275 lines
21 KiB
XML
275 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne & Haring, Elisabeth" docDate="2021" docId="418C14069E195FBCB8E7E2B00CAC0B07" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Charpentieria dyodon subsp. thomasiana" docType="treatment" docVersion="4" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>De Mattia, Willy</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reier, Susanne</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Haring, Elisabeth</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1077</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>175</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
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||
<mods:identifier type="ZooBank">C28AD65A76F242CFBED7DFB3702CABCE</mods:identifier>
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<mods:identifier type="Pensoft-UUID">734088641608531C8E2CC69397B000ED</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="191645955" LSID="urn:lsid:plazi:treatment:418C14069E195FBCB8E7E2B00CAC0B07" httpUri="http://treatment.plazi.org/id/418C14069E195FBCB8E7E2B00CAC0B07" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="418C1406-9E19-5FBC-B8E7-E2B00CAC0B07" authority="(Kuester, 1850)" baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
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Charpentieria dyodon thomasiana (
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<normalizedToken originalValue="Küster">Kuester</normalizedToken>
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, 1850)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">Figs 41.1-41.4, 42.3, 42.4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="W. De Mattia, J. Macor" country="Italy" elevation="1100" latitude="45.63028" location="Santuario di Oropa." longLatPrecision="1" longitude="7.978353" municipality="Biella" specimenCount="1" stateProvince="Piedmont">
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<collectingCountry name="Italy">Italy</collectingCountry>
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,
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<collectingRegion country="Italy" name="Piemonte">Piedmont</collectingRegion>
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,
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<collectingMunicipality>Biella</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:418C14069E195FBCB8E7E2B00CAC0B07:F8278824FC4FBF69DDBAFBF741B434BB" country="Italy" latitude="45.63028" longLatPrecision="1" longitude="7.978353" municipality="Biella" name="Santuario di Oropa." stateProvince="Piedmont">Santuario di Oropa.</location>
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</quantity>
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asl
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</elevation>
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<geoCoordinate degrees="45" direction="north" minutes="37" orientation="latitude" precision="1" seconds="49.01" value="45.63028">45°37'49.01"N</geoCoordinate>
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,
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<geoCoordinate degrees="07" direction="east" minutes="58" orientation="longitude" precision="1" seconds="42.07" value="7.978353">07°58'42.07"E</geoCoordinate>
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,
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<collectorName>W. De Mattia</collectorName>
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and
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<collectorName>J. Macor</collectorName>
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leg. and det., 3 dissected spm
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</materialsCitation>
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.
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Italy" county="Biella" elevation="1200" latitude="45.692326" location="Pra del Vecia" longLatPrecision="1" longitude="7.967325" municipality="Sentiero" specimenCount="1" stateProvince="Piedmont">
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<collectingCountry name="Italy">Italy</collectingCountry>
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,
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<collectingRegion country="Italy" name="Piemonte">Piedmont</collectingRegion>
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,
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<collectingCounty>Biella</collectingCounty>
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,
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<collectingMunicipality>Sentiero</collectingMunicipality>
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per
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<location LSID="urn:lsid:plazi:treatment:418C14069E195FBCB8E7E2B00CAC0B07:B358FEC85D15F47DFEC16D06C3D01684" country="Italy" county="Biella" latitude="45.692326" longLatPrecision="1" longitude="7.967325" municipality="Sentiero" name="Pra del Vecia" stateProvince="Piedmont">
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<normalizedToken originalValue="Prà">Pra</normalizedToken>
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del Vecia
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</location>
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, Piedicavallo.
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
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asl
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</elevation>
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,
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<geoCoordinate degrees="45" direction="north" minutes="41" orientation="latitude" precision="1" seconds="32.37" value="45.692326">45°41'32.37"N</geoCoordinate>
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,
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<geoCoordinate degrees="07" direction="east" minutes="58" orientation="longitude" precision="1" seconds="2.37" value="7.967325">07°58'2.37"E</geoCoordinate>
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, leg. and det., 2 dissected spm
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
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<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon thomasiana</emphasis>
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</taxonomicName>
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is found in the surroundings of Oropa and Piedicavallo, surroundings of Biella (Piedmont).
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</paragraph>
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<caption doi="10.3897/zookeys.1077.67081.figure40" httpUri="https://binary.pensoft.net/fig/622832" pageId="0" pageNumber="1" start="Figure 40" startId="F40">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 40.</emphasis>
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<taxonomicName authority="dyodon" authorityName="dyodon (S. Studer" authorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="dyodon">
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<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon dyodon</emphasis>
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</taxonomicName>
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(S. Studer, 1820), Verbano
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<emphasis bold="true" pageId="0" pageNumber="1">40.1</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">40.2</emphasis>
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internal distal part of genital organs.
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<taxonomicName baseAuthorityName="Stabile" baseAuthorityYear="1859" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. alpina" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="alpina">
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<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon alpina</emphasis>
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</taxonomicName>
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(Stabile, 1859). Val di Lanzo, Usseglio
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<emphasis bold="true" pageId="0" pageNumber="1">40.3</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">40.4</emphasis>
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internal distal part of genital organs.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs">
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<paragraph pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Fig.
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<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">41</figureCitation>
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.1, 41.3).
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</emphasis>
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The FO is half as long as the V. The FDBC is shorter than the BC+SDBC (SDBC+BC). The BC+SDBC is club-like and longer than the V with slight distinction between the SDBC and the BC. The apex is pointed. The D is slightly longer than the BC+SDBC, thinner with a round apex. The V is long and cylindrical. The PC is longer than the V. The PR is long and robust. The E is slightly shorter than the P and thin.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs">
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<paragraph pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Fig.
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<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">41</figureCitation>
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.2, 41.4).
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</emphasis>
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The A and the P are smooth, with a very fine granulated sculpturing. The V shows many smooth longitudinal pleats. The smooth simple PP is simple, elongated with a pointed apex. Its base is partially connected to the ELP and transversally extends along the transition wall, narrowing the transition passage. The epiphallar formula is: PP(ELP). The E shows two main longitudinal moderately fringed pleats that proximally fade before the VD.
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</paragraph>
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<caption doi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1" start="Figure 41" startId="F41">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 41.</emphasis>
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<taxonomicName baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
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<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon thomasiana</emphasis>
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</taxonomicName>
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(
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<normalizedToken originalValue="Küster">Kuester</normalizedToken>
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, 1850), Piedicavallo
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<emphasis bold="true" pageId="0" pageNumber="1">41.1</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">41.2</emphasis>
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internal distal part of genital organs. Oropa, Biella
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<emphasis bold="true" pageId="0" pageNumber="1">41.3</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">41.4</emphasis>
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internal distal part of genital organs.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="brief genital anatomical description of the subspecific taxa of charpentieria ornata and charpentieria itala">
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<paragraph pageId="0" pageNumber="1">
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Brief genital anatomical description of the subspecific taxa of
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<taxonomicName baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria ornata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ornata">
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<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria ornata</emphasis>
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</taxonomicName>
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and
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<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
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<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The Southern Alpine
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<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
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<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
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</taxonomicName>
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is distributed from Como Lake to the Garda Lake, Trentino, Veneto (including Colli Berici and Colli Euganei) and the Pre-Alps of Friuli.
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="169 - 203" refId="B58" refString="Nordsieck, H, 1963b. Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen. Archiv fuer Molluskenkunde 92: 169 - 203" title="Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen." volume="92" year="1963 b">Nordsieck (1963b)</bibRefCitation>
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provided, for the eastern border of its distribution, the axis Pordenone-Ampezzo. Recent collecting extended its eastern distribution as far Udine and Cividale in Friuli (WDM, personal unpublished data). Its eastern distribution range overlaps with
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<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
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<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
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</taxonomicName>
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and in few localities they are found syntopic (e.g., San Romedio, Trento, where
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<taxonomicName authority="stenzii" lsidName="C. stenzii subsp. stenzii" pageId="0" pageNumber="1" rank="subSpecies" species="stenzii" subSpecies="stenzii">
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<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii stenzii</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="C. itala subsp. serravalensis" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="serravalensis">
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<emphasis italics="true" pageId="0" pageNumber="1">C. itala serravalensis</emphasis>
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</taxonomicName>
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are found syntopic). Concerning the nomenclature of
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<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
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<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
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</taxonomicName>
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, we adopted the system by Scheel and Hausdorf (2012) that considered
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<taxonomicName lsidName="C. clavata" pageId="0" pageNumber="1" rank="species" species="clavata">
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<emphasis italics="true" pageId="0" pageNumber="1">C. clavata</emphasis>
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</taxonomicName>
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as a subspecies of
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<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
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<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption doi="10.3897/zookeys.1077.67081.figure42" httpUri="https://binary.pensoft.net/fig/622834" pageId="0" pageNumber="1" start="Figure 42" startId="F42">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 42.</emphasis>
|
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<taxonomicName authority="dyodon" authorityName="dyodon (S. Studer" authorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="dyodon">
|
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<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon dyodon</emphasis>
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</taxonomicName>
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(S. Studer, 1820), Verbano
|
||
<emphasis bold="true" pageId="0" pageNumber="1">42.1</emphasis>
|
||
shell.
|
||
<taxonomicName baseAuthorityName="Stabile" baseAuthorityYear="1859" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. alpina" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="alpina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon alpina</emphasis>
|
||
</taxonomicName>
|
||
(Stabile, 1859). Val di Lanzo, Usseglio
|
||
<emphasis bold="true" pageId="0" pageNumber="1">42.2</emphasis>
|
||
shell.
|
||
<taxonomicName baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon thomasiana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
|
||
, 1850), Piedicavallo
|
||
<emphasis bold="true" pageId="0" pageNumber="1">42.3</emphasis>
|
||
shell. Oropa, Biella
|
||
<emphasis bold="true" pageId="0" pageNumber="1">42.4</emphasis>
|
||
shell.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Its shell is weakly striated to frankly ribbed, with evident sutural papillae and a strong lunella, which is remarkably curved. Twelve out of the seventeen currently valid subspecies of
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
were anatomically investigated, revealing a common overall arrangement of the genital organs. The pseudopapilla can be either smooth or markedly wrinkled and occupies almost the whole penial volume.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |