222 lines
33 KiB
XML
222 lines
33 KiB
XML
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<mods:title id="4089FD4569AB1BFB949C6DAEA01CB54A">Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae 2697</mods:title>
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<mods:namePart id="2EEB6F67929008924B739230570A4F13">Martynov, Alexander</mods:namePart>
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<mods:title id="7B682E0CDA6DEBFCE61C45C6A00F5516">Zootaxa</mods:title>
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<mods:date id="8F50A78B5A7EE49E4F107F6530350D58">2010</mods:date>
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<mods:number id="CD435AEF88597FF4F4634DE95AFA3CE5">2010-12-03</mods:number>
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<mods:number id="560CD7EA59595A9B223893DD1A649A82">2697</mods:number>
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<subSubSection id="7CEEBB89FFE0521CFF48F8E68653F89A" box="[151,604,1802,1829]" pageId="86" pageNumber="87" type="nomenclature">
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<paragraph id="344BE802FFE0521CFF48F8E68653F89A" blockId="86.[151,604,1802,1863]" box="[151,604,1802,1829]" pageId="86" pageNumber="87">
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<heading id="6F035F6EFFE0521CFF48F8E68653F89A" bold="true" box="[151,604,1802,1829]" fontSize="11" level="1" pageId="86" pageNumber="87" reason="1">
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<emphasis id="06803410FFE0521CFF48F8E68653F89A" bold="true" box="[151,604,1802,1829]" pageId="86" pageNumber="87">
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Genus
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<taxonomicName id="F3F49381FFE0521CFF31F8E78653F89A" ID-CoL="7PQZD" authority="H. L. Clark, 1915" authorityName="H. L. Clark" authorityYear="1915" box="[238,604,1802,1829]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="86" pageNumber="87" phylum="Echinodermata" rank="genus">
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<emphasis id="06803410FFE0521CFF31F8E7858DF89B" bold="true" box="[238,386,1803,1829]" italics="true" pageId="86" pageNumber="87">Ophioprium</emphasis>
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H.L.
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<bibRefCitation id="506595F3FFE0521CFE15F8E68653F89A" author="Clark, H. L." box="[458,604,1802,1828]" pageId="86" pageNumber="87" pagination="165 - 376" refId="ref85989" refString="Clark, H. L. (1915) Catalogue of recent Ophiurans based on collections of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology, Harvard 25, 165 - 376." type="journal article" year="1915">Clark, 1915</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7CEEBB89FFE0521CFF48F8C185ACF8F9" box="[151,419,1837,1863]" pageId="86" pageNumber="87" type="description">
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<paragraph id="344BE802FFE0521CFF48F8C185ACF8F9" blockId="86.[151,604,1802,1863]" box="[151,419,1837,1863]" pageId="86" pageNumber="87">
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<figureCitation id="ACCFF487FFE0521CFF48F8C18542F8F9" box="[151,333,1837,1863]" captionStart="FIGURE 57" captionStartId="83.[151,255,1564,1588]" captionTargetBox="[192,1395,194,1543]" captionTargetId="figure-210@83.[192,1395,194,1543]" captionTargetPageId="83" captionText="FIGURE 57. Genera Ophioprium and Ophiopristis, external views. A-B, Ophioprium cervicornis (Lyman, 1883), holotype MCZ 2043, 10 mm dd, dorsal and ventral views respectively; C-D, Ophiopristis hirsuta (Lyman, 1875), holotype MCZ 1950, 5.8 mm dd, dorsal and ventral views respectively; E. Ophioprium cervicornis (Lyman, 1883), holotype MCZ 2043, arm spine articulations, proximal segments; F. Ophiopristis hirsuta (Lyman, 1875), paratype MCZ 1951, 5 mm dd, ventral view. Photos: Alexander Martynov." pageId="86" pageNumber="87">Figures 57A, B</figureCitation>
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;
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<figureCitation id="ACCFF487FFE0521CFE85F8C185ACF8F9" box="[346,419,1837,1863]" captionStart-0="FIGURE 61" captionStart-1="FIGURE 62" captionStartId-0="88.[151,255,1911,1935]" captionStartId-1="89.[151,255,1163,1187]" captionTargetBox-0="[158,1428,679,1890]" captionTargetBox-1="[173,1410,182,1157]" captionTargetId-0="figure-295@88.[158,1428,679,1890]" captionTargetId-1="figure-329@89.[173,1410,182,1159]" captionTargetPageId-0="88" captionTargetPageId-1="89" captionText-0="FIGURE 61. Ophioprium permixtum (Koehler, 1914), USNM E 05300, 15 mm dd, external views. A. dorsal view; B. ventral view; C. dorsal disk, details; D. dorsal disk, details showing small radial shields; E. ventral view, details oral and adoral shield papillae; F. interradial view showing long wide genital slits; G. ventral view, showing details of the teeth. Photos: Tatiana Korshunova." captionText-1="FIGURE 62. Ophioprium permixtum (Koehler, 1914), USNM E 05300, 15 mm dd, details. A-B, lateral plate and spine articulations, proximal segments; C, G, proximal spine, section showing small cavities; D. proximal vertebra, dorsal view; E. same, ventral view; F. same, distal view; H. proximal spine; I. proximal spine, ventral view. Scale bars: A–B, D– F, 300 µ; C, G, K, 100 µ; I, 1 mm. Photos: Alexander Martynov." pageId="86" pageNumber="87">61–62</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="7CEEBB89FFE0521CFF48F894864FF831" box="[151,576,1912,1935]" pageId="86" pageNumber="87" type="reference_group">
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<paragraph id="344BE802FFE0521CFF48F894864FF831" blockId="86.[151,974,1912,1967]" box="[151,576,1912,1935]" pageId="86" pageNumber="87">
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<treatmentCitationGroup id="14E4CF2CFFE0521CFF48F894864FF831" box="[151,576,1912,1935]" pageId="86" pageNumber="87">
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<treatmentCitation id="B555CE13FFE0521CFF48F894864FF831" author="Clark, H. L." box="[151,576,1912,1935]" page="215" pageId="86" pageNumber="87" year="1915">
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<taxonomicName id="F3F49381FFE0521CFF48F894864FF831" ID-CoL="7PQZD" authority="H. L. Clark, 1915: 214 - 215" authorityName="H. L. Clark" authorityPageNumber="214 - 215" authorityYear="1915" box="[151,576,1912,1935]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="86" pageNumber="87" phylum="Echinodermata" rank="genus">
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<emphasis id="06803410FFE0521CFF48F8948516F831" box="[151,281,1912,1935]" italics="true" pageId="86" pageNumber="87">Ophioprium</emphasis>
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H.L.
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<bibRefCitation id="506595F3FFE0521CFE88F894864FF831" author="Clark, H. L." box="[343,576,1912,1935]" pageId="86" pageNumber="87" pagination="165 - 376" refId="ref85989" refString="Clark, H. L. (1915) Catalogue of recent Ophiurans based on collections of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology, Harvard 25, 165 - 376." type="journal article" year="1915">Clark, 1915: 214–215</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="7CEEBB89FFE05212FF48F87486A8FDCC" lastPageId="88" lastPageNumber="89" pageId="86" pageNumber="87" type="materials_examined">
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<paragraph id="344BE802FFE05212FF48F87486A8FDCC" blockId="86.[151,974,1912,1967]" lastBlockId="88.[151,1436,152,626]" lastPageId="88" lastPageNumber="89" pageId="86" pageNumber="87">
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<typeStatus id="EB4F56A0FFE0521CFF48F87484C3F811" box="[151,204,1944,1967]" pageId="86" pageNumber="87">Type</typeStatus>
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species:
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<taxonomicName id="F3F49381FFE0521CFEF2F87486C2F811" authority="Lyman, 1883" authorityName="Lyman" authorityYear="1883" box="[301,717,1944,1967]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="86" pageNumber="87" phylum="Echinodermata" rank="species" species="cervicornis">
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<emphasis id="06803410FFE0521CFEF2F8748638F811" box="[301,567,1944,1967]" italics="true" pageId="86" pageNumber="87">Ophiacantha cervicornis</emphasis>
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<bibRefCitation id="506595F3FFE0521CFDE1F87486C2F811" author="Lyman, T." box="[574,717,1944,1967]" pageId="86" pageNumber="87" pagination="227 - 287" refId="ref90277" refString="Lyman, T. (1883) Report on the Ophiuroidea. Bulletin of the Museum of Comparative Zoology at Harvard College 10, 227 - 287." type="journal article" year="1883">Lyman, 1883</bibRefCitation>
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</taxonomicName>
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, by original designation
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<emphasis id="06803410FFE1521DFF48F8BA8518F8CE" bold="true" box="[151,279,1878,1904]" pageId="87" pageNumber="88">Diagnosis.</emphasis>
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The disk is covered with numerous small, uniform scales and a thin skin layer. The radial shields are small, rounded, hardly conspicuous in dried specimens. The adradial and abradial genital plates are well defined. Details of the articulations of genital plates and radial shields are unknown. The genital slits are long and conspicuous, genital plates border more than half of the slits. The jaw bears numerous spiniform oral papillae, similar in shape to the small cluster of ventralmost teeth. Two adoral shield papillae and one adjacent oral papilla of each jaw are narrow, spiniform, 3–4 times longer than the oral papillae, placed around the second tentacle pore. The half-jaws, judged from the length of the dental plate, are high. The dental plate is small and elongated, without folds and with few rounded sockets, alternately placed. The teeth are spiniform and few in number (besides the apical papillae there are 4–5 teeth). All teeth are similar in shape. The oral shield is lozenge-shaped with a distinct distal lobe. The adoral shields have distal bilobed wings, proximally tapered. Dorsal and ventral arm plates are well developed. The arm spine articulations are placed almost transversally in relation to the lateral plate. The muscle opening is larger than the nerve opening. A prominent distal perforated lobe occupies the distal part of the articulation. The sigmoidal fold is absent. The arm spines are long, flattened, solid over most of their length but contain some holes of various sizes, not hooked distally. The tentacles scales are thin, spiniform, placed on both lateral and ventral arm plates. The tentacles pores are large. The vertebrae have a rounded distal keel, with deep dorsal medial furrow. The articulation is zygospondylous. The podial basins are large.
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</paragraph>
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</subSubSection>
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||
<caption id="608BB88AFFE1521DFF48F99A80C3F893" pageId="87" pageNumber="88" startId="87.[151,255,1654,1678]" targetBox="[158,1428,197,1633]" targetPageId="87">
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||
<paragraph id="344BE802FFE1521DFF48F99A80C3F893" blockId="87.[151,1437,1654,1838]" pageId="87" pageNumber="88">
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||
<emphasis id="06803410FFE1521DFF48F99A8528F930" bold="true" box="[151,295,1654,1678]" pageId="87" pageNumber="88">FIGURE 60.</emphasis>
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<taxonomicName id="F3F49381FFE1521DFEF1F99B86CEF933" authority="(Koehler, 1904)" baseAuthorityName="Koehler" baseAuthorityYear="1904" box="[302,705,1654,1678]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="87" pageNumber="88" phylum="Echinodermata" rank="species" species="procera">
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<emphasis id="06803410FFE1521DFEF1F99B8601F930" box="[302,526,1655,1678]" italics="true" pageId="87" pageNumber="88">Ophiopristis procera</emphasis>
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(Koehler, 1904)
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</taxonomicName>
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, syntype MNHN EcOs 20406, 12 mm dd, external view and details. A. dorsal view; B. ventral view; C. proximal arm segments, ventral view showing oval tentacle scales; D. same, dorsal view, showing arm spine articulations and spines; E–F, lateral arm plate and spine articulations, proximal segments; G. same, details; H. distal vertebra, dorsal view; I. same, ventral view; J. proximal vertebra, dorsal view; K. same, distal view; L. hollow proximal spine transversally sectioned; M. proximal spine, ventral view; N. proximal spine. Scale bars: E–I, 100 µ; J, N, 1 mm; K, M 300 µ; L, 100 µ. A–C Tatiana Korshunova; D–N Alexander Martynov.
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</paragraph>
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</caption>
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<caption id="608BB88AFFEE5212FF48F89B85C3F851" pageId="88" pageNumber="89" startId="88.[151,255,1911,1935]" targetBox="[158,1428,679,1890]" targetPageId="88">
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<paragraph id="344BE802FFEE5212FF48F89B85C3F851" blockId="88.[151,1437,1911,2031]" pageId="88" pageNumber="89">
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<emphasis id="06803410FFEE5212FF48F89B8527F831" bold="true" box="[151,296,1911,1935]" pageId="88" pageNumber="89">FIGURE 61.</emphasis>
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<taxonomicName id="F3F49381FFEE5212FEEFF89486ECF831" authority="(Koehler, 1914)" baseAuthorityName="Koehler" baseAuthorityYear="1914" box="[304,739,1912,1935]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="88" pageNumber="89" phylum="Echinodermata" rank="species" species="permixtum">
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<emphasis id="06803410FFEE5212FEEFF8948621F831" box="[304,558,1912,1935]" italics="true" pageId="88" pageNumber="89">Ophioprium permixtum</emphasis>
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(
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<bibRefCitation id="506595F3FFEE5212FDE3F89486D4F831" author="Koehler, R." box="[572,731,1912,1935]" pageId="88" pageNumber="89" pagination="1 - 173" refId="ref88727" refString="Koehler, R. (1914) A contribution to the study of Ophiurans of the United States National Museum. Bulletin of the United States National Museum 84, 1 - 173." type="journal article" year="1914">Koehler, 1914</bibRefCitation>
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)
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</taxonomicName>
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, USNM E 05300, 15 mm dd, external views. A. dorsal view; B. ventral view; C. dorsal disk, details; D. dorsal disk, details showing small radial shields; E. ventral view, details oral and adoral shield papillae; F. interradial view showing long wide genital slits; G. ventral view, showing details of the teeth. Photos: Tatiana Korshunova.
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</paragraph>
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</caption>
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<caption id="608BB88AFFEF5213FF48FB67874DFABC" pageId="89" pageNumber="90" startId="89.[151,255,1163,1187]" targetBox="[173,1410,182,1157]" targetPageId="89">
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<paragraph id="344BE802FFEF5213FF48FB67874DFABC" blockId="89.[151,1436,1163,1283]" pageId="89" pageNumber="90">
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<emphasis id="06803410FFEF5213FF48FB678528FB1D" bold="true" box="[151,295,1163,1187]" pageId="89" pageNumber="90">FIGURE 62.</emphasis>
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<taxonomicName id="F3F49381FFEF5213FEF1FB6786D1FB1C" authority="(Koehler, 1914)" baseAuthorityName="Koehler" baseAuthorityYear="1914" box="[302,734,1163,1186]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="permixtum">
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<emphasis id="06803410FFEF5213FEF1FB678625FB1C" box="[302,554,1163,1186]" italics="true" pageId="89" pageNumber="90">Ophioprium permixtum</emphasis>
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||
(
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<bibRefCitation id="506595F3FFEF5213FDE6FB6786D9FB1C" author="Koehler, R." box="[569,726,1163,1186]" pageId="89" pageNumber="90" pagination="1 - 173" refId="ref88727" refString="Koehler, R. (1914) A contribution to the study of Ophiurans of the United States National Museum. Bulletin of the United States National Museum 84, 1 - 173." type="journal article" year="1914">Koehler, 1914</bibRefCitation>
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)
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</taxonomicName>
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||
, USNM E 05300, 15 mm dd, details. A-B, lateral plate and spine articulations, proximal segments; C, G, proximal spine, section showing small cavities; D. proximal vertebra, dorsal view; E. same, ventral view; F. same, distal view; H. proximal spine; I. proximal spine, ventral view. Scale bars: A–B, D– F, 300 µ; C, G, K, 100 µ; I, 1 mm. Photos: Alexander Martynov.
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</paragraph>
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</caption>
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<subSubSection id="7CEEBB89FFEF5213FF19FADC809BF995" pageId="89" pageNumber="90" type="discussion">
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<paragraph id="344BE802FFEF5213FF19FADC809BF995" blockId="89.[151,1436,1328,1691]" pageId="89" pageNumber="90">
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<emphasis id="06803410FFEF5213FF19FADC8530FAF4" bold="true" box="[198,319,1328,1354]" pageId="89" pageNumber="90">Remarks.</emphasis>
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Several different species have been included in the genus
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<taxonomicName id="F3F49381FFEF5213FC37FADD8078FAF4" authorityName="H. L. Clark" authorityYear="1915" box="[1000,1143,1329,1354]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="genus">
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<emphasis id="06803410FFEF5213FC37FADD8078FAF4" box="[1000,1143,1329,1354]" italics="true" pageId="89" pageNumber="90">Ophioprium</emphasis>
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</taxonomicName>
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(H.L.
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<bibRefCitation id="506595F3FFEF5213FB1BFADD8145FAF5" author="Clark, H. L." box="[1220,1354,1329,1355]" pageId="89" pageNumber="90" pagination="165 - 376" refId="ref85989" refString="Clark, H. L. (1915) Catalogue of recent Ophiurans based on collections of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology, Harvard 25, 165 - 376." type="journal article" year="1915">Clark 1915</bibRefCitation>
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;
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<bibRefCitation id="506595F3FFEF5213FA89FADD84DAFACE" author="Baker, A. N." pageId="89" pageNumber="90" pagination="21 - 51" refId="ref84962" refString="Baker, A. N. (1979) Some Ophiuroidea from the Tasman Sea and adjacent waters. New Zealand Journal of Zoology 6, 21 - 51." type="journal article" year="1979">Baker 1979</bibRefCitation>
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||
;
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||
<bibRefCitation id="506595F3FFEF5213FF3DFABA85F1FACE" author="Baker, A. N. & Devaney, D. M." box="[226,510,1366,1392]" pageId="89" pageNumber="90" pagination="155 - 178" refId="ref84992" refString="Baker, A. N. & Devaney, D. M. (1981) New records of Ophiuroidea (Echinodermata) from Southern Australia, including new species of Ophiacantha and Ophionereis. Transactions of the Royal Society of Southern Australia, 155 - 178." type="book chapter" year="1981">Baker & Devaney 1981</bibRefCitation>
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).
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<bibRefCitation id="506595F3FFEF5213FDCCFABA86C0FACE" author="Paterson, G. L. J." box="[531,719,1366,1392]" pageId="89" pageNumber="90" pagination="1 - 162" refId="ref91221" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History), Zoology Series 49, 1 - 162." type="book chapter" year="1985">Paterson (1985)</bibRefCitation>
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excluded from this genus
|
||
<taxonomicName id="F3F49381FFEF5213FBD6FABA8198FACE" authority="Baker, 1979" authorityName="Baker" authorityYear="1979" box="[1033,1431,1366,1392]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="larissae">
|
||
<emphasis id="06803410FFEF5213FBD6FABA80F2FAD1" box="[1033,1277,1366,1391]" italics="true" pageId="89" pageNumber="90">Ophioprium larissae</emphasis>
|
||
<bibRefCitation id="506595F3FFEF5213FADAFABA8198FACE" author="Baker, A. N." box="[1285,1431,1366,1392]" pageId="89" pageNumber="90" pagination="21 - 51" refId="ref84962" refString="Baker, A. N. (1979) Some Ophiuroidea from the Tasman Sea and adjacent waters. New Zealand Journal of Zoology 6, 21 - 51." type="journal article" year="1979">Baker, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3F49381FFEF5213FF48FA9785CBFA2B" authority="Baker, 1979" authorityName="Baker" authorityYear="1979" box="[151,452,1403,1429]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="kapalae">
|
||
<emphasis id="06803410FFEF5213FF48FA97852FFA2A" box="[151,288,1403,1428]" italics="true" pageId="89" pageNumber="90">O. kapalae</emphasis>
|
||
<bibRefCitation id="506595F3FFEF5213FEF5FA9785CBFA2B" author="Baker, A. N." box="[298,452,1403,1429]" pageId="89" pageNumber="90" pagination="21 - 51" refId="ref84962" refString="Baker, A. N. (1979) Some Ophiuroidea from the Tasman Sea and adjacent waters. New Zealand Journal of Zoology 6, 21 - 51." type="journal article" year="1979">Baker, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3F49381FFEF5213FE0CFA978767FA2B" authority="(H. L. Clark, 1909)" baseAuthorityName="H. L. Clark" baseAuthorityYear="1909" box="[467,872,1403,1429]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiopristis" kingdom="Animalia" order="Ophiacanthida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="axiologus">
|
||
<emphasis id="06803410FFEF5213FE0CFA97867CFA2A" box="[467,627,1403,1428]" italics="true" pageId="89" pageNumber="90">O. axiologus</emphasis>
|
||
(H.L.
|
||
<bibRefCitation id="506595F3FFEF5213FD15FA978750FA2B" author="Clark, H. L." box="[714,863,1403,1429]" pageId="89" pageNumber="90" pagination="519 - 564" refId="ref85888" refString="Clark, H. L. (1909) Scientific results of the trawling expedition of H. M. C. S. Thetis. Echinodermata. Memoirs of the Australian Museum, 4, 519 - 564." type="journal article" year="1909">Clark, 1909</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3F49381FFEF5213FC77FA9780D9FA2B" authority="(Lyman, 1878)" baseAuthorityName="Lyman" baseAuthorityYear="1878" box="[936,1238,1403,1429]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="rosea">
|
||
<emphasis id="06803410FFEF5213FC77FA97801DFA2A" box="[936,1042,1403,1428]" italics="true" pageId="89" pageNumber="90">O. rosea</emphasis>
|
||
(
|
||
<bibRefCitation id="506595F3FFEF5213FBFAFA9780C2FA2B" author="Lyman, T." box="[1061,1229,1403,1429]" pageId="89" pageNumber="90" pagination="65 - 168" refId="ref90096" refString="Lyman, T. (1878) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard 5, 65 - 168." type="journal article" year="1878">Lyman, 1878</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but retained
|
||
<taxonomicName id="F3F49381FFEF5213FAA1FA9785F2FA05" authority="H. L. Clark, 1915" authorityName="H. L. Clark" authorityYear="1915" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="imperfectum">
|
||
<emphasis id="06803410FFEF5213FAA1FA978526FA04" italics="true" pageId="89" pageNumber="90">O. imperfectum</emphasis>
|
||
H.L.
|
||
<bibRefCitation id="506595F3FFEF5213FEB0FA4D85F2FA05" author="Clark, H. L." box="[367,509,1441,1467]" pageId="89" pageNumber="90" pagination="165 - 376" refId="ref85989" refString="Clark, H. L. (1915) Catalogue of recent Ophiurans based on collections of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology, Harvard 25, 165 - 376." type="journal article" year="1915">Clark, 1915</bibRefCitation>
|
||
</taxonomicName>
|
||
. Re-examination of the
|
||
<typeStatus id="EB4F56A0FFEF5213FCC7FA4D8771FA05" box="[792,894,1441,1467]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="F3F49381FFEF5213FC79FA4D8053FA04" authorityName="H. L. Clark" authorityYear="1915" box="[934,1116,1441,1466]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="imperfectum">
|
||
<emphasis id="06803410FFEF5213FC79FA4D8053FA04" box="[934,1116,1441,1466]" italics="true" pageId="89" pageNumber="90">O. imperfectum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="ACCFF487FFEF5213FBB2FA4D811CFA05" box="[1133,1299,1441,1467]" captionStart="FIGURE 52" captionStartId="76.[151,255,1590,1614]" captionTargetBox="[158,1428,194,1569]" captionTargetId="figure-217@76.[158,1428,194,1569]" captionTargetPageId="76" captionText="FIGURE 52. Ophiophrura tripapillata (Stöhr & Segonzac, 2005) comb.nov. (A–B, D–E, holotype MNHN EcOs 22795, 10 mm dd), Ophiophrura imperfecta (H.L. Clark, 1915) comb.nov. (F, H–J, holotype MSZ 2167, ca. 7 mm dd), and Ophiophrura liodisca H.L. Clark, 1911, holotype USNM 25672, 8 mm dd (C, G). A, H, dorsal view; B, G, I, ventral views; C, F, dorsal disk details; D-E, G, J, arm segments, ventral views, details showing pattern of several tentacle scales. A–B, D–E Sabine Stöhr; C, G Tatiana Korshunova; F, H, I–J Alexander Martynov." pageId="89" pageNumber="90">Figs 52F, H–J</figureCitation>
|
||
) during the present study reveals considerable differences from the type species of the genus
|
||
<taxonomicName id="F3F49381FFEF5213FB91FA2A80D2FA61" authorityName="H. L. Clark" authorityYear="1915" box="[1102,1245,1478,1503]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFEF5213FB91FA2A80D2FA61" box="[1102,1245,1478,1503]" italics="true" pageId="89" pageNumber="90">Ophioprium</emphasis>
|
||
</taxonomicName>
|
||
. Thus, only two species remain within the genus. Both species are very similar and their taxonomy needs further study. Verrill’s authorship of the genus
|
||
<taxonomicName id="F3F49381FFEF5213FDC9F9FD86AAF994" authorityName="H. L. Clark" authorityYear="1915" box="[534,677,1553,1578]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="06803410FFEF5213FDC9F9FD86AAF994" box="[534,677,1553,1578]" italics="true" pageId="89" pageNumber="90">Ophioprium</emphasis>
|
||
</taxonomicName>
|
||
was incorrectly listed by
|
||
<bibRefCitation id="506595F3FFEF5213FC0CF9FD809FF995" author="Paterson, G. L. J." box="[979,1168,1553,1579]" pageId="89" pageNumber="90" pagination="1 - 162" refId="ref91221" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History), Zoology Series 49, 1 - 162." type="book chapter" year="1985">Paterson (1985)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7CEEBB89FFEF5213FF19F9DA84CAF925" pageId="89" pageNumber="90" type="materials_examined">
|
||
<paragraph id="344BE802FFEF5213FF19F9DA84CAF925" blockId="89.[151,1436,1328,1691]" pageId="89" pageNumber="90">
|
||
<emphasis id="06803410FFEF5213FF19F9DA85A8F9EE" bold="true" box="[198,423,1590,1616]" pageId="89" pageNumber="90">Material studied.</emphasis>
|
||
<taxonomicName id="F3F49381FFEF5213FE66F9DA87A4F9EE" authority="(Lyman, 1883)" baseAuthorityName="Lyman" baseAuthorityYear="1883" box="[441,939,1590,1616]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="cervicornis">
|
||
<emphasis id="06803410FFEF5213FE66F9DA86D1F9F1" box="[441,734,1590,1615]" italics="true" pageId="89" pageNumber="90">Ophioprium cervicornis</emphasis>
|
||
(
|
||
<bibRefCitation id="506595F3FFEF5213FD28F9DA87ADF9EE" author="Lyman, T." box="[759,930,1590,1616]" pageId="89" pageNumber="90" pagination="227 - 287" refId="ref90277" refString="Lyman, T. (1883) Report on the Ophiuroidea. Bulletin of the Museum of Comparative Zoology at Harvard College 10, 227 - 287." type="journal article" year="1883">Lyman, 1883</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<materialsCitation id="849CE25FFFEF5213FC1EF9DA867AF9CB" collectionCode="MCZ" pageId="89" pageNumber="90" specimenCode="MCZ 2043" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="EB4F56A0FFEF5213FC1EF9DA8021F9EE" box="[961,1070,1590,1616]" pageId="89" pageNumber="90" type="holotype">Holotype</typeStatus>
|
||
<specimenCode id="64524079FFEF5213FB9FF9DA80C4F9EE" box="[1088,1227,1590,1616]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="89" pageNumber="90" type="Museum">MCZ 2043</specimenCode>
|
||
(
|
||
<figureCitation id="ACCFF487FFEF5213FB3AF9DA8183F9EE" box="[1253,1420,1590,1616]" captionStart="FIGURE 57" captionStartId="83.[151,255,1564,1588]" captionTargetBox="[192,1395,194,1543]" captionTargetId="figure-210@83.[192,1395,194,1543]" captionTargetPageId="83" captionText="FIGURE 57. Genera Ophioprium and Ophiopristis, external views. A-B, Ophioprium cervicornis (Lyman, 1883), holotype MCZ 2043, 10 mm dd, dorsal and ventral views respectively; C-D, Ophiopristis hirsuta (Lyman, 1875), holotype MCZ 1950, 5.8 mm dd, dorsal and ventral views respectively; E. Ophioprium cervicornis (Lyman, 1883), holotype MCZ 2043, arm spine articulations, proximal segments; F. Ophiopristis hirsuta (Lyman, 1875), paratype MCZ 1951, 5 mm dd, ventral view. Photos: Alexander Martynov." pageId="89" pageNumber="90">Figs 57A, B</figureCitation>
|
||
);
|
||
<taxonomicName id="F3F49381FFEF5213FF48F9B7867AF9CB" authority="(Koehler, 1914)" baseAuthorityName="Koehler" baseAuthorityYear="1914" box="[151,629,1627,1653]" class="Ophiuroidea" family="Ophioscolecidae" genus="Ophioprium" kingdom="Animalia" order="Ophioscolecida" pageId="89" pageNumber="90" phylum="Echinodermata" rank="species" species="permixtum">
|
||
<emphasis id="06803410FFEF5213FF48F9B785A2F9CA" box="[151,429,1627,1652]" italics="true" pageId="89" pageNumber="90">Ophioprium permixtum</emphasis>
|
||
(
|
||
<bibRefCitation id="506595F3FFEF5213FE61F9B78663F9CB" author="Koehler, R." box="[446,620,1627,1653]" pageId="89" pageNumber="90" pagination="1 - 173" refId="ref88727" refString="Koehler, R. (1914) A contribution to the study of Ophiurans of the United States National Museum. Bulletin of the United States National Museum 84, 1 - 173." type="journal article" year="1914">Koehler, 1914</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="849CE25FFFEF5213FD5EF9B784CEF925" accessionNumber="E 05300" collectionCode="USNM" pageId="89" pageNumber="90" specimenCode="USNM 032296" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="EB4F56A0FFEF5213FD5EF9B786E0F9CB" box="[641,751,1627,1653]" pageId="89" pageNumber="90" type="holotype">Holotype</typeStatus>
|
||
<specimenCode id="64524079FFEF5213FD28F9B787BEF9CB" box="[759,945,1627,1653]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="89" pageNumber="90" type="Museum">USNM 032296</specimenCode>
|
||
;
|
||
<specimenCount id="22F2238BFFEF5213FC62F9B78050F9CB" box="[957,1119,1627,1653]" pageId="89" pageNumber="90" type="generic">one specimen</specimenCount>
|
||
<collectionCode id="52E570C7FFEF5213FBB7F9B780B2F9CB" box="[1128,1213,1627,1653]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="89" pageNumber="90" type="Museum">USNM</collectionCode>
|
||
E 05300 (
|
||
<figureCitation id="ACCFF487FFEF5213FAE6F9B78199F9CB" box="[1337,1430,1627,1653]" captionStart="FIGURE 61" captionStartId="88.[151,255,1911,1935]" captionTargetBox="[158,1428,679,1890]" captionTargetId="figure-295@88.[158,1428,679,1890]" captionTargetPageId="88" captionText="FIGURE 61. Ophioprium permixtum (Koehler, 1914), USNM E 05300, 15 mm dd, external views. A. dorsal view; B. ventral view; C. dorsal disk, details; D. dorsal disk, details showing small radial shields; E. ventral view, details oral and adoral shield papillae; F. interradial view showing long wide genital slits; G. ventral view, showing details of the teeth. Photos: Tatiana Korshunova." pageId="89" pageNumber="90">Figs 61</figureCitation>
|
||
;
|
||
<figureCitation id="ACCFF487FFEF5213FF48F96D84B7F925" box="[151,184,1665,1691]" captionStart="FIGURE 62" captionStartId="89.[151,255,1163,1187]" captionTargetBox="[173,1410,182,1157]" captionTargetId="figure-329@89.[173,1410,182,1159]" captionTargetPageId="89" captionText="FIGURE 62. Ophioprium permixtum (Koehler, 1914), USNM E 05300, 15 mm dd, details. A-B, lateral plate and spine articulations, proximal segments; C, G, proximal spine, section showing small cavities; D. proximal vertebra, dorsal view; E. same, ventral view; F. same, distal view; H. proximal spine; I. proximal spine, ventral view. Scale bars: A–B, D– F, 300 µ; C, G, K, 100 µ; I, 1 mm. Photos: Alexander Martynov." pageId="89" pageNumber="90">62</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |