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<mods:title id="5BFF80A7EF6561F65C2C9F3BFC4B63EA">Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae 2697</mods:title>
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<paragraph id="344BE802FFE2521EFF48FCB1862BFCC9" blockId="84.[151,548,861,922]" box="[151,548,861,887]" pageId="84" pageNumber="85">
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<emphasis id="06803410FFE2521EFF48FCB1862BFCC9" bold="true" box="[151,548,861,887]" pageId="84" pageNumber="85">
Genus
<taxonomicName id="F3F49381FFE2521EFF31FCB1862BFCC9" ID-CoL="7PQZC" authority="Verrill, 1899" authorityName="Verrill" authorityYear="1899" box="[238,548,861,887]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="84" pageNumber="85" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE2521EFF31FCB1858DFCC9" bold="true" box="[238,386,861,887]" italics="true" pageId="84" pageNumber="85">Ophiopristis</emphasis>
<bibRefCitation id="506595F3FFE2521EFE56FCB1862BFCC9" author="Verrill, A. E." box="[393,548,861,887]" pageId="84" pageNumber="85" pagination="1 - 86" refId="ref92761" refString="Verrill, A. E. (1899) Report on the Ophiuroidea; collected by the Bahama Expedition from the University of Iowa in 1893. Bulletin from the Laboratories of Natural History of the State University of Iowa 5, 1 - 86." type="journal article" year="1899">Verrill, 1899</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="7CEEBB89FFE2521EFF48FC6C85CDFC24" box="[151,450,896,922]" pageId="84" pageNumber="85" type="description">
<paragraph id="344BE802FFE2521EFF48FC6C85CDFC24" blockId="84.[151,548,861,922]" box="[151,450,896,922]" pageId="84" pageNumber="85">
<figureCitation id="ACCFF487FFE2521EFF48FC6C8563FC24" box="[151,364,896,922]" captionStart="FIGURE 57" captionStartId="83.[151,255,1564,1588]" captionTargetBox="[192,1395,194,1543]" captionTargetId="figure-210@83.[192,1395,194,1543]" captionTargetPageId="83" captionText="FIGURE 57. Genera Ophioprium and Ophiopristis, external views. A-B, Ophioprium cervicornis (Lyman, 1883), holotype MCZ 2043, 10 mm dd, dorsal and ventral views respectively; C-D, Ophiopristis hirsuta (Lyman, 1875), holotype MCZ 1950, 5.8 mm dd, dorsal and ventral views respectively; E. Ophioprium cervicornis (Lyman, 1883), holotype MCZ 2043, arm spine articulations, proximal segments; F. Ophiopristis hirsuta (Lyman, 1875), paratype MCZ 1951, 5 mm dd, ventral view. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figures 57CD, F</figureCitation>
;
<figureCitation id="ACCFF487FFE2521EFEA6FC6C85CDFC24" box="[377,450,896,922]" captionStart-0="FIGURE 58" captionStart-1="FIGURE 59" captionStart-2="FIGURE 60" captionStartId-0="85.[151,255,1371,1395]" captionStartId-1="86.[151,255,1557,1581]" captionStartId-2="87.[151,255,1654,1678]" captionTargetBox-0="[158,1428,343,1349]" captionTargetBox-1="[158,1428,647,1535]" captionTargetBox-2="[158,1428,197,1633]" captionTargetId-0="figure-246@85.[158,1428,343,1349]" captionTargetId-1="figure-290@86.[158,1428,647,1535]" captionTargetId-2="figure-194@87.[158,1428,197,1633]" captionTargetPageId-0="85" captionTargetPageId-1="86" captionTargetPageId-2="87" captionText-0="FIGURE 58. Ophiopristis hirsuta (Lyman, 1875), USNM 6429, 10 mm dd, details. A. arm spine articulations, proximal segments; B. same, details; C. hollow spine transversally sectioned; D. spine; E. same, ventral view; F. proximal vertebra, dorsal view; G. same, proximal view; H. same, distal view. Scale bars: AB, FH, 300 µ; C, E, 100 µ; D, 1 mm. Photos: Alexander Martynov." captionText-1="FIGURE 59. Ophiopristis luctosa (Koehler, 1904), MNHN EcOs 22978, 4.5 mm dd, external view and details. A. dorsal view; B. ventral view; C. lateral arm plate and spine articulations, proximal segments; DE, arm spine articulations, details; F. hollow spine transversally sectioned; G. spine, proximal segments; H. proximal vertebra, dorsal view; I. same, ventral view; J. same, proximal view; K. same, distal view; L. spine, ventral view. Scale bars: C-E, H-K, 100 µ; F, 30 µ; L, 300 µ. AB Tatiana Korshunova; CM Alexander Martynov." captionText-2="FIGURE 60. Ophiopristis procera (Koehler, 1904), syntype MNHN EcOs 20406, 12 mm dd, external view and details. A. dorsal view; B. ventral view; C. proximal arm segments, ventral view showing oval tentacle scales; D. same, dorsal view, showing arm spine articulations and spines; EF, lateral arm plate and spine articulations, proximal segments; G. same, details; H. distal vertebra, dorsal view; I. same, ventral view; J. proximal vertebra, dorsal view; K. same, distal view; L. hollow proximal spine transversally sectioned; M. proximal spine, ventral view; N. proximal spine. Scale bars: EI, 100 µ; J, N, 1 mm; K, M 300 µ; L, 100 µ. AC Tatiana Korshunova; DN Alexander Martynov." pageId="84" pageNumber="85">5860</figureCitation>
</paragraph>
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<subSubSection id="7CEEBB89FFE2521EFF48FC2685F5FC5F" box="[151,506,970,993]" pageId="84" pageNumber="85" type="reference_group">
<paragraph id="344BE802FFE2521EFF48FC2685F5FC5F" blockId="84.[151,929,970,1025]" box="[151,506,970,993]" pageId="84" pageNumber="85">
<treatmentCitationGroup id="14E4CF2CFFE2521EFF48FC2685F5FC5F" box="[151,506,970,993]" pageId="84" pageNumber="85">
<treatmentCitation id="B555CE13FFE2521EFF48FC2685DAFC5F" author="Verrill, A. E." box="[151,469,970,993]" page="39" pageId="84" pageNumber="85" year="1899">
<taxonomicName id="F3F49381FFE2521EFF48FC2685DAFC5F" ID-CoL="7PQZC" authority="Verrill, 1899: 39" authorityName="Verrill" authorityPageNumber="39" authorityYear="1899" box="[151,469,970,993]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="84" pageNumber="85" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE2521EFF48FC268513FC5F" box="[151,284,970,993]" italics="true" pageId="84" pageNumber="85">Ophiopristis</emphasis>
<bibRefCitation id="506595F3FFE2521EFEFCFC2685DAFC5F" author="Verrill, A. E." box="[291,469,970,993]" pageId="84" pageNumber="85" pagination="1 - 86" refId="ref92761" refString="Verrill, A. E. (1899) Report on the Ophiuroidea; collected by the Bahama Expedition from the University of Iowa in 1893. Bulletin from the Laboratories of Natural History of the State University of Iowa 5, 1 - 86." type="journal article" year="1899">Verrill, 1899: 39</bibRefCitation>
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, 40
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<subSubSection id="7CEEBB89FFE2521EFF48FC0687AEFBBF" box="[151,929,1002,1025]" pageId="84" pageNumber="85" type="materials_examined">
<paragraph id="344BE802FFE2521EFF48FC0687AEFBBF" blockId="84.[151,929,970,1025]" box="[151,929,1002,1025]" pageId="84" pageNumber="85">
<typeStatus id="EB4F56A0FFE2521EFF48FC0684C3FBBF" box="[151,204,1002,1025]" pageId="84" pageNumber="85">Type</typeStatus>
species:
<taxonomicName id="F3F49381FFE2521EFEF2FC0686AEFBBF" authority="Lyman, 1875" authorityName="Lyman" authorityYear="1875" box="[301,673,1002,1025]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="84" pageNumber="85" phylum="Echinodermata" rank="species" species="hirsuta">
<emphasis id="06803410FFE2521EFEF2FC068605FBBF" box="[301,522,1002,1025]" italics="true" pageId="84" pageNumber="85">Ophiacantha hirsuta</emphasis>
<bibRefCitation id="506595F3FFE2521EFDCDFC0686AEFBBF" author="Lyman, T." box="[530,673,1002,1025]" pageId="84" pageNumber="85" pagination="1 - 34" refId="ref90050" refString="Lyman, T. (1875) Zoological results of the Hassler Expedition. II. Ophiuridae and Astrophytidae, including those dredged by the late Dr. William Stimpson. Ilustrated Catalogue Museum Comparative Zoology, Harvard College, 8, 1 - 34." type="journal article" year="1875">Lyman, 1875</bibRefCitation>
</taxonomicName>
, by original designation
</paragraph>
</subSubSection>
<subSubSection id="7CEEBB89FFE2521EFF48FBC386F6F8B1" pageId="84" pageNumber="85" type="diagnosis">
<paragraph id="344BE802FFE2521EFF48FBC386F6F8B1" blockId="84.[151,1436,1071,2031]" pageId="84" pageNumber="85">
<emphasis id="06803410FFE2521EFF48FBC38517FBF7" bold="true" box="[151,280,1071,1097]" pageId="84" pageNumber="85">Diagnosis.</emphasis>
The disk is covered with numerous small, uniform scales; a skin layer is not evident. The dorsal disk surface is covered with long, densely spaced spinelets. The radial shields are well-defined, elongated, but concealed under the disk plates. The adradial and abradial genital plates are well-defined. The genital slits are long and conspicuous, genital plates border approximately half of the slits. The articulation surface of the radial shield is a broad, low elevation. The jaw bears numerous short conical oral papillae similar in shape to the single (or double) apical papillae. The adoral shield papillae and adjacent oral papillae are slightly longer than other oral papillae. The half-jaws are relatively low. The adradial sides of the half-jaws distally bear a few convoluted folds. The dental plate is elongated without folds and with few elongate narrow slit-shaped sockets, alternately placed. The sockets for the apical papillae are similar in shape to all others. The teeth are broad, conical to rectangular. There are 12 ventralmost teeth. Generally all teeth are similar in shape, but the dorsalmost teeth are usually longer. The oral shield is broadly arrow-shaped with a short distal lobe. The adoral shields have distal wings, proximally tapered. Dorsal and ventral arm plates are well developed. The arm spine articulations are placed at a small angle in relation to the lateral plate. The muscle opening is larger than the nerve opening. There is a volute-shaped perforated lobe occupying the dorsal and distal part of the articulation. The sigmoidal fold is well-defined. The proximal edge of the spine articulation is entire but connected with the main part of the lateral arm plate by a short ridge. The arm spines are relatively long, rounded, hollow over almost their entire length, not hooked distally. Tentacle scales are well-defined, oval or lanceolate, single or double. The tentacle pores of moderate size. The vertebrae have a narrow keel that is distally abruptly truncated; the dorsal medial furrow is moderately expressed. The articulation is zygospondylous. Podial basins are moderate in size.
</paragraph>
</subSubSection>
<subSubSection id="7CEEBB89FFE2521EFF19F8F686F8F81A" pageId="84" pageNumber="85" type="materials_examined">
<paragraph id="344BE802FFE2521EFF19F8F686F8F81A" blockId="84.[151,1436,1071,2031]" pageId="84" pageNumber="85">
<emphasis id="06803410FFE2521EFF19F8F685AFF88A" bold="true" box="[198,416,1818,1844]" pageId="84" pageNumber="85">Material studied.</emphasis>
<taxonomicName id="F3F49381FFE2521EFE75F8F78756F88A" authority="(Lyman, 1875)" baseAuthorityName="Lyman" baseAuthorityYear="1875" box="[426,857,1818,1844]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="84" pageNumber="85" phylum="Echinodermata" rank="species" species="hirsuta">
<emphasis id="06803410FFE2521EFE75F8F78695F88A" box="[426,666,1819,1844]" italics="true" pageId="84" pageNumber="85">Ophiopristis hirsuta</emphasis>
(
<bibRefCitation id="506595F3FFE2521EFD72F8F6875FF88A" author="Lyman, T." box="[685,848,1818,1844]" pageId="84" pageNumber="85" pagination="1 - 34" refId="ref90050" refString="Lyman, T. (1875) Zoological results of the Hassler Expedition. II. Ophiuridae and Astrophytidae, including those dredged by the late Dr. William Stimpson. Ilustrated Catalogue Museum Comparative Zoology, Harvard College, 8, 1 - 34." type="journal article" year="1875">Lyman, 1875</bibRefCitation>
)
</taxonomicName>
,
<materialsCitation id="849CE25FFFE2521EFCB7F8F686E4F8C1" collectionCode="MNHN" pageId="84" pageNumber="85" specimenCode="MCZ 1950, USNM 6429" specimenCount="3" typeStatus="holotype">
<typeStatus id="EB4F56A0FFE2521EFCB7F8F687DAF88A" box="[872,981,1818,1844]" pageId="84" pageNumber="85" type="holotype">Holotype</typeStatus>
<specimenCode id="64524079FFE2521EFC00F8F6806CF88A" box="[991,1123,1818,1844]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="84" pageNumber="85" type="Museum">MCZ 1950</specimenCode>
(
<figureCitation id="ACCFF487FFE2521EFBA9F8F68120F88A" box="[1142,1327,1818,1844]" captionStart="FIGURE 57" captionStartId="83.[151,255,1564,1588]" captionTargetBox="[192,1395,194,1543]" captionTargetId="figure-210@83.[192,1395,194,1543]" captionTargetPageId="83" captionText="FIGURE 57. Genera Ophioprium and Ophiopristis, external views. A-B, Ophioprium cervicornis (Lyman, 1883), holotype MCZ 2043, 10 mm dd, dorsal and ventral views respectively; C-D, Ophiopristis hirsuta (Lyman, 1875), holotype MCZ 1950, 5.8 mm dd, dorsal and ventral views respectively; E. Ophioprium cervicornis (Lyman, 1883), holotype MCZ 2043, arm spine articulations, proximal segments; F. Ophiopristis hirsuta (Lyman, 1875), paratype MCZ 1951, 5 mm dd, ventral view. Photos: Alexander Martynov." pageId="84" pageNumber="85">Figs 57CD, F</figureCitation>
);
<specimenCode id="64524079FFE2521EFA98F8F684DAF8E4" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="84" pageNumber="85" type="Museum">USNM 6429</specimenCode>
,
<specimenCount id="22F2238BFFE2521EFF3EF8AC8568F8E4" box="[225,359,1856,1882]" pageId="84" pageNumber="85" type="generic">1 specimen</specimenCount>
(
<figureCitation id="ACCFF487FFE2521EFEA6F8AC85DDF8E4" box="[377,466,1856,1882]" captionStart="FIGURE 58" captionStartId="85.[151,255,1371,1395]" captionTargetBox="[158,1428,343,1349]" captionTargetId="figure-246@85.[158,1428,343,1349]" captionTargetPageId="85" captionText="FIGURE 58. Ophiopristis hirsuta (Lyman, 1875), USNM 6429, 10 mm dd, details. A. arm spine articulations, proximal segments; B. same, details; C. hollow spine transversally sectioned; D. spine; E. same, ventral view; F. proximal vertebra, dorsal view; G. same, proximal view; H. same, distal view. Scale bars: AB, FH, 300 µ; C, E, 100 µ; D, 1 mm. Photos: Alexander Martynov." pageId="84" pageNumber="85">Fig. 58</figureCitation>
);
<taxonomicName id="F3F49381FFE2521EFE36F8AC87AEF8E4" authority="(Koehler, 1904)" baseAuthorityName="Koehler" baseAuthorityYear="1904" box="[489,929,1856,1882]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="84" pageNumber="85" phylum="Echinodermata" rank="species" species="luctosa">
<emphasis id="06803410FFE2521EFE36F8AC86D6F8E7" box="[489,729,1856,1881]" italics="true" pageId="84" pageNumber="85">Ophiopristis luctosa</emphasis>
(Koehler, 1904)
</taxonomicName>
,
<collectionCode id="52E570C7FFE2521EFC71F8AC8007F8E4" box="[942,1032,1856,1882]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="84" pageNumber="85">MNHN</collectionCode>
EcOs 2297822979,
<specimenCount id="22F2238BFFE2521EFAD5F8AC8193F8E4" box="[1290,1436,1856,1882]" pageId="84" pageNumber="85" type="generic">2 specimens</specimenCount>
(
<figureCitation id="ACCFF487FFE2521EFF7FF88984F1F8C1" box="[160,254,1893,1919]" captionStart="FIGURE 59" captionStartId="86.[151,255,1557,1581]" captionTargetBox="[158,1428,647,1535]" captionTargetId="figure-290@86.[158,1428,647,1535]" captionTargetPageId="86" captionText="FIGURE 59. Ophiopristis luctosa (Koehler, 1904), MNHN EcOs 22978, 4.5 mm dd, external view and details. A. dorsal view; B. ventral view; C. lateral arm plate and spine articulations, proximal segments; DE, arm spine articulations, details; F. hollow spine transversally sectioned; G. spine, proximal segments; H. proximal vertebra, dorsal view; I. same, ventral view; J. same, proximal view; K. same, distal view; L. spine, ventral view. Scale bars: C-E, H-K, 100 µ; F, 30 µ; L, 300 µ. AB Tatiana Korshunova; CM Alexander Martynov." pageId="84" pageNumber="85">Fig. 59</figureCitation>
);
<taxonomicName id="F3F49381FFE2521EFEC9F88986E4F8C1" authority="(Koehler, 1904)" baseAuthorityName="Koehler" baseAuthorityYear="1904" box="[278,747,1893,1919]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="84" pageNumber="85" phylum="Echinodermata" rank="species" species="procera">
<emphasis id="06803410FFE2521EFEC9F8898615F8C0" box="[278,538,1893,1918]" italics="true" pageId="84" pageNumber="85">Ophiopristis procera</emphasis>
(Koehler, 1904)
</taxonomicName>
</materialsCitation>
,
<materialsCitation id="849CE25FFFE2521EFD26F88986FCF81A" collectionCode="MNHN" pageId="84" pageNumber="85" specimenCount="4" typeStatus="syntype">
<typeStatus id="EB4F56A0FFE2521EFD26F8898752F8C1" box="[761,861,1893,1919]" pageId="84" pageNumber="85" type="syntype">Syntype</typeStatus>
EcOs 20406 (
<figureCitation id="ACCFF487FFE2521EFBCEF8898060F8C1" box="[1041,1135,1893,1919]" captionStart="FIGURE 60" captionStartId="87.[151,255,1654,1678]" captionTargetBox="[158,1428,197,1633]" captionTargetId="figure-194@87.[158,1428,197,1633]" captionTargetPageId="87" captionText="FIGURE 60. Ophiopristis procera (Koehler, 1904), syntype MNHN EcOs 20406, 12 mm dd, external view and details. A. dorsal view; B. ventral view; C. proximal arm segments, ventral view showing oval tentacle scales; D. same, dorsal view, showing arm spine articulations and spines; EF, lateral arm plate and spine articulations, proximal segments; G. same, details; H. distal vertebra, dorsal view; I. same, ventral view; J. proximal vertebra, dorsal view; K. same, distal view; L. hollow proximal spine transversally sectioned; M. proximal spine, ventral view; N. proximal spine. Scale bars: EI, 100 µ; J, N, 1 mm; K, M 300 µ; L, 100 µ. AC Tatiana Korshunova; DN Alexander Martynov." pageId="84" pageNumber="85">Fig. 60</figureCitation>
);
<taxonomicName id="F3F49381FFE2521EFB58F889855CF81A" authority="(Koehler, 1905)" baseAuthorityName="Koehler" baseAuthorityYear="1905" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="84" pageNumber="85" phylum="Echinodermata" rank="species" species="dissidens">
<emphasis id="06803410FFE2521EFB58F8898193F8C0" box="[1159,1436,1893,1918]" italics="true" pageId="84" pageNumber="85">Ophiopristis dissidens</emphasis>
(
<bibRefCitation id="506595F3FFE2521EFF40F8668545F81A" author="Koehler, R." box="[159,330,1930,1956]" pageId="84" pageNumber="85" pagination="1 - 142" refId="ref88606" refString="Koehler, R. (1905) Ophiures littorales. Siboga-Expeditie Monograph, 45, 1 - 142," type="journal article" year="1905">Koehler, 1905</bibRefCitation>
)
</taxonomicName>
,
<collectionCode id="52E570C7FFE2521EFE80F86685B6F81A" box="[351,441,1930,1956]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="84" pageNumber="85">MNHN</collectionCode>
EcOs 22478,
<specimenCount id="22F2238BFFE2521EFD80F86686FCF81A" box="[607,755,1930,1956]" pageId="84" pageNumber="85" type="generic">4 specimens</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7CEEBB89FFE2521FFF19F84385DCF8A5" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="discussion">
<paragraph id="344BE802FFE2521FFF19F84384EBFE9C" blockId="84.[151,1436,1071,2031]" lastBlockId="85.[151,1436,152,290]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<emphasis id="06803410FFE2521EFF19F8438530F877" bold="true" box="[198,319,1967,1993]" pageId="84" pageNumber="85">Remarks.</emphasis>
<bibRefCitation id="506595F3FFE2521EFE99F85C8658F874" author="O'Hara, T. D. &amp; Stohr, S." box="[326,599,1968,1994]" pageId="84" pageNumber="85" pagination="33 - 141" refId="ref91113" refString="O'Hara, T. D. &amp; Stohr, S. (2006) Deep water Ophiuroidea (Echinodermata) of New Caledonia: Ophiacanthidae and Hemieuryalidae. In Richer De Forges, B. &amp; Justine, J. - L. (Eds), Tropical Deep-Sea Benthos. Volume 24. Memoires du Museum national d'Histoire Naturelle 193, 33 - 141." type="journal article" year="2006">OHara &amp; Stöhr (2006)</bibRefCitation>
recently reviewed the genus
<taxonomicName id="F3F49381FFE2521EFC74F85C8032F877" authorityName="Verrill" authorityYear="1899" box="[939,1085,1968,1993]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="84" pageNumber="85" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE2521EFC74F85C8032F877" box="[939,1085,1968,1993]" italics="true" pageId="84" pageNumber="85">Ophiopristis</emphasis>
</taxonomicName>
and further delineated it from other ophiacanthid genera, initially suggested by
<bibRefCitation id="506595F3FFE2521EFD0CF839879FF851" author="Paterson, G. L. J." box="[723,912,2005,2031]" pageId="84" pageNumber="85" pagination="1 - 162" refId="ref91221" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History), Zoology Series 49, 1 - 162." type="book chapter" year="1985">Paterson (1985)</bibRefCitation>
. Among the species included into this genus, a small Australian species
<taxonomicName id="F3F49381FFE3521FFE02FF748677FF0F" baseAuthorityName="H. L. Clark" baseAuthorityYear="1909" box="[477,632,152,177]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="species" species="axiologus">
<emphasis id="06803410FFE3521FFE02FF748677FF0F" box="[477,632,152,177]" italics="true" pageId="85" pageNumber="86">O. axiologus</emphasis>
</taxonomicName>
differs considerably from all previously known species by bottleshaped arm spines, disk covered with short stellate stumps instead of relatively long spinelets and possibly represents another genus (
<bibRefCitation id="506595F3FFE3521FFE14FF0E86DCFF42" author="O'Hara, T. D. &amp; Stohr, S." box="[459,723,226,252]" pageId="85" pageNumber="86" pagination="33 - 141" refId="ref91113" refString="O'Hara, T. D. &amp; Stohr, S. (2006) Deep water Ophiuroidea (Echinodermata) of New Caledonia: Ophiacanthidae and Hemieuryalidae. In Richer De Forges, B. &amp; Justine, J. - L. (Eds), Tropical Deep-Sea Benthos. Volume 24. Memoires du Museum national d'Histoire Naturelle 193, 33 - 141." type="journal article" year="2006">OHara &amp; Stöhr 2006</bibRefCitation>
).
<taxonomicName id="F3F49381FFE3521FFD38FF0E87FAFF45" baseAuthorityName="H. L. Clark" baseAuthorityYear="1909" box="[743,1013,226,251]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="species" species="axiologus">
<emphasis id="06803410FFE3521FFD38FF0E87FAFF45" box="[743,1013,226,251]" italics="true" pageId="85" pageNumber="86">Ophiopristis axiologus</emphasis>
</taxonomicName>
is therefore not included in the key below.
</paragraph>
<caption id="608BB88AFFE3521FFF48FAB785DDFA6C" pageId="85" pageNumber="86" startId="85.[151,255,1371,1395]" targetBox="[158,1428,343,1349]" targetPageId="85">
<paragraph id="344BE802FFE3521FFF48FAB785DDFA6C" blockId="85.[151,1437,1371,1490]" pageId="85" pageNumber="86">
<emphasis id="06803410FFE3521FFF48FAB78529FACD" bold="true" box="[151,294,1371,1395]" pageId="85" pageNumber="86">FIGURE 58.</emphasis>
<taxonomicName id="F3F49381FFE3521FFEF2FAB786A3FACC" authority="(Lyman, 1875)" baseAuthorityName="Lyman" baseAuthorityYear="1875" box="[301,684,1371,1394]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="species" species="hirsuta">
<emphasis id="06803410FFE3521FFEF2FAB7860BFACC" box="[301,516,1371,1394]" italics="true" pageId="85" pageNumber="86">Ophiopristis hirsuta</emphasis>
(
<bibRefCitation id="506595F3FFE3521FFDCCFAB786ABFACC" author="Lyman, T." box="[531,676,1371,1394]" pageId="85" pageNumber="86" pagination="1 - 34" refId="ref90050" refString="Lyman, T. (1875) Zoological results of the Hassler Expedition. II. Ophiuridae and Astrophytidae, including those dredged by the late Dr. William Stimpson. Ilustrated Catalogue Museum Comparative Zoology, Harvard College, 8, 1 - 34." type="journal article" year="1875">Lyman, 1875</bibRefCitation>
)
</taxonomicName>
, USNM 6429, 10 mm dd, details. A. arm spine articulations, proximal segments; B. same, details; C. hollow spine transversally sectioned; D. spine; E. same, ventral view; F. proximal vertebra, dorsal view; G. same, proximal view; H. same, distal view. Scale bars: AB, FH, 300 µ; C, E, 100 µ; D, 1 mm. Photos: Alexander Martynov.
</paragraph>
</caption>
<paragraph id="344BE802FFE3521FFF18FA17861BF94B" blockId="85.[151,1436,1531,1819]" pageId="85" pageNumber="86">
There are a number of species of the genus
<taxonomicName id="F3F49381FFE3521FFCD2FA1787BAF9AA" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[781,949,1531,1556]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE3521FFCD2FA1787BAF9AA" box="[781,949,1531,1556]" italics="true" pageId="85" pageNumber="86">Ophiacantha</emphasis>
</taxonomicName>
, which possess characters similar to
<taxonomicName id="F3F49381FFE3521FFF48F9CD8539F984" authorityName="Verrill" authorityYear="1899" box="[151,310,1569,1594]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE3521FFF48F9CD8539F984" box="[151,310,1569,1594]" italics="true" pageId="85" pageNumber="86">Ophiopristis</emphasis>
</taxonomicName>
, including long numerous spinelets and the pattern of the oral papillae, for instance
<taxonomicName id="F3F49381FFE3521FFAA1F9CD8608F9DE" authority="H. L. Clark, 1911" authorityName="H. L. Clark" authorityYear="1911" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="species" species="hylacantha">
<emphasis id="06803410FFE3521FFAA1F9CD852BF9E1" italics="true" pageId="85" pageNumber="86">O. hylacantha</emphasis>
H.L.
<bibRefCitation id="506595F3FFE3521FFEADF9AA8608F9DE" author="Clark, H. L." box="[370,519,1606,1632]" pageId="85" pageNumber="86" pagination="1 - 302" refId="ref85928" refString="Clark, H. L. (1911) North Pacific Ophiurans in the collection of the United States National Museum. Bulletin of the United States National Museum 75, 1 - 302." type="journal article" year="1911">Clark, 1911</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="F3F49381FFE3521FFDE9F9AA87D4F9DE" authority="H. L. Clark, 1911" authorityName="H. L. Clark" authorityYear="1911" box="[566,987,1606,1632]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="species" species="atopostoma">
<emphasis id="06803410FFE3521FFDE9F9AA86FEF9E1" box="[566,753,1606,1631]" italics="true" pageId="85" pageNumber="86">O. atopostoma</emphasis>
H.L.
<bibRefCitation id="506595F3FFE3521FFC9EF9AA87D4F9DE" author="Clark, H. L." box="[833,987,1606,1632]" pageId="85" pageNumber="86" pagination="1 - 302" refId="ref85928" refString="Clark, H. L. (1911) North Pacific Ophiurans in the collection of the United States National Museum. Bulletin of the United States National Museum 75, 1 - 302." type="journal article" year="1911">Clark, 1911</bibRefCitation>
</taxonomicName>
. H.L.
<bibRefCitation id="506595F3FFE3521FFBF2F9AA80D9F9DE" author="Clark, H. L." box="[1069,1238,1606,1632]" pageId="85" pageNumber="86" pagination="165 - 376" refId="ref85989" refString="Clark, H. L. (1915) Catalogue of recent Ophiurans based on collections of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology, Harvard 25, 165 - 376." type="journal article" year="1915">Clark (1915)</bibRefCitation>
, while creating
<taxonomicName id="F3F49381FFE3521FFF48F9878526F93A" authorityName="Verrill" authorityYear="1899" box="[151,297,1643,1668]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE3521FFF48F9878526F93A" box="[151,297,1643,1668]" italics="true" pageId="85" pageNumber="86">Ophioprium</emphasis>
</taxonomicName>
and delineating it from Verrills
<taxonomicName id="F3F49381FFE3521FFD65F987875FF93A" authorityName="Verrill" authorityYear="1899" box="[698,848,1643,1668]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE3521FFD65F987875FF93A" box="[698,848,1643,1668]" italics="true" pageId="85" pageNumber="86">Ophiopristis</emphasis>
</taxonomicName>
, noted that the former differs considerably from other ophiacanthids, whereas
<taxonomicName id="F3F49381FFE3521FFE2AF97D8687F914" authorityName="Verrill" authorityYear="1899" box="[501,648,1681,1706]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE3521FFE2AF97D8687F914" box="[501,648,1681,1706]" italics="true" pageId="85" pageNumber="86">Ophiopristis</emphasis>
</taxonomicName>
should be considered as a synonym of
<taxonomicName id="F3F49381FFE3521FFB86F97D80FDF914" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[1113,1266,1681,1706]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="genus">
<emphasis id="06803410FFE3521FFB86F97D80FDF914" box="[1113,1266,1681,1706]" italics="true" pageId="85" pageNumber="86">Ophiacantha</emphasis>
</taxonomicName>
. The situation needs further study, involving delineation of the more than 130 currently known
<taxonomicName id="F3F49381FFE3521FFBBEF95A8155F96E" box="[1121,1370,1718,1744]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="85" pageNumber="86" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="06803410FFE3521FFBBEF95A80F3F971" box="[1121,1276,1718,1743]" italics="true" pageId="85" pageNumber="86">Ophiacantha</emphasis>
species
</taxonomicName>
from some other ophiacanthid genera.
</paragraph>
<paragraph id="344BE802FFE3521FFF19F8EC85DCF8A5" blockId="85.[151,1436,1531,1819]" box="[198,467,1792,1819]" pageId="85" pageNumber="86">
<emphasis id="06803410FFE3521FFF19F8EC85BAF8A4" bold="true" box="[198,437,1792,1818]" pageId="85" pageNumber="86">Number of species:</emphasis>
8.
</paragraph>
</subSubSection>
</treatment>
</document>