468B3FFF2F64752C0FB37FB48FB29" box="[968,1169,1159,1184]" pageId="6" pageNumber="403" value="2007-06-30" valueMax="2007-07-31" valueMin="2007-06-30">30.vi–31.vii.2007</date>
, E. Yamamoto, Malaise trap.
<emphasis id="78CE9261FFF2F64751D8FB1CFECEFB4C" bold="true" box="[208,279,1196,1221]" pageId="6" pageNumber="403">
<collectingCountry id="32AD0EE3FFF2F64751D8FB1CFECEFB4C" box="[208,279,1196,1221]" name="India" pageId="6" pageNumber="403">Kochi</collectingCountry>
</emphasis>
:
<specimenCount id="5CBC85FAFFF2F647502EFB1CFEADFB4C" box="[294,372,1196,1221]" pageId="6" pageNumber="403" type="male">1 male</specimenCount>
, Okawa-mura, Funato, Yoshino-gawa,
<quantity id="8D42E396FFF2F6475222FB1CFCAAFB4D" box="[810,883,1196,1220]" metricMagnitude="2" metricUnit="m" metricValue="3.78" pageId="6" pageNumber="403" unit="m" value="378.0">378 m</quantity>
,
<date id="3E0468B3FFF2F6475277FB1CFC22FB4C" box="[895,1019,1196,1221]" pageId="6" pageNumber="403" value="2004-03-27">27.iii.2004</date>
, K. Nio.
</paragraph>
</subSubSection>
<subSubSection id="02A01DF8FFF2F64751CFFB7FFC7EFAB9" pageId="6" pageNumber="403" type="distribution">
<paragraph id="4A054E73FFF2F64751CFFB7FFDD7FA84" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="403">
<emphasis id="78CE9261FFF2F64751CFFB7FFE80FB61" bold="true" box="[199,345,1231,1256]" pageId="6" pageNumber="403">Distribution</emphasis>
(
<figureCitation id="D28152F6FFF2F6475064FB7FFE1BFB61" box="[364,450,1231,1256]" captionStart="FIGURE 21" captionStartId="19.[151,250,1367,1389]" captionTargetBox="[152,1414,187,1366]" captionTargetId="figure@19.[151,1436,171,1368]" captionTargetPageId="19" captionText="FIGURE 21. Distribution of the 5 species of the Lepidostoma coreanum Species Complex of the genus Lepidostoma. The plot in North Korea was referred from Kumanski &amp; Weaver (1992)." httpUri="https://zenodo.org/record/256233/files/figure.png" pageId="6" pageNumber="403">Fig. 21</figureCitation>
).
<collectingCountry id="32AD0EE3FFF2F64750D2FB7FFDB0FB61" box="[474,617,1231,1256]" name="North Korea" pageId="6" pageNumber="403">North Korea</collectingCountry>
(Province Pyongan Pukudo), Russian Far East (Primorye, Khasanski),
<collectingCountry id="32AD0EE3FFF2F647519FFB45FF0EFA85" box="[151,215,1269,1292]" name="Japan" pageId="6" pageNumber="403">Japan</collectingCountry>
(central Honshu, Shikoku).
</paragraph>
<paragraph id="4A054E73FFF2F64751CFFAA7FC7EFAB9" blockId="6.[151,1437,151,1365]" box="[199,935,1303,1328]" pageId="6" pageNumber="403">
<emphasis id="78CE9261FFF2F64751CFFAA7FEFAFAB9" bold="true" box="[199,291,1303,1328]" pageId="6" pageNumber="403">Habitat</emphasis>
. Adults were collected from small water flows in forests.
</paragraph>
</subSubSection>
<subSubSection id="02A01DF8FFF2F64751CFFA8CFD60FADC" box="[199,697,1340,1365]" pageId="6" pageNumber="403" type="description">
<paragraph id="4A054E73FFF2F64751CFFA8CFD60FADC" blockId="6.[151,1437,151,1365]" box="[199,697,1340,1365]" pageId="6" pageNumber="403">
<emphasis id="78CE9261FFF2F64751CFFA8CFEA5FADC" bold="true" box="[199,380,1340,1365]" pageId="6" pageNumber="403">Japanese name</emphasis>
. Korea-sunatsutsu-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>
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