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<mods:title id="D3919B54C82F3B4748C6462502EDC299">Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae 2697</mods:title>
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<taxonomicName id="F3F49381FFF85204FF48F996858CF92A" ID-CoL="49YWY" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[151,387,1658,1684]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioplexa" kingdom="Animalia" order="Ophioscolecida" pageId="78" pageNumber="79" phylum="Echinodermata" rank="species" species="condita" status="gen. et sp. nov.">
<emphasis id="06803410FFF85204FF48F996858CF92A" bold="true" box="[151,387,1658,1684]" italics="true" pageId="78" pageNumber="79">Ophioplexa condita</emphasis>
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<figureCitation id="ACCFF487FFF85204FF48F9718596F909" box="[151,409,1693,1719]" captionStart="FIGURE 4" captionStartId="13.[151,256,1339,1363]" captionTargetBox="[282,1312,194,1330]" captionTargetId="figure-434@13.[282,1312,194,1330]" captionTargetPageId="13" captionText="FIGURE 4. Genital plates and radial shields of various taxa of the families Asteronychidae, Gorgonocephalidae and Ophiomyxidae. A. Gorgonocephalus arcticus Leach, 1819, ZMMU D-613, 39 mm dd, adradial and abradial genital plates in situ, showing articulation surface, dorsal view; B. same, articulation surface of the radial shield, ventral view; C. Asteronyx loveni Müller &amp; Troschel, 1842, ZMMU D-833, 17.5 mm dd, articulation surface of the adradial genital plate, dorsal view; D. same, articulation surface of the radial shield, ventral view. E. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, articulation surface of the adradial genital plate, dorsal view; F. same, radial shield showing articulation surface, ventral view; G. same, articulating adradial genital plate and radial shield, dorsal view; H. Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, dissected disk showing most proximal vertebrae and absence of the both genital plates and radial shield, dorsal view; I. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, adradial and abradial genital plates in situ, ventral view; J. Ophioplexa condita gen. et sp. nov., paratype ca. 17 mm dd, ZMMU D-792, dissected disk showing articulating adradial genital plate and radial shield in situ, dorsal view; K. same, separate articulating adradial genital plate and radial shield, dorsal view; L. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, adradial genital plate, dorsal view; M. Ophiologimus cf. secundus Koehler, 1914, 4 mm dd, ZMMU D-814, adradial genital plate, dorsal view; N. Ophioplexa condita gen. et sp. nov., separate adradial genital plate, dorsal view; O. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, adradial and abradial genital plates, dorsal view; P. Ophioplexa condita gen. et sp. nov., separate articulating adradial genital plate and radial shield, dorsal view; Q. articulation surface of the adradial genital plate, dorsal view; R. radial shield, ventral view, showing distally the distinctive excavated articulation surface; S. articulation surface of the radial shield, ventral view; T. Ophiocymbium ninae sp. nov., paratype 5 mm dd, radial shield, ventral view; U. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, same, articulation surface of the radial shield, proximal view; Scale bars: A– B,1 mm; C–D, 300 µ; E–F, 100 µ; G, 200 µ; H, 300 µ; I, 200 µ; J, 1000 µ; K, 300 µ; L, 100 µ; M, 300 µ; N, 300 µ; O, 200 µ; P, 100 µ; Q–T, 100 µ; U, 200 µ. Photos: Alexander Martynov." pageId="78" pageNumber="79">Figures 4J, K, N, P–S</figureCitation>
; 
<figureCitation id="ACCFF487FFF85204FE7AF97185F4F909" box="[421,507,1693,1719]" captionStart="FIGURE 17" captionStartId="27.[151,255,1036,1060]" captionTargetBox="[161,1426,186,1015]" captionTargetId="figure-399@27.[161,1426,186,1015]" captionTargetPageId="27" captionText="FIGURE 17. Vertebrae of various taxa of the family Ophiomyxidae. A. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, proximal segments, dorsal and ventral views; B. same, distal segments, dorsal and ventral views; C. same, proximal segments, proximal view; D. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, 10.5 mm dd, proximal segments, dorsal view; E. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, proximal segments, dorsal view; F. Ophioscolex glacialis, proximal segments, distal view; G. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, proximal segments, proximal view; H. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, ventral view; I. same, proximal view; J. same, distal view; K. Ophiomyxa pentagona, ZMMU D-553, 21 mm dd, distal view; L. Ophioplexa condita gen. et sp. nov., holotype ca. 18 mm dd, ZMMU D-788, proximal segments, dorsal view; M. same, ventral view; N. same, proximal view; O. same, distal view; P. Ophiocymbium tanyae sp. nov., holotype 9.5 mm dd, ZMMU D-799, proximal segments, dorsal view; Q. same, ventral view; R. same, distal view; S. same, proximal view. Scale bars: A, 1000 µ; B, C, 200 µ, D, 200 µ; E, 300 µ; F, 200 µ; G, 200 µ; H, 300 µ; I, 100 µ; J, 100 µ; K, 1000 µ; L–S, 300 µ. Photos: Alexander Martynov." pageId="78" pageNumber="79">17L–O</figureCitation>
; 
<figureCitation id="ACCFF487FFF85204FDD8F9718651F909" box="[519,606,1693,1719]" captionStart="FIGURE 54" captionStartId="79.[151,257,1724,1748]" captionTargetBox="[158,1428,194,1703]" captionTargetId="figure-131@79.[158,1428,194,1703]" captionTargetPageId="79" captionText="FIGURE 54. Ophioplexa condita gen. nov., sp. nov. and Ophioscolex glacialis Müller &amp; Troschel, 1842, external views. A. Ophioplexa condita gen. nov., sp. nov., external view of the holotype ca. 18 mm dd, dorsal, ZMMU D-788, B. same ventral; C. a paratype 11.8 mm dd, dorsal view, ZMMU D-789; D. same, ventral view; E. holotype, ventral view of the arm; F. a paratype 8 mm dd, dorsal view, ZMMU D-793; G. same ventral; H. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-118, ethanol specimen 34 mm dd; I. Ophioscolex glacialis, ZMMU D-866, dried specimen 22 mm dd, showing changing in the external appearance compare to living and ethanol specimens; J. same, oral frame. Photos: Tatiana Korshunova." pageId="78" pageNumber="79">54A–E</figureCitation>
; 
<figureCitation id="ACCFF487FFF85204FDB5F97186CBF909" box="[618,708,1693,1719]" captionStart="FIGURE 55" captionStartId="80.[151,258,1764,1788]" captionTargetBox="[209,1364,194,1740]" captionTargetId="figure-174@80.[209,1364,194,1740]" captionTargetPageId="80" captionText="FIGURE 55. Ophioplexa condita gen. nov., sp. nov. and Ophiocymbium ninae sp. nov., details. A. Ophioplexa condita gen. nov., sp. nov., paratype 19 mm dd, ZMMU D-791, disk ventral view; B. same oral frame showing clusters of the ventral teeth (apical papillae); C. same, one interradius showing apical, oral and adoral shield papillae; D-F, same, most proximal arm segments showing tentacle scales and spines; G-H, arm spine articulations, distal segments; I. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-803, details, part of the oral frame and ventral arm side; J. same, jaw and oral papillae, one interradius; K. ventral arm view showing spines and tentacle scales; L. most proximal segments showing details of tentacle scales and spines; M-N, arm spine articulations, proximal segments; Scale bars: A– B, 1 mm; C–D, 500 µ; E–F, 200 µ; G–H, 100 µ; I, 1 mm; J–N, 300 µ. Photos: Alexander Martynov." pageId="78" pageNumber="79">55A–H</figureCitation>
; 
<figureCitation id="ACCFF487FFF85204FD0FF97186FFF909" box="[720,752,1693,1719]" captionStart="FIGURE 56" captionStartId="81.[151,255,1449,1473]" captionTargetBox="[188,1398,194,1427]" captionTargetId="figure-315@81.[188,1398,194,1428]" captionTargetPageId="81" captionText="FIGURE 56. Ophioplexa condita gen. nov., sp. nov. Arm spine articulations, spines, tentacle scales, dental plates and jaws of a paratype of ca. 17 mm dd, ZMMU D-792. A. two proximal arm segments adjoining disk; B. same, showing hollow spines; C. ventral view of the artciulations of the most proximal segments under the disk in situ including attaching area of the tentacle scales; D. separate lateral arm plate of the most proximal segments showing distinct shape of the ventralmost ridge; E. same, showing variability of following articulation; F. articulations of a middle segment; G. spine of a middle segment; H. separate lateral arm plate of the proximal segment showing more typical pattern of the articulation; I. a separate tentacle scale of the most proximal segment; J. same, basal part showing a small opening, presumably for nerve; K. basal part of a spine showing an opening for nerve; L. dental plate with several teeth; M, N, dental plates; O. jaw, abradial view; P. jaw, adradial view. Scale bars: A, 300; B, 100 µ; C, 300 µ; D, 100 µ; E, 100 µ; F,G, 100 µ; H, 300 µ; I,J, 100 µ; K, 30 µ; L–P, 300 µ. Photos: Alexander Martynov." pageId="78" pageNumber="79">56</figureCitation>
; 
<figureCitation id="ACCFF487FFF85204FD23F9718725F909" box="[764,810,1693,1719]" captionStart="FIGURE 67" captionStartId="98.[151,255,1494,1518]" captionTargetBox="[188,1398,679,1473]" captionTargetId="figure-460@98.[188,1398,679,1473]" captionTargetPageId="98" captionText="FIGURE 67. External views of various taxa of the families Ophiacanthidae and Ophiomyxidae. A. Ophiotoma alberti (Koehler, 1896), ZMMU D-810, 12 mm dd, dorsal view showing ventral teeth, soft disk damaged in centre after fixation; B. same, ventral view; C. same, ventro-lateral view showing long genital slits bordered by well-defined genital plates; D. Ophiotoma assimilis Koehler, 1904, ZMMU D-807, 18.5 mm dd, dorsal view; E. same, ventral view; F. Ophioplexa condita gen. nov., sp. nov., paratype 19 mm dd, ZMMU D-790, ventro-lateral view, the grooves between arm bases are not the genital slits, genital plates are not evident; G. Ophiocymbium rarispinum sp. nov., holotype 9.2 mm dd, ZMMU D-798, 9.2 mm dd. Photos: Tatiana Korshunova." pageId="78" pageNumber="79">67F</figureCitation>
</paragraph>
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<emphasis id="06803410FFF85204FF48F90B8504F8BF" bold="true" box="[151,267,1767,1793]" pageId="78" pageNumber="79">Material.</emphasis>
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, dried, 
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D-788, 
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“ Akademik Kurchatov”, cruise 11, sta. 896, 
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– 
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, depth 5651– 
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, Sigsbee trawl. One 
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D- 789, same sta. as 
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D-790, same sta. as 
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. One 
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, mounted on SEM stub, 
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D-791, same sta. as 
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. One 
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disarticulated and mounted on SEM stub, 
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D-792, same sta. as 
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D-793, 
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“Akademik Kurchatov”, cruise 11, sta. 870, 
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– 
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<geoCoordinate id="51C08EC5FFF85204FBC4F84E8088F802" box="[1051,1159,1954,1980]" degrees="25" direction="west" minutes="1" orientation="longitude" pageId="78" pageNumber="79" precision="925" value="-25.016666">25° 1' W</geoCoordinate>
, depth 4704– 
<quantity id="F30C45E7FFF85204FAE7F84E8198F802" box="[1336,1431,1954,1980]" metricMagnitude="3" metricUnit="m" metricValue="4.68" pageId="78" pageNumber="79" unit="m" value="4680.0">
<elevation id="BFD90F31FFF85204FAE7F84E8198F802" box="[1336,1431,1954,1980]" metricMagnitude="3" metricUnit="m" metricValue="4.68" pageId="78" pageNumber="79" unit="m" value="4680.0">4680 m</elevation>
</quantity>
, Sigsbee trawl.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="608BB88AFFF95205FF48F95085C3F82A" pageId="79" pageNumber="80" startId="79.[151,257,1724,1748]" targetBox="[158,1428,194,1703]" targetPageId="79">
<paragraph id="344BE802FFF95205FF48F95085C3F82A" blockId="79.[151,1437,1724,1940]" pageId="79" pageNumber="80">
<emphasis id="06803410FFF95205FF48F95086E3F96A" bold="true" box="[151,748,1724,1749]" pageId="79" pageNumber="80">
FIGURE 54. 
<taxonomicName id="F3F49381FFF95205FEEBF951861CF96B" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[308,531,1725,1749]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioplexa" kingdom="Animalia" order="Ophioscolecida" pageId="79" pageNumber="80" phylum="Echinodermata" rank="species" species="condita" status="gen. nov., sp. nov.">
<emphasis id="06803410FFF95205FEEBF951861CF96B" bold="true" box="[308,531,1725,1749]" italics="true" pageId="79" pageNumber="80">Ophioplexa condita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFFF95205FDC4F95086E3F96A" box="[539,748,1724,1748]" pageId="79" pageNumber="80" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="F3F49381FFF95205FCFBF9518139F96A" authority="Muller &amp; Troschel, 1842" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[804,1334,1725,1748]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="79" pageNumber="80" phylum="Echinodermata" rank="species" species="glacialis">
<emphasis id="06803410FFF95205FCFBF951801AF96A" box="[804,1045,1725,1748]" italics="true" pageId="79" pageNumber="80">Ophioscolex glacialis</emphasis>
<bibRefCitation id="506595F3FFF95205FBC1F9518139F96A" author="Muller, J. &amp; Troschel, F. H." box="[1054,1334,1725,1748]" pageId="79" pageNumber="80" refId="ref90891" refString="Muller, J., Troschel, F. H. (1842) System der Asteriden. 1. Asteridae. 2. Ophiuridae. Braunschweig." type="book" year="1842">Müller &amp; Troschel, 1842</bibRefCitation>
</taxonomicName>
, external views. A. 
<emphasis id="06803410FFF95205FEDDF93186A4F94A" bold="true" box="[258,683,1756,1781]" pageId="79" pageNumber="80">
<taxonomicName id="F3F49381FFF95205FEDDF93185D6F94B" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[258,473,1757,1781]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioplexa" kingdom="Animalia" order="Ophioscolecida" pageId="79" pageNumber="80" phylum="Echinodermata" rank="species" species="condita" status="gen. nov., sp. nov.">
<emphasis id="06803410FFF95205FEDDF93185D6F94B" bold="true" box="[258,473,1757,1781]" italics="true" pageId="79" pageNumber="80">Ophioplexa condita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFFF95205FE00F93086A9F94A" box="[479,678,1756,1780]" pageId="79" pageNumber="80" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
,
</emphasis>
external view of the holotype ca. 18 mm dd, dorsal, ZMMU D-788, B. same ventral; C. a paratype 11.8 mm dd, dorsal view, ZMMU D-789; D. same, ventral view; E. holotype, ventral view of the arm; F. a paratype 8 mm dd, dorsal view, ZMMU D-793; G. same ventral; H. 
<taxonomicName id="F3F49381FFF95205FBE9F8F18537F8EA" authority="Muller &amp; Troschel, 1842" authorityName="Muller &amp; Troschel" authorityYear="1842" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="79" pageNumber="80" phylum="Echinodermata" rank="species" species="glacialis">
<emphasis id="06803410FFF95205FBE9F8F18125F88A" box="[1078,1322,1821,1844]" italics="true" pageId="79" pageNumber="80">Ophioscolex glacialis</emphasis>
<bibRefCitation id="506595F3FFF95205FAECF8F18537F8EA" author="Muller, J. &amp; Troschel, F. H." pageId="79" pageNumber="80" refId="ref90891" refString="Muller, J., Troschel, F. H. (1842) System der Asteriden. 1. Asteridae. 2. Ophiuridae. Braunschweig." type="book" year="1842">Müller &amp; Troschel, 1842</bibRefCitation>
</taxonomicName>
, ZMMU D-118, ethanol specimen 34 mm dd; I. 
<taxonomicName id="F3F49381FFF95205FCE4F8D1808DF8EA" authority=", ZMMU" authorityName="ZMMU" box="[827,1154,1853,1876]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioscolex" kingdom="Animalia" order="Ophioscolecida" pageId="79" pageNumber="80" phylum="Echinodermata" rank="species" species="glacialis">
<emphasis id="06803410FFF95205FCE4F8D1802DF8EA" box="[827,1058,1853,1876]" italics="true" pageId="79" pageNumber="80">Ophioscolex glacialis</emphasis>
, ZMMU
</taxonomicName>
D-866, dried specimen 22 mm dd, showing changing in the external appearance compare to living and ethanol specimens; J. same, oral frame. Photos: Tatiana Korshunova.
</paragraph>
</caption>
<caption id="608BB88AFFE6521AFF48F90880B0F865" pageId="80" pageNumber="81" startId="80.[151,258,1764,1788]" targetBox="[209,1364,194,1740]" targetPageId="80">
<paragraph id="344BE802FFE6521AFF48F90880B0F865" blockId="80.[151,1436,1764,2012]" pageId="80" pageNumber="81">
<emphasis id="06803410FFE6521AFF48F9088026F942" bold="true" box="[151,1065,1764,1788]" pageId="80" pageNumber="81">
FIGURE 55. 
<taxonomicName id="F3F49381FFE6521AFEEAF9088618F942" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[309,535,1764,1788]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioplexa" kingdom="Animalia" order="Ophioscolecida" pageId="80" pageNumber="81" phylum="Echinodermata" rank="species" species="condita" status="gen. nov., sp. nov.">
<emphasis id="06803410FFE6521AFEEAF9088618F942" bold="true" box="[309,535,1764,1788]" italics="true" pageId="80" pageNumber="81">Ophioplexa condita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFFE6521AFDFFF90886FCF942" box="[544,755,1764,1788]" pageId="80" pageNumber="81" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="F3F49381FFE6521AFCEFF9088026F942" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[816,1065,1764,1788]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiocymbium" kingdom="Animalia" order="Ophioscolecida" pageId="80" pageNumber="81" phylum="Echinodermata" rank="species" species="ninae" status="sp. nov.">
<emphasis id="06803410FFE6521AFCEFF9088026F942" bold="true" box="[816,1065,1764,1788]" italics="true" pageId="80" pageNumber="81">Ophiocymbium ninae</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="1DB3896BFFE6521AFBEEF9088085F945" box="[1073,1162,1764,1788]" pageId="80" pageNumber="81" rank="species">
<emphasis id="06803410FFE6521AFBEEF908808AF942" bold="true" box="[1073,1157,1764,1788]" pageId="80" pageNumber="81">sp. nov</emphasis>
.
</taxonomicNameLabel>
, details. A. 
<emphasis id="06803410FFE6521AFAC8F90885B2F8A2" bold="true" pageId="80" pageNumber="81">
<taxonomicName id="F3F49381FFE6521AFAC8F90884E7F8A2" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioplexa" kingdom="Animalia" order="Ophioscolecida" pageId="80" pageNumber="81" phylum="Echinodermata" rank="species" species="condita" status="gen. nov., sp. nov.">
<emphasis id="06803410FFE6521AFAC8F90884E7F8A2" bold="true" italics="true" pageId="80" pageNumber="81">Ophioplexa condita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFFE6521AFF30F8E885B7F8A2" box="[239,440,1796,1820]" pageId="80" pageNumber="81" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
,
</emphasis>
paratype 19 mm dd, ZMMU D-791, disk ventral view; B. same oral frame showing clusters of the ventral teeth (apical papillae); C. same, one interradius showing apical, oral and adoral shield papillae; D-F, same, most proximal arm segments showing tentacle scales and spines; G-H, arm spine articulations, distal segments; I. 
<taxonomicName id="F3F49381FFE6521AFF48F888858BF8C2" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[151,388,1892,1916]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophiocymbium" kingdom="Animalia" order="Ophioscolecida" pageId="80" pageNumber="81" phylum="Echinodermata" rank="species" species="ninae" status="sp. nov.">Ophiocymbium ninae</taxonomicName>
<taxonomicNameLabel id="1DB3896BFFE6521AFE54F88885EFF8C5" box="[395,480,1892,1916]" pageId="80" pageNumber="81" rank="species">
<emphasis id="06803410FFE6521AFE54F88885D5F8C2" bold="true" box="[395,474,1892,1916]" pageId="80" pageNumber="81">sp. nov</emphasis>
.
</taxonomicNameLabel>
, paratype 5 mm dd, ZMMU D-803, details, part of the oral frame and ventral arm side; J. same, jaw and oral papillae, one interradius; K. ventral arm view showing spines and tentacle scales; L. most proximal segments showing details of tentacle scales and spines; M-N, arm spine articulations, proximal segments; Scale bars: A– B, 1 mm; C–D, 500 µ; E–F, 200 µ; G–H, 100 µ; I, 1 mm; J–N, 300 µ. Photos: Alexander Martynov.
</paragraph>
</caption>
<caption id="608BB88AFFE7521BFF48FA45801FF95F" pageId="81" pageNumber="82" startId="81.[151,255,1449,1473]" targetBox="[188,1398,194,1427]" targetPageId="81">
<paragraph id="344BE802FFE7521BFF48FA45801FF95F" blockId="81.[151,1437,1449,1762]" pageId="81" pageNumber="82">
<emphasis id="06803410FFE7521BFF48FA4586D6FA7F" bold="true" box="[151,729,1449,1473]" pageId="81" pageNumber="82">
FIGURE 56. 
<taxonomicName id="F3F49381FFE7521BFEF0FA458607FA7F" authority="Martynov, 2010" authorityName="Martynov" authorityYear="2010" box="[303,520,1449,1473]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioplexa" kingdom="Animalia" order="Ophioscolecida" pageId="81" pageNumber="82" phylum="Echinodermata" rank="species" species="condita" status="gen. nov., sp. nov.">
<emphasis id="06803410FFE7521BFEF0FA458607FA7F" bold="true" box="[303,520,1449,1473]" italics="true" pageId="81" pageNumber="82">Ophioplexa condita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1DB3896BFFE7521BFDD0FA4586D6FA7F" box="[527,729,1449,1473]" pageId="81" pageNumber="82" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
Arm spine articulations, spines, tentacle scales, dental plates and jaws of a paratype of ca. 17 mm dd, ZMMU D-792. A. two proximal arm segments adjoining disk; B. same, showing hollow spines; C. ventral view of the artciulations of the most proximal segments under the disk in situ including attaching area of the tentacle scales; D. separate lateral arm plate of the most proximal segments showing distinct shape of the ventralmost ridge; E. same, showing variability of following articulation; F. articulations of a middle segment; G. spine of a middle segment; H. separate lateral arm plate of the proximal segment showing more typical pattern of the articulation; I. a separate tentacle scale of the most proximal segment; J. same, basal part showing a small opening, presumably for nerve; K. basal part of a spine showing an opening for nerve; L. dental plate with several teeth; M, N, dental plates; O. jaw, abradial view; P. jaw, adradial view. Scale bars: A, 300; B, 100 µ; C, 300 µ; D, 100 µ; E, 100 µ; F,G, 100 µ; H, 300 µ; I,J, 100 µ; K, 30 µ; L–P, 300 µ. Photos: Alexander Martynov.
</paragraph>
</caption>
<subSubSection id="7CEEBB89FFE7521BFF19F8E58116F8F7" pageId="81" pageNumber="82" type="etymology">
<paragraph id="344BE802FFE7521BFF19F8E58116F8F7" blockId="81.[151,1436,1801,2014]" pageId="81" pageNumber="82">
<emphasis id="06803410FFE7521BFF19F8E5855CF89D" bold="true" box="[198,339,1801,1827]" pageId="81" pageNumber="82">Etymology.</emphasis>
from Latin 
<emphasis id="06803410FFE7521BFE39F8E68646F89D" box="[486,585,1802,1827]" italics="true" pageId="81" pageNumber="82">conditus</emphasis>
, hide, concealed, in reference to some morphological (small concealed radial shields and genital plates) and ecological (abyssal environment) features of the new species.
</paragraph>
</subSubSection>
<subSubSection id="7CEEBB89FFE7521EFF18F8B88621FCB9" lastPageId="84" lastPageNumber="85" pageId="81" pageNumber="82" type="description">
<paragraph id="344BE802FFE75218FF18F8B88161FC92" blockId="81.[151,1436,1801,2014]" lastBlockId="82.[151,1436,152,1970]" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
<emphasis id="06803410FFE7521BFF18F8B8862FF8D0" bold="true" box="[199,544,1876,1902]" pageId="81" pageNumber="82">
Description of the 
<typeStatus id="EB4F56A0FFE7521BFE71F8B88614F8D0" box="[430,539,1876,1902]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
.
</emphasis>
The disk is damaged, approximately 
<quantity id="F30C45E7FFE7521BFC06F8B88025F8D0" box="[985,1066,1876,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="81" pageNumber="82" unit="mm" value="18.0">18 mm</quantity>
dd, presumably flattened when alive, round and not indented interradially. The disk plates are numerous, most of them elongated and narrow, irregular-rhomboidal and polygonal, entirely devoid of spines and granules. There is a thin skin layer, which is conspicuous in ethanol specimens, but almost undetectable when dried. Radial shields are very small and entirely concealed under disc scales (see a detailed radial shield description under internal and other characters below). The interradii are slightly swollen, ventrally covered with numerous scales similar to the dorsal ones. Each jaw bears three to five irregularly placed, narrow, spiniform ventralmost teeth, which are not distinguished from more dorsally placed teeth, nor from adjacent lateral oral papillae. Distal oral papillae and adoral shield papillae are similar in size and shape, and placed along both edges of a jaw in a characteristic slightly curved row around the second tentacle pore, approximately 
<quantity id="F30C45E7FFE45218FC05FEBE8029FED2" box="[986,1062,338,364]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="82" pageNumber="83" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">5–6 in</quantity>
number (including 2–3 adoral papillae in the strict sense). Thus there is a continuous series of papillae from the teeth to the adoral shield papillae. Deeper in the mouth opening on the jaws there is a distinct group of papillae, placed laterally to the other oral papillae, 
<quantity id="F30C45E7FFE45218FE5DFE2E85C1FE62" box="[386,462,450,476]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="82" pageNumber="83" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">2–4 in</quantity>
number. Numerous spiniform teeth are arranged in a dorsalwards tapering coneshape on the dental plate. There are 17–20 teeth (including “apical papillae”). The oral shield is wide, triangular, with straight or slightly convex, distal edge, about as long as wide, completely separated from the first lateral arm plate by the adoral shields. Distally the oral shield is contiguous with a supplementary oral shield in form of a narrow elongated plate or just an elevation formed by adjacent interradial scales. The madreporite is similar in shape to the others, but slightly larger and has a swollen round area in the middle, conspicuously smaller than the entire shield. Adoral shields wing-like laterally, widely adjoining the arm, rapidly narrowing towards the jaws, thus retaining only a narrow bar between jaws and oral shield. True genital slits are small openings under the adoral shields, but long furrows between the proximalmost arm segments and the ventral disk interradius form false slits. Arm length is about 1.4 times the disk diameter.
</paragraph>
<paragraph id="344BE802FFE45218FF18FCD4818CFA4C" blockId="82.[151,1436,152,1970]" pageId="82" pageNumber="83">The dorsal arm plates are well developed but thin, fan-shaped, moderate in size. Few proximal plates are wide, rapidly narrowing towards the middle of the arm, contiguous along most of the arm length. The proximal edge of the dorsal arm plate is almost straight throughout the length of the arm, but triangular on the basal segments. A few proximal dorsal plates have a pointed proximal angle, whereas the majority of them possess a short straight proximal edge. Two arms show irregular bands of numerous elongated scales intermingled with the lateral and dorsal plates. These proximal scales are similar to dorsal disk scales. The second dorsal arm plate near the disk is narrow, elongated transversally, bar-shaped or oval. On two arms the proximalmost dorsal plates are absent for 4–5 segments, whereas in one arm small scales almost entirely conceal the first two proximal segments. Arms are weakly noded and form a low lateral ridge on which the arm spine articulations are placed. There are two or four arm spines on the most proximal segments under the disk, six to seven rather adpressed spines on the free proximal and middle segments, and five distally. Dorsal and ventral spines are similar in size. Arm spines are flattened, pointed apically. The proximal edge of the ventral arm plate is straight, whereas the distal edge is convex with a small pit in the middle. Ventral plates throughout the length of the arm are contiguous. The tentacle pores are larger and more conspicuous proximally than towards the distal end of the arm. Basally on the part of the lateral plate that encircles the tentacle pores, three flattened, apically rounded tentacle scales are placed. The tentacle scales are shorter than the arm spines, but conspicuous and large enough to cover the whole tentacle pore. Many of the tentacle scales are broken or have fallen off. Towards the distal end of the arm the number of tentacle scales is reduced to two, and the inner scale is shorter than the outer one. The ventral arm plates do not bear any tentacle scales.</paragraph>
<paragraph id="344BE802FFE45218FF11FA118621F80C" blockId="82.[151,1436,152,1970]" pageId="82" pageNumber="83">
<emphasis id="06803410FFE45218FF11FA1185C3F9A9" bold="true" box="[206,460,1533,1559]" pageId="82" pageNumber="83">
<typeStatus id="EB4F56A0FFE45218FF11FA11854FF9A9" box="[206,320,1533,1559]" pageId="82" pageNumber="83" type="paratype">Paratype</typeStatus>
variations.
</emphasis>
A 
<typeStatus id="EB4F56A0FFE45218FE2EFA118656F9A9" box="[497,601,1533,1559]" pageId="82" pageNumber="83" type="paratype">paratype</typeStatus>
, from sta. 
<quantity id="F30C45E7FFE45218FD0AFA11875BF9A9" box="[725,852,1533,1559]" metricMagnitude="-1" metricUnit="m" metricValue="8.708" pageId="82" pageNumber="83" unit="mm" value="870.8">870, 8 mm</quantity>
dd, differs from the 
<typeStatus id="EB4F56A0FFE45218FB98FA1180A2F9A9" box="[1095,1197,1533,1559]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
in having an almost undamaged irregularly-pentagonal disk, evenly covered with numerous, mostly elongated scales, which continue dorsally onto 1–2 proximal arm segments. Almost all dorsal arm plates, except for the 1–2 most proximal, have a pointed, narrowly triangular proximal edge, and are separated by lateral plates, to a small degree proximally, but up to half of the dorsal plate length distally. The pattern of the oral papillae is similar to the 
<typeStatus id="EB4F56A0FFE45218FF1CF9548521F96C" box="[195,302,1720,1746]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
, but the papillae are significantly fewer, and the curved row of oral papillae around the second tentacle pore is evident only in some interradii. Numerous spiniform teeth do not differ from the 3–4 apical papillae. Two, rarely three papillae on the adoral shield. Oral shields are more lengthened than in the 
<typeStatus id="EB4F56A0FFE45218FAF1F8EE8197F8A2" box="[1326,1432,1794,1820]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
, and the madreporite is just an inconspicuous spot within the oral shield. Most of the oral shields have a remarkable feature of one or two rather conspicuous scales overlapping the shield distally, as a continuation of the supplementary oral shield – several scales adjacent distally to the oral shield. In other characters the 
<typeStatus id="EB4F56A0FFE45218FF48F87484F3F80C" box="[151,252,1944,1970]" pageId="82" pageNumber="83" type="paratype">paratype</typeStatus>
is similar to the 
<typeStatus id="EB4F56A0FFE45218FE1FF8748624F80C" box="[448,555,1944,1970]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="608BB88AFFE55219FF48F9F0866DF900" pageId="83" pageNumber="84" startId="83.[151,255,1564,1588]" targetBox="[192,1395,194,1543]" targetPageId="83">
<paragraph id="344BE802FFE55219FF48F9F0866DF900" blockId="83.[151,1436,1564,1726]" pageId="83" pageNumber="84">
<emphasis id="06803410FFE55219FF48F9F08528F98A" bold="true" box="[151,295,1564,1588]" pageId="83" pageNumber="84">FIGURE 57.</emphasis>
Genera 
<taxonomicName id="F3F49381FFE55219FE5DF9F1860BF98A" authorityName="Verrill" authorityYear="1899" box="[386,516,1565,1588]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="83" pageNumber="84" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE55219FE5DF9F1860BF98A" box="[386,516,1565,1588]" italics="true" pageId="83" pageNumber="84">Ophioprium</emphasis>
</taxonomicName>
and 
<taxonomicName id="F3F49381FFE55219FDE7F9F186B2F98A" authorityName="Verrill" authorityYear="1899" box="[568,701,1565,1588]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="83" pageNumber="84" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE55219FDE7F9F186B2F98A" box="[568,701,1565,1588]" italics="true" pageId="83" pageNumber="84">Ophiopristis</emphasis>
</taxonomicName>
, external views. A-B, 
<taxonomicName id="F3F49381FFE55219FC70F9F18157F98A" authority="(Lyman, 1883)" baseAuthorityName="Lyman" baseAuthorityYear="1883" box="[943,1368,1565,1588]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="83" pageNumber="84" phylum="Echinodermata" rank="species" species="cervicornis">
<emphasis id="06803410FFE55219FC70F9F180BFF98A" box="[943,1200,1565,1588]" italics="true" pageId="83" pageNumber="84">Ophioprium cervicornis</emphasis>
(
<bibRefCitation id="506595F3FFE55219FB60F9F1815EF98A" author="Lyman, T." box="[1215,1361,1565,1588]" pageId="83" pageNumber="84" pagination="227 - 287" refId="ref90277" refString="Lyman, T. (1883) Report on the Ophiuroidea. Bulletin of the Museum of Comparative Zoology at Harvard College 10, 227 - 287." type="journal article" year="1883">Lyman, 1883</bibRefCitation>
)
</taxonomicName>
, holotype MCZ 2043, 10 mm dd, dorsal and ventral views respectively; C-D, 
<taxonomicName id="F3F49381FFE55219FC71F9AC813DF9E8" authority="(Lyman, 1875)" baseAuthorityName="Lyman" baseAuthorityYear="1875" box="[942,1330,1599,1623]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="83" pageNumber="84" phylum="Echinodermata" rank="species" species="hirsuta">
<emphasis id="06803410FFE55219FC71F9AC8088F9E9" box="[942,1159,1600,1623]" italics="true" pageId="83" pageNumber="84">Ophiopristis hirsuta</emphasis>
(
<bibRefCitation id="506595F3FFE55219FB48F9D38125F9E8" author="Lyman, T." box="[1175,1322,1599,1622]" pageId="83" pageNumber="84" pagination="1 - 34" refId="ref90050" refString="Lyman, T. (1875) Zoological results of the Hassler Expedition. II. Ophiuridae and Astrophytidae, including those dredged by the late Dr. William Stimpson. Ilustrated Catalogue Museum Comparative Zoology, Harvard College, 8, 1 - 34." type="journal article" year="1875">Lyman, 1875</bibRefCitation>
)
</taxonomicName>
, holotype MCZ 1950, 5.8 mm dd, dorsal and ventral views respectively; E. 
<taxonomicName id="F3F49381FFE55219FC92F98E80FBF9C7" authority="(Lyman, 1883)" baseAuthorityName="Lyman" baseAuthorityYear="1883" box="[845,1268,1634,1657]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="83" pageNumber="84" phylum="Echinodermata" rank="species" species="cervicornis">
<emphasis id="06803410FFE55219FC92F98E8042F9C7" box="[845,1101,1634,1657]" italics="true" pageId="83" pageNumber="84">Ophioprium cervicornis</emphasis>
(
<bibRefCitation id="506595F3FFE55219FB84F98E80E3F9C7" author="Lyman, T." box="[1115,1260,1634,1657]" pageId="83" pageNumber="84" pagination="227 - 287" refId="ref90277" refString="Lyman, T. (1883) Report on the Ophiuroidea. Bulletin of the Museum of Comparative Zoology at Harvard College 10, 227 - 287." type="journal article" year="1883">Lyman, 1883</bibRefCitation>
)
</taxonomicName>
, holotype MCZ 2043, arm spine articulations, proximal segments; F. 
<taxonomicName id="F3F49381FFE55219FD15F9698047F922" authority="(Lyman, 1875)" baseAuthorityName="Lyman" baseAuthorityYear="1875" box="[714,1096,1669,1692]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="83" pageNumber="84" phylum="Echinodermata" rank="species" species="hirsuta">
<emphasis id="06803410FFE55219FD15F96987AFF922" box="[714,928,1669,1692]" italics="true" pageId="83" pageNumber="84">Ophiopristis hirsuta</emphasis>
(
<bibRefCitation id="506595F3FFE55219FC70F969804FF922" author="Lyman, T." box="[943,1088,1669,1692]" pageId="83" pageNumber="84" pagination="1 - 34" refId="ref90050" refString="Lyman, T. (1875) Zoological results of the Hassler Expedition. II. Ophiuridae and Astrophytidae, including those dredged by the late Dr. William Stimpson. Ilustrated Catalogue Museum Comparative Zoology, Harvard College, 8, 1 - 34." type="journal article" year="1875">Lyman, 1875</bibRefCitation>
)
</taxonomicName>
, paratype MCZ 1951, 5 mm dd, ventral view. Photos: Alexander Martynov.
</paragraph>
</caption>
<paragraph id="344BE802FFE5521EFF11F91E872CFED2" blockId="83.[151,1436,1778,2028]" lastBlockId="84.[151,1436,152,775]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
Another 
<typeStatus id="EB4F56A0FFE55219FEE9F91E8591F8B2" box="[310,414,1778,1804]" pageId="83" pageNumber="84" type="paratype">paratype</typeStatus>
, also from sta. 870, 5.5 mm dd, differs from the 
<typeStatus id="EB4F56A0FFE55219FC09F91E8032F8B2" box="[982,1085,1778,1804]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
and other 
<typeStatus id="EB4F56A0FFE55219FB66F91E8111F8B2" box="[1209,1310,1778,1804]" pageId="83" pageNumber="84" type="paratype">paratype</typeStatus>
in size, an irregularly pentagonal disk, evenly covered with numerous polygonal and oval scales, which are continued dorsally on 2–3 proximal arm segments. The pattern of the oral papillae is also distinct from the 
<typeStatus id="EB4F56A0FFE55219FADCF8D18166F8E9" box="[1283,1385,1853,1879]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
and from the 
<typeStatus id="EB4F56A0FFE55219FEDBF88E8566F8C2" box="[260,361,1890,1916]" pageId="83" pageNumber="84" type="paratype">paratype</typeStatus>
from the same station (870), but these differences clearly reflect the juvenile nature of the present specimen. The papillae are significantly smaller in number, and the curved row of oral papillae around the second tentacle pore is indistinct and can be traced only in two interradii. One or two distalmost oral papillae are block shaped. At least one papilla on the adoral shields is present in one interradius, whereas in the other interradii it has most probably fallen off. Oral shields are more elongated in length than in the 
<typeStatus id="EB4F56A0FFE2521EFF48FF51850EFF69" box="[151,257,189,215]" pageId="84" pageNumber="85" type="holotype">holotype</typeStatus>
, and the madreporite is just an inconspicuous spot within one oral shield. Most of the oral shields are overlapped by a conspicuous scale distally. On the most proximal lateral arm plates under the disk there are 2 spines, on the proximal lateral plates 4–5 spines, towards the middle 5–6. Spines are flattened, all similar in size. Tentacle scales are similar to the 
<typeStatus id="EB4F56A0FFE2521EFD83FEC186C9FEF9" box="[604,710,301,327]" pageId="84" pageNumber="85" type="holotype">holotype</typeStatus>
, but many are broken or have fallen off. In the middle part of the arm the number of tentacle scales is reduced to two.
</paragraph>
<paragraph id="344BE802FFE2521EFF18FE9B8621FCB9" blockId="84.[151,1436,152,775]" pageId="84" pageNumber="85">
<emphasis id="06803410FFE2521EFF18FE9B86C5FE2F" bold="true" box="[199,714,375,401]" pageId="84" pageNumber="85">Internal and microstructural characters.</emphasis>
Radial shield and genital plate articulate very tightly, unlike most extant ophiuroids, and have the appearance of an entire plate broken by a fissure (
<figureCitation id="ACCFF487FFE2521EFB69FE718158FE09" box="[1206,1367,413,439]" captionStart="FIGURE 4" captionStartId="13.[151,256,1339,1363]" captionTargetBox="[282,1312,194,1330]" captionTargetId="figure-434@13.[282,1312,194,1330]" captionTargetPageId="13" captionText="FIGURE 4. Genital plates and radial shields of various taxa of the families Asteronychidae, Gorgonocephalidae and Ophiomyxidae. A. Gorgonocephalus arcticus Leach, 1819, ZMMU D-613, 39 mm dd, adradial and abradial genital plates in situ, showing articulation surface, dorsal view; B. same, articulation surface of the radial shield, ventral view; C. Asteronyx loveni Müller &amp; Troschel, 1842, ZMMU D-833, 17.5 mm dd, articulation surface of the adradial genital plate, dorsal view; D. same, articulation surface of the radial shield, ventral view. E. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, articulation surface of the adradial genital plate, dorsal view; F. same, radial shield showing articulation surface, ventral view; G. same, articulating adradial genital plate and radial shield, dorsal view; H. Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, dissected disk showing most proximal vertebrae and absence of the both genital plates and radial shield, dorsal view; I. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, adradial and abradial genital plates in situ, ventral view; J. Ophioplexa condita gen. et sp. nov., paratype ca. 17 mm dd, ZMMU D-792, dissected disk showing articulating adradial genital plate and radial shield in situ, dorsal view; K. same, separate articulating adradial genital plate and radial shield, dorsal view; L. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, adradial genital plate, dorsal view; M. Ophiologimus cf. secundus Koehler, 1914, 4 mm dd, ZMMU D-814, adradial genital plate, dorsal view; N. Ophioplexa condita gen. et sp. nov., separate adradial genital plate, dorsal view; O. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, adradial and abradial genital plates, dorsal view; P. Ophioplexa condita gen. et sp. nov., separate articulating adradial genital plate and radial shield, dorsal view; Q. articulation surface of the adradial genital plate, dorsal view; R. radial shield, ventral view, showing distally the distinctive excavated articulation surface; S. articulation surface of the radial shield, ventral view; T. Ophiocymbium ninae sp. nov., paratype 5 mm dd, radial shield, ventral view; U. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, same, articulation surface of the radial shield, proximal view; Scale bars: A– B,1 mm; C–D, 300 µ; E–F, 100 µ; G, 200 µ; H, 300 µ; I, 200 µ; J, 1000 µ; K, 300 µ; L, 100 µ; M, 300 µ; N, 300 µ; O, 200 µ; P, 100 µ; Q–T, 100 µ; U, 200 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 4J, K, P</figureCitation>
). The articulation surface of the radial shield is distinctly excavated ventrally, with a distal groove (
<figureCitation id="ACCFF487FFE2521EFB06FE2E8158FE62" box="[1241,1367,450,476]" captionStart="FIGURE 4" captionStartId="13.[151,256,1339,1363]" captionTargetBox="[282,1312,194,1330]" captionTargetId="figure-434@13.[282,1312,194,1330]" captionTargetPageId="13" captionText="FIGURE 4. Genital plates and radial shields of various taxa of the families Asteronychidae, Gorgonocephalidae and Ophiomyxidae. A. Gorgonocephalus arcticus Leach, 1819, ZMMU D-613, 39 mm dd, adradial and abradial genital plates in situ, showing articulation surface, dorsal view; B. same, articulation surface of the radial shield, ventral view; C. Asteronyx loveni Müller &amp; Troschel, 1842, ZMMU D-833, 17.5 mm dd, articulation surface of the adradial genital plate, dorsal view; D. same, articulation surface of the radial shield, ventral view. E. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, articulation surface of the adradial genital plate, dorsal view; F. same, radial shield showing articulation surface, ventral view; G. same, articulating adradial genital plate and radial shield, dorsal view; H. Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, dissected disk showing most proximal vertebrae and absence of the both genital plates and radial shield, dorsal view; I. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, adradial and abradial genital plates in situ, ventral view; J. Ophioplexa condita gen. et sp. nov., paratype ca. 17 mm dd, ZMMU D-792, dissected disk showing articulating adradial genital plate and radial shield in situ, dorsal view; K. same, separate articulating adradial genital plate and radial shield, dorsal view; L. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, adradial genital plate, dorsal view; M. Ophiologimus cf. secundus Koehler, 1914, 4 mm dd, ZMMU D-814, adradial genital plate, dorsal view; N. Ophioplexa condita gen. et sp. nov., separate adradial genital plate, dorsal view; O. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, adradial and abradial genital plates, dorsal view; P. Ophioplexa condita gen. et sp. nov., separate articulating adradial genital plate and radial shield, dorsal view; Q. articulation surface of the adradial genital plate, dorsal view; R. radial shield, ventral view, showing distally the distinctive excavated articulation surface; S. articulation surface of the radial shield, ventral view; T. Ophiocymbium ninae sp. nov., paratype 5 mm dd, radial shield, ventral view; U. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, same, articulation surface of the radial shield, proximal view; Scale bars: A– B,1 mm; C–D, 300 µ; E–F, 100 µ; G, 200 µ; H, 300 µ; I, 200 µ; J, 1000 µ; K, 300 µ; L, 100 µ; M, 300 µ; N, 300 µ; O, 200 µ; P, 100 µ; Q–T, 100 µ; U, 200 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 4R, S</figureCitation>
). The articulation surface of the adradial genital plate lacks distinct condyles (
<figureCitation id="ACCFF487FFE2521EFC3EFE04804FFDBC" box="[993,1088,488,514]" captionStart="FIGURE 4" captionStartId="13.[151,256,1339,1363]" captionTargetBox="[282,1312,194,1330]" captionTargetId="figure-434@13.[282,1312,194,1330]" captionTargetPageId="13" captionText="FIGURE 4. Genital plates and radial shields of various taxa of the families Asteronychidae, Gorgonocephalidae and Ophiomyxidae. A. Gorgonocephalus arcticus Leach, 1819, ZMMU D-613, 39 mm dd, adradial and abradial genital plates in situ, showing articulation surface, dorsal view; B. same, articulation surface of the radial shield, ventral view; C. Asteronyx loveni Müller &amp; Troschel, 1842, ZMMU D-833, 17.5 mm dd, articulation surface of the adradial genital plate, dorsal view; D. same, articulation surface of the radial shield, ventral view. E. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-794, 9.5 mm dd, articulation surface of the adradial genital plate, dorsal view; F. same, radial shield showing articulation surface, ventral view; G. same, articulating adradial genital plate and radial shield, dorsal view; H. Ophiocymbium tanyae sp. nov., paratype 11 mm dd, ZMMU D-801, dissected disk showing most proximal vertebrae and absence of the both genital plates and radial shield, dorsal view; I. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, adradial and abradial genital plates in situ, ventral view; J. Ophioplexa condita gen. et sp. nov., paratype ca. 17 mm dd, ZMMU D-792, dissected disk showing articulating adradial genital plate and radial shield in situ, dorsal view; K. same, separate articulating adradial genital plate and radial shield, dorsal view; L. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, adradial genital plate, dorsal view; M. Ophiologimus cf. secundus Koehler, 1914, 4 mm dd, ZMMU D-814, adradial genital plate, dorsal view; N. Ophioplexa condita gen. et sp. nov., separate adradial genital plate, dorsal view; O. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, adradial and abradial genital plates, dorsal view; P. Ophioplexa condita gen. et sp. nov., separate articulating adradial genital plate and radial shield, dorsal view; Q. articulation surface of the adradial genital plate, dorsal view; R. radial shield, ventral view, showing distally the distinctive excavated articulation surface; S. articulation surface of the radial shield, ventral view; T. Ophiocymbium ninae sp. nov., paratype 5 mm dd, radial shield, ventral view; U. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, same, articulation surface of the radial shield, proximal view; Scale bars: A– B,1 mm; C–D, 300 µ; E–F, 100 µ; G, 200 µ; H, 300 µ; I, 200 µ; J, 1000 µ; K, 300 µ; L, 100 µ; M, 300 µ; N, 300 µ; O, 200 µ; P, 100 µ; Q–T, 100 µ; U, 200 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Fig. 4Q</figureCitation>
). The abradial genital scale is absent. Jaws are slightly elongated (
<figureCitation id="ACCFF487FFE2521EFD9BFDE186D8FD99" box="[580,727,525,551]" captionStart="FIGURE 56" captionStartId="81.[151,255,1449,1473]" captionTargetBox="[188,1398,194,1427]" captionTargetId="figure-315@81.[188,1398,194,1428]" captionTargetPageId="81" captionText="FIGURE 56. Ophioplexa condita gen. nov., sp. nov. Arm spine articulations, spines, tentacle scales, dental plates and jaws of a paratype of ca. 17 mm dd, ZMMU D-792. A. two proximal arm segments adjoining disk; B. same, showing hollow spines; C. ventral view of the artciulations of the most proximal segments under the disk in situ including attaching area of the tentacle scales; D. separate lateral arm plate of the most proximal segments showing distinct shape of the ventralmost ridge; E. same, showing variability of following articulation; F. articulations of a middle segment; G. spine of a middle segment; H. separate lateral arm plate of the proximal segment showing more typical pattern of the articulation; I. a separate tentacle scale of the most proximal segment; J. same, basal part showing a small opening, presumably for nerve; K. basal part of a spine showing an opening for nerve; L. dental plate with several teeth; M, N, dental plates; O. jaw, abradial view; P. jaw, adradial view. Scale bars: A, 300; B, 100 µ; C, 300 µ; D, 100 µ; E, 100 µ; F,G, 100 µ; H, 300 µ; I,J, 100 µ; K, 30 µ; L–P, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 56O, P</figureCitation>
). Adradial sides of the jaws distally bear few sharp straight folds (
<figureCitation id="ACCFF487FFE2521EFF3DFDDE8544FDF2" box="[226,331,562,588]" captionStart="FIGURE 56" captionStartId="81.[151,255,1449,1473]" captionTargetBox="[188,1398,194,1427]" captionTargetId="figure-315@81.[188,1398,194,1428]" captionTargetPageId="81" captionText="FIGURE 56. Ophioplexa condita gen. nov., sp. nov. Arm spine articulations, spines, tentacle scales, dental plates and jaws of a paratype of ca. 17 mm dd, ZMMU D-792. A. two proximal arm segments adjoining disk; B. same, showing hollow spines; C. ventral view of the artciulations of the most proximal segments under the disk in situ including attaching area of the tentacle scales; D. separate lateral arm plate of the most proximal segments showing distinct shape of the ventralmost ridge; E. same, showing variability of following articulation; F. articulations of a middle segment; G. spine of a middle segment; H. separate lateral arm plate of the proximal segment showing more typical pattern of the articulation; I. a separate tentacle scale of the most proximal segment; J. same, basal part showing a small opening, presumably for nerve; K. basal part of a spine showing an opening for nerve; L. dental plate with several teeth; M, N, dental plates; O. jaw, abradial view; P. jaw, adradial view. Scale bars: A, 300; B, 100 µ; C, 300 µ; D, 100 µ; E, 100 µ; F,G, 100 µ; H, 300 µ; I,J, 100 µ; K, 30 µ; L–P, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Fig. 56P</figureCitation>
). The dental plate is large with rounded sockets and irregular folds (
<figureCitation id="ACCFF487FFE2521EFBB0FDDE8104FDF2" box="[1135,1291,562,588]" captionStart="FIGURE 56" captionStartId="81.[151,255,1449,1473]" captionTargetBox="[188,1398,194,1427]" captionTargetId="figure-315@81.[188,1398,194,1428]" captionTargetPageId="81" captionText="FIGURE 56. Ophioplexa condita gen. nov., sp. nov. Arm spine articulations, spines, tentacle scales, dental plates and jaws of a paratype of ca. 17 mm dd, ZMMU D-792. A. two proximal arm segments adjoining disk; B. same, showing hollow spines; C. ventral view of the artciulations of the most proximal segments under the disk in situ including attaching area of the tentacle scales; D. separate lateral arm plate of the most proximal segments showing distinct shape of the ventralmost ridge; E. same, showing variability of following articulation; F. articulations of a middle segment; G. spine of a middle segment; H. separate lateral arm plate of the proximal segment showing more typical pattern of the articulation; I. a separate tentacle scale of the most proximal segment; J. same, basal part showing a small opening, presumably for nerve; K. basal part of a spine showing an opening for nerve; L. dental plate with several teeth; M, N, dental plates; O. jaw, abradial view; P. jaw, adradial view. Scale bars: A, 300; B, 100 µ; C, 300 µ; D, 100 µ; E, 100 µ; F,G, 100 µ; H, 300 µ; I,J, 100 µ; K, 30 µ; L–P, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 56M, N</figureCitation>
). Arm spine articulations compressed transversally (
<figureCitation id="ACCFF487FFE2521EFD5BFDB4872EFDCC" box="[644,801,600,626]" captionStart="FIGURE 55" captionStartId="80.[151,258,1764,1788]" captionTargetBox="[209,1364,194,1740]" captionTargetId="figure-174@80.[209,1364,194,1740]" captionTargetPageId="80" captionText="FIGURE 55. Ophioplexa condita gen. nov., sp. nov. and Ophiocymbium ninae sp. nov., details. A. Ophioplexa condita gen. nov., sp. nov., paratype 19 mm dd, ZMMU D-791, disk ventral view; B. same oral frame showing clusters of the ventral teeth (apical papillae); C. same, one interradius showing apical, oral and adoral shield papillae; D-F, same, most proximal arm segments showing tentacle scales and spines; G-H, arm spine articulations, distal segments; I. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-803, details, part of the oral frame and ventral arm side; J. same, jaw and oral papillae, one interradius; K. ventral arm view showing spines and tentacle scales; L. most proximal segments showing details of tentacle scales and spines; M-N, arm spine articulations, proximal segments; Scale bars: A– B, 1 mm; C–D, 500 µ; E–F, 200 µ; G–H, 100 µ; I, 1 mm; J–N, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 55G–H</figureCitation>
; 
<figureCitation id="ACCFF487FFE2521EFCEFFDB48780FDCC" box="[816,911,600,626]" captionStart="FIGURE 56" captionStartId="81.[151,255,1449,1473]" captionTargetBox="[188,1398,194,1427]" captionTargetId="figure-315@81.[188,1398,194,1428]" captionTargetPageId="81" captionText="FIGURE 56. Ophioplexa condita gen. nov., sp. nov. Arm spine articulations, spines, tentacle scales, dental plates and jaws of a paratype of ca. 17 mm dd, ZMMU D-792. A. two proximal arm segments adjoining disk; B. same, showing hollow spines; C. ventral view of the artciulations of the most proximal segments under the disk in situ including attaching area of the tentacle scales; D. separate lateral arm plate of the most proximal segments showing distinct shape of the ventralmost ridge; E. same, showing variability of following articulation; F. articulations of a middle segment; G. spine of a middle segment; H. separate lateral arm plate of the proximal segment showing more typical pattern of the articulation; I. a separate tentacle scale of the most proximal segment; J. same, basal part showing a small opening, presumably for nerve; K. basal part of a spine showing an opening for nerve; L. dental plate with several teeth; M, N, dental plates; O. jaw, abradial view; P. jaw, adradial view. Scale bars: A, 300; B, 100 µ; C, 300 µ; D, 100 µ; E, 100 µ; F,G, 100 µ; H, 300 µ; I,J, 100 µ; K, 30 µ; L–P, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">56A–H</figureCitation>
). Some proximal articulations may have a distinct nerve opening (
<figureCitation id="ACCFF487FFE2521EFE6AFD918642FD29" box="[437,589,637,663]" captionStart="FIGURE 56" captionStartId="81.[151,255,1449,1473]" captionTargetBox="[188,1398,194,1427]" captionTargetId="figure-315@81.[188,1398,194,1428]" captionTargetPageId="81" captionText="FIGURE 56. Ophioplexa condita gen. nov., sp. nov. Arm spine articulations, spines, tentacle scales, dental plates and jaws of a paratype of ca. 17 mm dd, ZMMU D-792. A. two proximal arm segments adjoining disk; B. same, showing hollow spines; C. ventral view of the artciulations of the most proximal segments under the disk in situ including attaching area of the tentacle scales; D. separate lateral arm plate of the most proximal segments showing distinct shape of the ventralmost ridge; E. same, showing variability of following articulation; F. articulations of a middle segment; G. spine of a middle segment; H. separate lateral arm plate of the proximal segment showing more typical pattern of the articulation; I. a separate tentacle scale of the most proximal segment; J. same, basal part showing a small opening, presumably for nerve; K. basal part of a spine showing an opening for nerve; L. dental plate with several teeth; M, N, dental plates; O. jaw, abradial view; P. jaw, adradial view. Scale bars: A, 300; B, 100 µ; C, 300 µ; D, 100 µ; E, 100 µ; F,G, 100 µ; H, 300 µ; I,J, 100 µ; K, 30 µ; L–P, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 56D–E</figureCitation>
). Few ventralmost articulations on the proximalmost segments have a distinctive slit-shaped appearance (
<figureCitation id="ACCFF487FFE2521EFDE5FD4E86C0FD02" box="[570,719,674,700]" captionStart="FIGURE 56" captionStartId="81.[151,255,1449,1473]" captionTargetBox="[188,1398,194,1427]" captionTargetId="figure-315@81.[188,1398,194,1428]" captionTargetPageId="81" captionText="FIGURE 56. Ophioplexa condita gen. nov., sp. nov. Arm spine articulations, spines, tentacle scales, dental plates and jaws of a paratype of ca. 17 mm dd, ZMMU D-792. A. two proximal arm segments adjoining disk; B. same, showing hollow spines; C. ventral view of the artciulations of the most proximal segments under the disk in situ including attaching area of the tentacle scales; D. separate lateral arm plate of the most proximal segments showing distinct shape of the ventralmost ridge; E. same, showing variability of following articulation; F. articulations of a middle segment; G. spine of a middle segment; H. separate lateral arm plate of the proximal segment showing more typical pattern of the articulation; I. a separate tentacle scale of the most proximal segment; J. same, basal part showing a small opening, presumably for nerve; K. basal part of a spine showing an opening for nerve; L. dental plate with several teeth; M, N, dental plates; O. jaw, abradial view; P. jaw, adradial view. Scale bars: A, 300; B, 100 µ; C, 300 µ; D, 100 µ; E, 100 µ; F,G, 100 µ; H, 300 µ; I,J, 100 µ; K, 30 µ; L–P, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 56C–E</figureCitation>
). Vertebrae short, distinctly keeled and with well developed zygospondylous articulation (
<figureCitation id="ACCFF487FFE2521EFDDCFD248692FD5C" box="[515,669,712,738]" captionStart="FIGURE 17" captionStartId="27.[151,255,1036,1060]" captionTargetBox="[161,1426,186,1015]" captionTargetId="figure-399@27.[161,1426,186,1015]" captionTargetPageId="27" captionText="FIGURE 17. Vertebrae of various taxa of the family Ophiomyxidae. A. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, proximal segments, dorsal and ventral views; B. same, distal segments, dorsal and ventral views; C. same, proximal segments, proximal view; D. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, 10.5 mm dd, proximal segments, dorsal view; E. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, proximal segments, dorsal view; F. Ophioscolex glacialis, proximal segments, distal view; G. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, proximal segments, proximal view; H. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, ventral view; I. same, proximal view; J. same, distal view; K. Ophiomyxa pentagona, ZMMU D-553, 21 mm dd, distal view; L. Ophioplexa condita gen. et sp. nov., holotype ca. 18 mm dd, ZMMU D-788, proximal segments, dorsal view; M. same, ventral view; N. same, proximal view; O. same, distal view; P. Ophiocymbium tanyae sp. nov., holotype 9.5 mm dd, ZMMU D-799, proximal segments, dorsal view; Q. same, ventral view; R. same, distal view; S. same, proximal view. Scale bars: A, 1000 µ; B, C, 200 µ, D, 200 µ; E, 300 µ; F, 200 µ; G, 200 µ; H, 300 µ; I, 100 µ; J, 100 µ; K, 1000 µ; L–S, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 17L–O</figureCitation>
). Vertebral dorsal median groove almost indistinct (
<figureCitation id="ACCFF487FFE2521EFAC0FD248181FD5C" box="[1311,1422,712,738]" captionStart="FIGURE 17" captionStartId="27.[151,255,1036,1060]" captionTargetBox="[161,1426,186,1015]" captionTargetId="figure-399@27.[161,1426,186,1015]" captionTargetPageId="27" captionText="FIGURE 17. Vertebrae of various taxa of the family Ophiomyxidae. A. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, proximal segments, dorsal and ventral views; B. same, distal segments, dorsal and ventral views; C. same, proximal segments, proximal view; D. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, 10.5 mm dd, proximal segments, dorsal view; E. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, proximal segments, dorsal view; F. Ophioscolex glacialis, proximal segments, distal view; G. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, proximal segments, proximal view; H. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, ventral view; I. same, proximal view; J. same, distal view; K. Ophiomyxa pentagona, ZMMU D-553, 21 mm dd, distal view; L. Ophioplexa condita gen. et sp. nov., holotype ca. 18 mm dd, ZMMU D-788, proximal segments, dorsal view; M. same, ventral view; N. same, proximal view; O. same, distal view; P. Ophiocymbium tanyae sp. nov., holotype 9.5 mm dd, ZMMU D-799, proximal segments, dorsal view; Q. same, ventral view; R. same, distal view; S. same, proximal view. Scale bars: A, 1000 µ; B, C, 200 µ, D, 200 µ; E, 300 µ; F, 200 µ; G, 200 µ; H, 300 µ; I, 100 µ; J, 100 µ; K, 1000 µ; L–S, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Fig. 17L</figureCitation>
). Podial basins are large (
<figureCitation id="ACCFF487FFE2521EFE70FD01862EFCB9" box="[431,545,749,775]" captionStart="FIGURE 17" captionStartId="27.[151,255,1036,1060]" captionTargetBox="[161,1426,186,1015]" captionTargetId="figure-399@27.[161,1426,186,1015]" captionTargetPageId="27" captionText="FIGURE 17. Vertebrae of various taxa of the family Ophiomyxidae. A. Ophioscolex glacialis Müller &amp; Troschel, 1842, ZMMU D-843, 18 mm dd, proximal segments, dorsal and ventral views; B. same, distal segments, dorsal and ventral views; C. same, proximal segments, proximal view; D. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, 10.5 mm dd, proximal segments, dorsal view; E. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, proximal segments, dorsal view; F. Ophioscolex glacialis, proximal segments, distal view; G. Ophiocymbium cavernosum Lyman, 1880, ZMMU D-795, proximal segments, proximal view; H. Ophiocymbium ninae sp. nov., paratype 5 mm dd, ZMMU D-804, ventral view; I. same, proximal view; J. same, distal view; K. Ophiomyxa pentagona, ZMMU D-553, 21 mm dd, distal view; L. Ophioplexa condita gen. et sp. nov., holotype ca. 18 mm dd, ZMMU D-788, proximal segments, dorsal view; M. same, ventral view; N. same, proximal view; O. same, distal view; P. Ophiocymbium tanyae sp. nov., holotype 9.5 mm dd, ZMMU D-799, proximal segments, dorsal view; Q. same, ventral view; R. same, distal view; S. same, proximal view. Scale bars: A, 1000 µ; B, C, 200 µ, D, 200 µ; E, 300 µ; F, 200 µ; G, 200 µ; H, 300 µ; I, 100 µ; J, 100 µ; K, 1000 µ; L–S, 300 µ. Photos: Alexander Martynov." pageId="84" pageNumber="85">Fig. 17M</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>