272 lines
23 KiB
XML
272 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.788.26282" ID-GBIF-Dataset="4878bde6-41c0-44fe-9437-1fdf5857a6ff" ID-PMC="PMC6189228" ID-Pensoft-Pub="1313-2970-788-201" ID-PubMed="30337830" ID-ZBK="6F7FD9E2E936440D9CD542D6F8961D2F" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-788-201" ModsDocOrigin="ZooKeys 788" ModsDocTitle="Six new species and one new subspecies of noctuid moths from western United States of America and Mexico (Lepidoptera, Noctuidae)" checkinTime="1539050135329" checkinUser="pensoft" docAuthor="Crabo, Lars G., Hammond, Paul C., Mustelin, Tomas & Wikle, David L." docDate="2018" docId="5DE44ECFD20ECB5FEB606E75B1A04DEF" docLanguage="en" docName="ZooKeys 788: 201-239" docOrigin="ZooKeys 788" docSource="http://dx.doi.org/10.3897/zookeys.788.26282" docTitle="Plagiomimicus yakama Crabo & Wikle, sp. n." docType="treatment" docUuid="D50EEB56-B99D-4FAB-8DB3-0960A8117802" docUuidSource="ZooBank" docVersion="6" lastPageNumber="210" masterDocId="214CFFB25134FFE1FFC6F4538A421B48" masterDocTitle="Six new species and one new subspecies of noctuid moths from western United States of America and Mexico (Lepidoptera, Noctuidae)" masterLastPageNumber="239" masterPageNumber="201" pageNumber="206" updateTime="1668166263055" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Six new species and one new subspecies of noctuid moths from western United States of America and Mexico (Lepidoptera, Noctuidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Crabo, Lars G.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hammond, Paul C.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mustelin, Tomas</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wikle, David L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>788</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>201</mods:start>
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<mods:end>239</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.788.26282</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.788.26282</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-788-201</mods:identifier>
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<mods:identifier type="ZBK">6F7FD9E2E936440D9CD542D6F8961D2F</mods:identifier>
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<mods:identifier type="ZooBank">6F7FD9E2E936440D9CD542D6F8961D2F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="148657009" LSID="urn:lsid:zoobank.org:act:D50EEB56-B99D-4FAB-8DB3-0960A8117802" httpUri="http://treatment.plazi.org/id/5DE44ECFD20ECB5FEB606E75B1A04DEF" lastPageId="9" lastPageNumber="210" pageId="5" pageNumber="206">
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<subSubSection pageId="5" pageNumber="206" type="nomenclature">
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<paragraph pageId="5" pageNumber="206">
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<taxonomicName LSID="http://zoobank.org/D50EEB56-B99D-4FAB-8DB3-0960A8117802" authority="Crabo & Wikle" class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus yakama" order="Lepidoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="yakama">Plagiomimicus yakama Crabo & Wikle</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="206">sp. n.</taxonomicNameLabel>
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Figs 11-13, 17, 18, 21, 24, 49
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="207" type="type locality">
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<paragraph pageId="6" pageNumber="207">
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<pageBreakToken pageId="6" pageNumber="207" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph pageId="6" pageNumber="207">USA, Washington, Yakima County, Satus Creek.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="207" type="type material">
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<paragraph pageId="6" pageNumber="207">Type Material.</paragraph>
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<paragraph pageId="6" pageNumber="207">
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Holotype, male. [USA], Washington, Yakima County, Satus Creek, 30 V [19]49, E. C. Johnston. CNC. Paratypes. 10 males, 3 females. USA: Washington: Kittitas County: Schnebly Coulee,
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<geoCoordinate direction="north" orientation="latitude" precision="555" value="46.95">46.95°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="555" value="-120.09">120.09°W</geoCoordinate>
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, 500 m, 2 VI 1990, L G Crabo leg (4 males, 2 females); same locality & collector, 3 VII 1990 (1 male); Schnebly Coulee, 46.955°, -120.095°, 500 m, 14 V 2010, L. G. Crabo leg.; Grant County: Wanapum Dam, 1.6 mi [2.6 km] N, 46.900°, -119.948°, 250 m, 14 V 2010, L. G. Crabo leg (1 m); Yakima County: Satus Creek, 30 V [19]49, E. C. Johnston, (2 m); same locality, date, & collector / Genitalia CNC slide # 17068 (1 m); same locality, date, & collector / Genitalia slide by P6 USNM 45.585 (1 f); same locality, date, & collector / Slide No. 10,789 (1 m). CNC, DLW, LGC.
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</paragraph>
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<paragraph pageId="6" pageNumber="207">The type series is restricted to Washington State.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="207" type="differential diagnosis">
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<paragraph pageId="6" pageNumber="207">Differential diagnosis.</paragraph>
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<paragraph pageId="6" pageNumber="207">
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<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus yakama" order="Lepidoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="yakama">Plagiomimicus yakama</taxonomicName>
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(Figs 11-13) is distinguished from the two other species in the
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<taxonomicName lsidName="P. tepperi" pageId="6" pageNumber="207" rank="species" species="tepperi">P. tepperi</taxonomicName>
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species-group found in the United States in having an inconspicuous subapical spot, not darker than the adjacent medial area. Fresh
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<taxonomicName lsidName="P. tepperi" pageId="6" pageNumber="207" rank="species" species="tepperi">P. tepperi</taxonomicName>
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(Figure 14) and
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<taxonomicName lsidName="P. mimica" pageId="6" pageNumber="207" rank="species" species="mimica">P. mimica</taxonomicName>
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(not shown) are greener than
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<taxonomicName lsidName="P. yakama" pageId="6" pageNumber="207" rank="species" species="yakama">P. yakama</taxonomicName>
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.
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<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus incomitatus" order="Lepidoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="incomitatus">Plagiomimicus incomitatus</taxonomicName>
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sp. n. (Figs 15, 16), only found in Mexico, has a dark smudged reniform stigma unlike the pale or absent stigma of
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<taxonomicName lsidName="P. yakama" pageId="6" pageNumber="207" rank="species" species="yakama">P. yakama</taxonomicName>
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.
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</paragraph>
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<caption pageId="6" pageNumber="207">
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<paragraph pageId="6" pageNumber="207">
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Figures 11-16.
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<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus" order="Lepidoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">Plagiomimicus</taxonomicName>
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adults. 11
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<taxonomicName lsidName="P. yakama" pageId="6" pageNumber="207" rank="subSpecies" species="yakama" subSpecies="yakama">P. yakama yakama</taxonomicName>
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, holotype male, USA, Washington, Yakima County, Satus Creek 12
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<taxonomicName lsidName="P. yakama" pageId="6" pageNumber="207" rank="subSpecies" species="yakama" subSpecies="mojave">P. yakama mojave</taxonomicName>
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, holotype male, USA, Arizona, Mohave County, Hualapai Mountains, Wikieup 9.7 km W 13
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<taxonomicName lsidName="P. yakama" pageId="6" pageNumber="207" rank="subSpecies" species="yakama" subSpecies="mojave">P. yakama mojave</taxonomicName>
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, male, USA, Nevada, Clark County, Charleston Mountains, Kyle Canyon 14
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<taxonomicName lsidName="P. tepperi" pageId="6" pageNumber="207" rank="species" species="tepperi">P. tepperi</taxonomicName>
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, male, USA Arizona, Maricopa County, Cave Creek 15
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<taxonomicName lsidName="P. incomitatus" pageId="6" pageNumber="207" rank="species" species="incomitatus">P. incomitatus</taxonomicName>
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, holotype male, Mexico, Baja California Sur, Punta Colorada 11.3 km S 16
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<taxonomicName lsidName="P. incomitatus" pageId="6" pageNumber="207" rank="species" species="incomitatus">P. incomitatus</taxonomicName>
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, female, same locality as last specimen.
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</paragraph>
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</caption>
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<paragraph pageId="6" pageNumber="207">
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The ampullae of the male valve of
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<taxonomicName lsidName="P. yakama" pageId="6" pageNumber="207" rank="species" species="yakama">P. yakama</taxonomicName>
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(Figs 17a, 18a) are nearly symmetrical, short and needlelike, arising perpendicular to the valve or at a slight angle. Those of
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<taxonomicName lsidName="P. tepperi" pageId="6" pageNumber="207" rank="species" species="tepperi">P. tepperi</taxonomicName>
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(Figure 19a) and
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<taxonomicName lsidName="P. mimica" pageId="6" pageNumber="207" rank="species" species="mimica">P. mimica</taxonomicName>
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(not shown) are longer, curved, and directed distad. Those of
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<taxonomicName lsidName="P. mimica" pageId="6" pageNumber="207" rank="species" species="mimica">P. mimica</taxonomicName>
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are also asymmetrical. The female of
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<taxonomicName lsidName="P. yakama" pageId="6" pageNumber="207" rank="species" species="yakama">P. yakama</taxonomicName>
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(Figure 21) has a longer and more strongly curved corpus bursae than
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<taxonomicName lsidName="P. tepperi" pageId="6" pageNumber="207" rank="species" species="tepperi">P. tepperi</taxonomicName>
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(Figure 22).
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</paragraph>
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<paragraph pageId="6" pageNumber="207">
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The barcodes of
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<taxonomicName lsidName="P. yakama" pageId="6" pageNumber="207" rank="species" species="yakama">P. yakama</taxonomicName>
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(BOLD:ACR9301) and
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<taxonomicName lsidName="P. tepperi" pageId="6" pageNumber="207" rank="species" species="tepperi">P. tepperi</taxonomicName>
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(BOLD:AAF2198) differ by at least 1.63 %.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207" type="description">
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<paragraph pageId="6" pageNumber="207">Description.</paragraph>
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<paragraph lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207">
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Nominate subspecies. Adults. Males and females similar in size and habitus. Head. Antenna filiform, ventral surface ciliate, dense (male), sparse (female); dorsal scales gray olive. Scape off-white. Eye round, bare. Frontal process sideways D-shaped, straight side ventrad, lateral and dorsal rims raised slightly, central process slightly caudal to
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<normalizedToken originalValue="“D”">"D"</normalizedToken>
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center, cone shaped, protruding slightly beyond edges; a transverse ridge caudal to process; frons and dorsal head scales short, tan off-white, palest near vertex. Labial palpus reaching dorsal eye, second segment long, third segment very short, scales short, light olive gray, darker than head. Haustellum normal. Thorax. Dorsum, including patagium and tegula, scales short, olive off-white; appearing uniform pale brownish olive gray similar to head and forewing base. Venter lighter. Legs: Pale olive gray; distal foretibia claw short, thornlike, tarsal segments equal length. Wings: Forewing: Length 11.5-12.5 mm; elongate with slightly pointed apex, lateral margin straight to CuA1, thence convex to posterior margin; scales mixed olive off-white, tan, and gray olive; base to antemedial line and basal postmedial area silver gray, distal postmedial area, terminal
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<pageBreakToken pageId="7" pageNumber="208" start="start">area</pageBreakToken>
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, and subapical spot slightly darker gray, medial area dark olive gray; cubital vein basal to postmedial line slightly lighter; basal and medial lines absent; antemedial and postmedial lines white, wide, slightly indistinct; antemedial line oblique from mid-costa to inner third posterior margin, slightly convex; postmedial line from outer third costa to R5 angled strongly distad, bent basad acutely on R5 to cubital vein at end of cell, thence parallel to antemedial line to outer ⅓ of posterior margin; subterminal line pale gray, white adjacent to subapical spot, slightly sinuous; terminal line thin, slightly darker than terminal area or absent; subapical spot slightly lighter than medial area, elongate, cau
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<pageBreakToken pageId="8" pageNumber="209" start="start">dal</pageBreakToken>
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margin smoothly convex; fringe olive off-white, base slightly darker; claviform and orbicular stigmata absent; reniform stigma absent or few pale anterior and darker posterior scales. Hindwing: Dorsum uniform medium gray, postmedial area slightly lighter in some specimens; terminal line thin, slightly darker; fringe whitish, base pale olive gray. Abdomen. Male lacking basal modifications; scales pale fuscous. Male genitalia: Uncus short, thick, curved slightly, point short, covered by sparse short fine setae. Juxta shield shape, height = width. Valve length 2.4-2.7
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<normalizedToken originalValue="×">x</normalizedToken>
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width, simple, outer margin convex, apex blunt, medial surface with sparse fine setae; cucullus unmodified, corona absent; sacculus ⅔
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<normalizedToken originalValue="×">x</normalizedToken>
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valve length, 0.4
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<normalizedToken originalValue="×">x</normalizedToken>
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width, basal process short, spike like, perpendicular to valve, mesial dorsum variable, smooth or with short broad triangular process; clasper base weak, origin near ventral margin; ampulla short, 0.14-0.18
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<normalizedToken originalValue="×">x</normalizedToken>
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valve width, right slightly longer, thin, acute, nearly perpendicular to valve; digitus absent. Phallus cylindrical, straight, length 4
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<normalizedToken originalValue="×">x</normalizedToken>
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width; vesica as long and slightly wider than phallus, straight beyond basal 120° bend, mesial and distal surface with large patch of similar-sized basally directed spike-like cornuti. Female genitalia: Papilla analis 2
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<normalizedToken originalValue="×">x</normalizedToken>
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segment A8 length, width 0.33
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<normalizedToken originalValue="×">x</normalizedToken>
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length, sclerotized, distal ⅓ tapered evenly to acute apex, setae short, sparse, membrane to segment A8 leathery; posterior apophysis 3.3
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<normalizedToken originalValue="×">x</normalizedToken>
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segment A8 length; anterior apophysis 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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posterior apophysis; ostium bursae sclerotized, cup shaped, slightly wider than ductus bursae; ductus bursae 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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segment A8 length, sclerotized lightly; corpus bursae elongate, length 3.5
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<normalizedToken originalValue="×">x</normalizedToken>
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width, posterior ⅓ sclerotized; anterior ⅔ membranous, lacking signa, constricted mesially to width of ductus bursae, anterior end bulbous, curved slightly rightward; appendix bursae short, truncate, moderately sclerotized, joined broadly to corpus bursae at junction with ductus bursae; ductus seminalis at ventral apex.
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</paragraph>
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<caption pageId="8" pageNumber="209">
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<paragraph pageId="8" pageNumber="209">
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Figures 17-20.
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<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus" order="Lepidoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">Plagiomimicus</taxonomicName>
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male genitalia. 17
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<taxonomicName lsidName="P. y." pageId="8" pageNumber="209" rank="subSpecies" species="y." subSpecies="yakama">P. y. yakama</taxonomicName>
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a valves b phallus with everted vesica 18
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<taxonomicName lsidName="P. yakama" pageId="8" pageNumber="209" rank="subSpecies" species="yakama" subSpecies="mojave">P. yakama mojave</taxonomicName>
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a valves b phallus with everted vesica 19
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<taxonomicName lsidName="P. tepperi" pageId="8" pageNumber="209" rank="species" species="tepperi">P. tepperi</taxonomicName>
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a valves b phallus with everted vesica 20
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<taxonomicName lsidName="P. incomitatus" pageId="8" pageNumber="209" rank="species" species="incomitatus">P. incomitatus</taxonomicName>
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a valves b phallus with everted vesica.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="209" type="geographic variation">
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<paragraph pageId="8" pageNumber="209">Geographic variation.</paragraph>
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<paragraph pageId="8" pageNumber="209">
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Populations of
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<taxonomicName lsidName="P. yakama" pageId="8" pageNumber="209" rank="species" species="yakama">P. yakama</taxonomicName>
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are arranged in northern and southern subspecies, described below.
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</paragraph>
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<paragraph pageId="8" pageNumber="209">
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Barcodes samples of
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<taxonomicName lsidName="P. yakama" pageId="8" pageNumber="209" rank="species" species="yakama">P. yakama</taxonomicName>
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exist for Washington (n = 2), Utah (n = 3), Nevada (n = 1), Arizona (n = 6), and California (n = 3). Washington samples differ from the others by 0.3 %.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="209" type="etymology">
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<paragraph pageId="8" pageNumber="209">Etymology.</paragraph>
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<paragraph pageId="8" pageNumber="209">The name refers to the Yakama people, the original human inhabitants of the type locality of this moth. It is a noun in apposition. The spelling of the Yakama Nation differs from the more familiar spellings of the city of Yakima and Yakima County, Washington.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="209" type="distribution">
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<paragraph pageId="8" pageNumber="209">Distribution and ecology.</paragraph>
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<paragraph pageId="8" pageNumber="209">
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<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus yakama" order="Lepidoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="yakama">Plagiomimicus yakama</taxonomicName>
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is found in two separate parts of the American West (Figure 49). The nominate subspecies occurs on the Columbia Plateau. Subspecies
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<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus yakama" order="Lepidoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="subSpecies" species="yakama" subSpecies="mojave">Plagiomimicus yakama mojave</taxonomicName>
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is found 1000 km farther south in the southern Great Basin and Mojave Desert.
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</paragraph>
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<paragraph pageId="8" pageNumber="209">
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This moth flies in dry desert habitats with exposed soil and rocks, favoring those with varied topography such as rim rock, coulees, and arroyos. Where known, the larva of
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<taxonomicName lsidName="P. yakama" pageId="8" pageNumber="209" rank="species" species="yakama">P. yakama</taxonomicName>
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feeds on
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Brickellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Brickellia" order="Asterales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">Brickellia</taxonomicName>
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species. The larva of subspecies
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<taxonomicName lsidName="P. y." pageId="8" pageNumber="209" rank="subSpecies" species="y." subSpecies="mojave">P. y. mojave</taxonomicName>
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is described below.
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</paragraph>
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<paragraph pageId="8" pageNumber="209">Adults fly during spring, from April to June, in all parts of the range. Southern populations have at least a partial fall brood.</paragraph>
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<caption pageId="8" pageNumber="209">
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<paragraph pageId="8" pageNumber="209">
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Figures 21-23.
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<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus" order="Lepidoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">Plagiomimicus</taxonomicName>
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female genitalia 21
|
||
<taxonomicName lsidName="P. y." pageId="8" pageNumber="209" rank="subSpecies" species="y." subSpecies="yakama">P. y. yakama</taxonomicName>
|
||
, female 22
|
||
<taxonomicName lsidName="P. tepperi" pageId="8" pageNumber="209" rank="species" species="tepperi">P. tepperi</taxonomicName>
|
||
, female 23
|
||
<taxonomicName lsidName="P. incomitatus" pageId="8" pageNumber="209" rank="species" species="incomitatus">P. incomitatus</taxonomicName>
|
||
, female.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="210" type="discussion">
|
||
<paragraph pageId="9" pageNumber="210">
|
||
<pageBreakToken pageId="9" pageNumber="210" start="start">Discussion</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="210">
|
||
Syntypes of
|
||
<taxonomicName class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schinia tepperi" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="tepperi">Schinia tepperi</taxonomicName>
|
||
Morrison, 1875 at MSU and the holotype of its synonym
|
||
<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus richii" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="richii">Plagiomimicus richii</taxonomicName>
|
||
Grote, 1886 at NHML, both described from
|
||
<normalizedToken originalValue="“Texas”">"Texas"</normalizedToken>
|
||
, match examined material of
|
||
<taxonomicName class="Insecta" family="Noctuidae" genus="Plagiomimicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiomimicus tepperi" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="tepperi">Plagiomimicus tepperi</taxonomicName>
|
||
from Texas, New Mexico, and southeastern and central Arizona. When restricted to these populations
|
||
<taxonomicName lsidName="P. tepperi" pageId="9" pageNumber="210" rank="species" species="tepperi">P. tepperi</taxonomicName>
|
||
is uniform in habitus and structure. Barcodes of
|
||
<taxonomicName lsidName="P. tepperi" pageId="9" pageNumber="210" rank="species" species="tepperi">P. tepperi</taxonomicName>
|
||
from southeast Arizona (n = 6) and New Mexico (n = 1) differ by 0.3 %. Barcodes of topotypical Texas populations have not been sampled. The range of
|
||
<taxonomicName lsidName="P. tepperi" pageId="9" pageNumber="210" rank="species" species="tepperi">P. tepperi</taxonomicName>
|
||
extends northwards to Colorado.
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="210">
|
||
In naming
|
||
<taxonomicName lsidName="P. yakama" pageId="9" pageNumber="210" rank="species" species="yakama">P. yakama</taxonomicName>
|
||
and restricting
|
||
<taxonomicName lsidName="P. tepperi" pageId="9" pageNumber="210" rank="species" species="tepperi">P. tepperi</taxonomicName>
|
||
in the above sense, the species in the
|
||
<taxonomicName lsidName="P. tepperi" pageId="9" pageNumber="210" rank="species" species="tepperi">P. tepperi</taxonomicName>
|
||
species-group become more uniform well defined entities. However, ongoing rearing and molecular work by DLW and David Wagner suggest that there could be additional unrecognized cryptic species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |