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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.144.62246" ID-GBIF-Dataset="3776e555-85f5-46ed-8ea1-98ca6a147ede" ID-PMC="PMC8222199" ID-Pensoft-Pub="1313-2970-1044-609" ID-Pensoft-UUID="C96ED07B505A52A8ADA354E1F70BB7C0" ID-PubMed="34183888" ID-ZooBank="66F01A49D32448A8AC2669BF3374894C" ModsDocID="1313-2970-1044-609" checkinTime="1623875354053" checkinUser="pensoft" docAuthor="Smith, Sarah M. &amp; Cognato, Anthony I." docDate="2021" docId="7A33407C57365418B68446B5FA164830" docLanguage="en" docName="ZooKeys 1044: 609-720" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.144.62246" docTitle="Coptoborus asperatus Smith &amp; Cognato 2021, sp. nov." docType="treatment" docUuid="E2AA8F62-3AEA-4398-9570-6151576AE9DD" docUuidSource="ZooBank" docVersion="4" id="C96ED07B505A52A8ADA354E1F70BB7C0" lastPageNumber="609" masterDocId="C96ED07B505A52A8ADA354E1F70BB7C0" masterDocTitle="A revision of the Neotropical genus Coptoborus Hopkins (Coleoptera, Curculionidae, Scolytinae, Xyleborini)" masterLastPageNumber="720" masterPageNumber="609" pageNumber="609" updateTime="1668150492206" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Neotropical genus Coptoborus Hopkins (Coleoptera, Curculionidae, Scolytinae, Xyleborini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
<mods:nameIdentifier type="email">camptocerus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cognato, Anthony I.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1044</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>609</mods:start>
<mods:end>720</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.144.62246</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.144.62246</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1044-609</mods:identifier>
<mods:identifier type="ZooBank">66F01A49D32448A8AC2669BF3374894C</mods:identifier>
<mods:identifier type="Pensoft-UUID">C96ED07B505A52A8ADA354E1F70BB7C0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182753903" LSID="urn:lsid:zoobank.org:act:E2AA8F62-3AEA-4398-9570-6151576AE9DD" httpUri="http://treatment.plazi.org/id/7A33407C57365418B68446B5FA164830" lastPageNumber="609" pageId="0" pageNumber="609">
<subSubSection pageId="0" pageNumber="609" type="nomenclature">
<paragraph pageId="0" pageNumber="609">
<taxonomicName LSID="http://zoobank.org/E2AA8F62-3AEA-4398-9570-6151576AE9DD" authority="Smith &amp; Cognato, 2021" authorityName="Smith &amp; Cognato" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Coptoborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptoborus asperatus" order="Coleoptera" pageId="0" pageNumber="609" phylum="Arthropoda" rank="species" species="asperatus" status="sp. nov.">Coptoborus asperatus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="609">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal, lateral, frontal and declivital view of Coptoborus amazonicus holotype, 2.8 mm (A-C, M), C. amplissimus holotype, 2.8 mm (D-F, N), C. asperatus holotype, 2.0 mm (G-I, O), C. attenuatus holotype, 2.0 mm (J-L, P). All photographs by SMS except (A-C, M) by A. V. Petrov and (J-L, P) by R. K. Osborn." figureDoi="10.3897/zookeys.144.62246.figure1" httpUri="https://binary.pensoft.net/fig/555836" pageId="0" pageNumber="609">Figure 1G-I, O</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="type material">
<paragraph pageId="0" pageNumber="609">Type material.</paragraph>
<paragraph pageId="0" pageNumber="609">
<emphasis bold="true" pageId="0" pageNumber="609">
<emphasis italics="true" pageId="0" pageNumber="609">Holotype</emphasis>
,
</emphasis>
female, Ecuador: Napo Prov. [= Orellana Prov.], Res[erva]. Ethnica Waorani, 1 km S. Okone Gare Camp, Trans[ect]. Ent[omology].,
<geoCoordinate degrees="00" direction="south" minutes="39" orientation="latitude" precision="15" seconds="10" value="-0.65277773">00°39'10&quot;S</geoCoordinate>
,
<geoCoordinate degrees="076" direction="west" minutes="26" orientation="longitude" precision="925" value="-76.433334">076°26'W</geoCoordinate>
, 220 m, October 1994, T.L. Erwin et al. collectors, insecticidal fogging, terra firme forest, trans[ect] 6, sta[tion] 3, Erwin lot #922 (ICB).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="diagnosis">
<paragraph pageId="0" pageNumber="609">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="609">
2.0 mm (n = 1), 2.86
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. This species is distinguished by the elytral apex attenuate and strongly emarginate, declivity convex, declivital interstriae 2 denticulate, elytral apex with interstriae 3 and 9 joining, forming a crenulate carina that continues submarginally to apex, crenulations equal in size, declivital interstriae 3 densely denticulate with fewer than ten denticles, declivital interstriae 1 with three rows of setae, and declivital striae not impressed.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="similar species">
<paragraph pageId="0" pageNumber="609">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="609">
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. carumbensis" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="carumbensis">
<emphasis italics="true" pageId="0" pageNumber="609">C. carumbensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="description">
<paragraph pageId="0" pageNumber="609">Description</paragraph>
<paragraph pageId="0" pageNumber="609">
<emphasis bold="true" pageId="0" pageNumber="609">
(female).
<emphasis italics="true" pageId="0" pageNumber="609">Holotype</emphasis>
</emphasis>
2.0 mm, 2.86
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Body uniformly brown, antennae and legs lighter.
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Head</emphasis>
: epistoma tuberculate. Frons dull, finely punctate, glabrous. Eyes broadly and moderately emarginate. Submentum narrow, triangular, slightly impressed. Antennal scape short and thick, much shorter than club. Pedicel shorter than funicle. Club circular, flat, type 3; segment 1 corneous, procurved on anterior face, occupying basal ~1/3; segment 2 narrow, subconvex, corneous; segments 1 and 2 present on posterior face.
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Pronotum</emphasis>
: 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. In dorsal view long and rounded frontally, type 7, sides parallel in basal 2/3, rounded anteriorly; anterior margin without serrations. In lateral view elongate, disc as long as anterior slope, type 7, summit indistinct, at midpoint. Anterior slope with densely spaced, broad fine asperities, becoming lower and more strongly transverse towards summit. Disc strongly shiny with moderately dense, minute punctures, some longer hair-like setae at margins. Lateral margins obliquely costate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Elytra</emphasis>
: 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as pronotum. Scutellum small. Elytra attenuate, parallel-sided in basal 2/3, then acutely tapered to apex, apex weakly emarginate. Disc smooth, shiny; strial punctures large, deep, each bearing a recumbent seta the length of a puncture; interstriae flat, sparsely, minutely punctate, unarmed, each puncture bearing a long, erect seta. Declivity gradual, occupying ~1/3 of elytra, shagreened, dull, declivital face convex; striae not impressed, strial punctures larger, deeper than those of disc, each puncture bearing a semi-erect seta as long as two punctures; interstriae flat, interstriae denticulate along their entire lengths, interstriae 3 sparsely denticulate, denticles separated by at least the width of three denticles and with eight or fewer denticles, interstrial setae erect, bristle-like, uniseriate, interstriae 1 with two additional rows of slightly shorter erect hair-like setae. Posterolateral margin with interstriae 3 and 9 joining, forming a granulate carina and continuing submarginally to apex.
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Legs</emphasis>
: protibiae obliquely triangular, broadest at apical 1/3; apical 1/2 of outer margin with five moderately sized socketed denticles, their length as long as basal width. Meso- and metatibiae flattened; outer margin evenly rounded with seven and six moderately sized socketed denticles, respectively.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="etymology">
<paragraph pageId="0" pageNumber="609">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="609">
L.
<emphasis italics="true" pageId="0" pageNumber="609">asperatus</emphasis>
= rough. In reference to the
<normalizedToken originalValue="species">species'</normalizedToken>
sculptured declivity. Adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="distribution">
<paragraph pageId="0" pageNumber="609">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="609">Ecuador (Orellana).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="biology_ecology">
<paragraph pageId="0" pageNumber="609">Biology.</paragraph>
<paragraph pageId="0" pageNumber="609">The holotype was collected by canopy fogging.</paragraph>
</subSubSection>
</treatment>
</document>