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<document ID-DOI="http://doi.org/10.5281/zenodo.3790614" ID-GBIF-Dataset="ab7d8c5f-f732-4f73-9e1f-5fe2bae6c9a7" ID-GBIF-Taxon="163754308" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790614" ID-ZooBank="BB883B1E-E58F-4074-92EB-7E814E78F678" approvalRequired="19" approvalRequired_for_textStreams="19" checkinTime="1587996592918" checkinUser="plazi" docAuthor="Hopp, Katie &amp; Caterino, Michael" docDate="2009" docId="4157F97AFF9EFFFDFF7EFD94FD9816FF" docLanguage="en" docName="ZK_article_2083.pdf" docOrigin="ZooKeys 24 (24)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5c452e69" docStyleName="zookeys.2008.journal_article" docTitle="Cephennium anophthalmicum Brendel B 1889" docType="treatment" docVersion="5" lastPageId="7" lastPageNumber="38" masterDocId="BD6E8102FF9DFFFAFFF2FF92FFBD1451" masterDocTitle="Seven new species of Cephennium Müller &amp; Kunze (Coleoptera, Staphylinidae, Scydmaeninae, Cephenniini) from California with a key to native North American species" masterLastPageNumber="54" masterPageNumber="31" pageId="3" pageNumber="34" updateTime="1643563761986" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seven new species of Cephennium Müller &amp; Kunze (Coleoptera, Staphylinidae, Scydmaeninae, Cephenniini) from California with a key to native North American species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hopp, Katie</mods:namePart>
<mods:affiliation>,,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael</mods:namePart>
<mods:affiliation>Clemson University, Clemson, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-10-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>24</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>24</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>54</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.24.247</mods:identifier>
<mods:identifier type="GBIF-Dataset">ab7d8c5f-f732-4f73-9e1f-5fe2bae6c9a7</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576549</mods:identifier>
<mods:identifier type="ZooBank">BB883B1E-E58F-4074-92EB-7E814E78F678</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790614" ID-GBIF-Taxon="163754308" ID-Zenodo-Dep="3790614" LSID="urn:lsid:plazi:treatment:4157F97AFF9EFFFDFF7EFD94FD9816FF" httpUri="http://treatment.plazi.org/id/4157F97AFF9EFFFDFF7EFD94FD9816FF" lastPageId="7" lastPageNumber="38" pageId="3" pageNumber="34">
<subSubSection pageId="3" pageNumber="34" type="nomenclature">
<paragraph blockId="3.[140,678,518,581]" box="[140,678,518,545]" pageId="3" pageNumber="34">
<heading box="[140,678,518,545]" fontSize="11" level="2" pageId="3" pageNumber="34" reason="3">
<taxonomicName authority="Brendel, 1889" authorityName="Brendel B" authorityYear="1889" box="[140,678,518,545]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis bold="true" box="[140,678,518,545]" pageId="3" pageNumber="34">
<emphasis bold="true" box="[140,502,518,545]" italics="true" pageId="3" pageNumber="34">Cephennium anophthalmicum</emphasis>
<bibRefCitation author="Brendel E" box="[509,678,518,545]" pageId="3" pageNumber="34" pagination="193 - 197" refId="ref11450" refString="Brendel E (1889) Descriptions of new Scydmaenidae and Pselaphidae. Entomologica Americana 5: 193 - 197. http: // www. biodiversitylibrary. org / item / 42392" type="journal article" year="1889">Brendel, 1889</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="3.[140,678,518,581]" box="[140,398,554,581]" pageId="3" pageNumber="34">Figs 1A, 2A, 3A, 5A, 6</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="34" type="materials_examined">
<paragraph blockId="3.[140,1108,624,1671]" pageId="3" pageNumber="34">
<emphasis bold="true" box="[140,317,624,650]" pageId="3" pageNumber="34">
<emphasis box="[140,198,624,650]" inLineHeading="true" pageId="3" pageNumber="34" reason="1">Type</emphasis>
Material.
</emphasis>
Not seen. A
<typeStatus box="[475,574,624,650]" pageId="3" pageNumber="34">holotype</typeStatus>
was not designated for this species. However, it was described from a single specimen from Alameda County that was sifted from vegetable debris together with a large number of
<taxonomicName baseAuthorityName="Mannerheim" baseAuthorityYear="1852" box="[708,987,695,721]" class="Insecta" family="Leiodidae" genus="Pinodytes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="cryptophagoides">
<emphasis box="[708,987,695,721]" italics="true" pageId="3" pageNumber="34">Pinodytes cryptophagoides</emphasis>
</taxonomicName>
(currently
<taxonomicName baseAuthorityName="Mannerheim" baseAuthorityYear="1852" box="[140,447,730,756]" class="Insecta" family="Leiodidae" genus="Catopocerus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="cryptophagoides">
<emphasis box="[140,447,730,756]" italics="true" pageId="3" pageNumber="34">Catopocerus cryptophagoides</emphasis>
</taxonomicName>
) by Marie Fuchs (
<bibRefCitation author="Brendel E" box="[663,825,730,756]" pageId="3" pageNumber="34" pagination="193 - 197" refId="ref11450" refString="Brendel E (1889) Descriptions of new Scydmaenidae and Pselaphidae. Entomologica Americana 5: 193 - 197. http: // www. biodiversitylibrary. org / item / 42392" type="journal article" year="1889">Brendel 1889</bibRefCitation>
). We attempted to track down this specimen but were not able to locate it at either the Academy of Natural Sciences in Philadelphia, the original repository of the Brendel collection, or the Museum of Comparative Zoology at Harvard University, the current repository for the Brendel collection. However, we did see a specimen from the
<collectionCode box="[729,815,871,897]" pageId="3" pageNumber="34">MCZC</collectionCode>
that was determined as
<taxonomicName authorityName="Brendel B" authorityYear="1889" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis italics="true" pageId="3" pageNumber="34">C. anophthalmicum</emphasis>
</taxonomicName>
from Alameda County. Because the label data do not exactly match the information presented by
<bibRefCitation author="Brendel E" box="[484,657,941,967]" pageId="3" pageNumber="34" pagination="193 - 197" refId="ref11450" refString="Brendel E (1889) Descriptions of new Scydmaenidae and Pselaphidae. Entomologica Americana 5: 193 - 197. http: // www. biodiversitylibrary. org / item / 42392" type="journal article" year="1889">Brendel (1889)</bibRefCitation>
we believe this is only a topotype, and not a primary type. We choose not to designate a
<typeStatus box="[697,787,977,1003]" pageId="3" pageNumber="34">neotype</typeStatus>
here as it is possible that the original type specimen is still in existence somewhere.
</paragraph>
<paragraph blockId="3.[140,1108,624,1671]" pageId="3" pageNumber="34">
<emphasis bold="true" box="[185,427,1046,1072]" pageId="3" pageNumber="34">
<emphasis box="[185,286,1046,1072]" inLineHeading="true" pageId="3" pageNumber="34" reason="1">Material</emphasis>
Examined.
</emphasis>
“
<materialsCitation ID-GBIF-Occurrence="2608963405" collectionCode="MCZC" county="Alameda" location="Laundry" municipality="Co." pageId="3" pageNumber="34" specimenCount="1">
<collectingCounty box="[449,551,1047,1073]" pageId="3" pageNumber="34">Alameda</collectingCounty>
;
<collectingMunicipality box="[568,610,1047,1073]" pageId="3" pageNumber="34">Co.</collectingMunicipality>
CAL.”/ “
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9EFFFDFF7EFD94FD9816FF:CC211EB7FF9EFFF9FD25FB85FC801060" box="[727,829,1047,1073]" county="Alameda" municipality="Co." name="Laundry" pageId="3" pageNumber="34">Laundry</location>
;
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9EFFFDFF7EFD94FD9816FF:CC211EB7FF9EFFF9FCBCFB85FC361060" box="[846,907,1047,1073]" county="Alameda" municipality="Co." name="Farm" pageId="3" pageNumber="34">Farm</location>
”/ “H. C. FALL; COLLECTION”/ “
<taxonomicName box="[371,511,1082,1108]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis box="[371,511,1082,1108]" italics="true" pageId="3" pageNumber="34">Cephennium</emphasis>
</taxonomicName>
;
<taxonomicName authorityName="Brendel B" authorityYear="1889" box="[526,711,1082,1108]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis box="[526,711,1082,1108]" italics="true" pageId="3" pageNumber="34">anophthalmicum</emphasis>
</taxonomicName>
; Brend.” (1
<collectionCode box="[853,943,1082,1108]" pageId="3" pageNumber="34">MCZC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608963386" collectingDate="1952-05-30" collectorName="Marin Co. Cal. &amp; H. B. Leech" location="Mill Valley" pageId="3" pageNumber="34" specimenCount="1">
“
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9EFFFDFF7EFD94FD9816FF:CC211EB7FF9EFFF9FC3CFBA8FBF21005" box="[974,1103,1082,1108]" name="Mill Valley" pageId="3" pageNumber="34">Mill Valley</location>
;
<collectorName box="[140,321,1117,1143]" pageId="3" pageNumber="34">Marin Co. Cal.</collectorName>
;
<date box="[336,456,1117,1144]" pageId="3" pageNumber="34" value="1952-05-30">
<collectingDate box="[336,456,1117,1144]" pageId="3" pageNumber="34" value="1952-05-30">30.V.1952</collectingDate>
</date>
”/ “By sifting; forest duff”/ “
<collectorName box="[786,922,1117,1143]" pageId="3" pageNumber="34">H.B. Leech</collectorName>
; Collector”/ red square label/ “
<taxonomicName box="[308,448,1152,1178]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis box="[308,448,1152,1178]" italics="true" pageId="3" pageNumber="34">Cephennium</emphasis>
</taxonomicName>
</materialsCitation>
; sp.
<specimenCount box="[508,523,1153,1179]" pageId="3" pageNumber="34" type="female">♀</specimenCount>
; Cl. Besuchet;
<materialsCitation ID-GBIF-Occurrence="2608963390" collectingDate="1961-05" collectionCode="CASC" location="CALIFORNIA" pageId="3" pageNumber="34" specimenCount="1" stateProvince="California">
det.
<date box="[760,851,1152,1179]" pageId="3" pageNumber="34" value="1961-05">
<collectingDate box="[760,851,1152,1179]" pageId="3" pageNumber="34" value="1961-05">V 1961</collectingDate>
</date>
”/ “Collection of the;
<collectingRegion box="[140,314,1188,1214]" country="United States of America" name="California" pageId="3" pageNumber="34">CALIFORNIA</collectingRegion>
ACADEMY; of SCIENCES, San; Francisco, Calif.” (1
<collectionCode box="[934,1012,1188,1214]" pageId="3" pageNumber="34">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608963406" collectingDate="1970-04-29" collectionCode="FMNH" collectorName="T. R. Haig" county="SanMateo Co." isTailEnumeration="true" location="Loma Mar" pageId="3" pageNumber="34" specimenCount="1" stateProvince="California">
“
<location LSID="urn:lsid:plazi:treatment:4157F97AFF9EFFFDFF7EFD94FD9816FF:CC211EB7FF9EFFF9FBE2FB36FF0210B0" county="SanMateo Co." name="Loma Mar" pageId="3" pageNumber="34">Loma Mar</location>
;
<collectingCounty box="[202,362,1223,1249]" pageId="3" pageNumber="34">SanMateo Co.</collectingCounty>
Calif;
<date box="[435,572,1223,1249]" pageId="3" pageNumber="34" value="1970-04-29">
<collectingDate box="[435,572,1223,1249]" pageId="3" pageNumber="34" value="1970-04-29">IV-29-1970</collectingDate>
</date>
”/ “ex. Redwood; Litter”/ “Collector;
<collectorName box="[991,1097,1223,1249]" pageId="3" pageNumber="34">T.R.Haig</collectorName>
” (1
<collectionCode box="[171,260,1258,1284]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="34" type="Museum">FMNH</collectionCode>
[female, disarticulated])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="35" pageId="3" pageNumber="34" type="diagnosis">
<paragraph blockId="3.[140,1108,624,1671]" pageId="3" pageNumber="34">
<emphasis bold="true" box="[185,319,1293,1319]" pageId="3" pageNumber="34">
<emphasis box="[185,314,1293,1319]" inLineHeading="true" pageId="3" pageNumber="34" reason="1">Diagnosis</emphasis>
.
</emphasis>
This species can be distinguished from its
<collectingRegion box="[857,979,1293,1319]" country="United States of America" name="California" pageId="3" pageNumber="34">California</collectingRegion>
congeners by the character combination of the absence of eyes, humeral angle of elytron bluntly angulate (Fig. 1A), and the absence of a basal elytral sutural ridge.
<taxonomicName authorityName="Brendel B" authorityYear="1889" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis italics="true" pageId="3" pageNumber="34">Cephennium anophthalmicum</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName authorityName="Hopp &amp; Caterino H" authorityYear="2009" box="[749,893,1399,1425]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="urbanum">
<emphasis box="[749,893,1399,1425]" italics="true" pageId="3" pageNumber="34">C. urbanum</emphasis>
</taxonomicName>
, but can be separated from it by the presence of a basal elytral sutural ridge (Fig. 1G), and the apex of the mesosternal keel divergent and crescent-shaped in
<taxonomicName authorityName="Hopp &amp; Caterino H" authorityYear="2009" box="[899,1043,1469,1496]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="urbanum">
<emphasis box="[899,1043,1469,1496]" italics="true" pageId="3" pageNumber="34">C. urbanum</emphasis>
</taxonomicName>
(
<figureCitation captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="3" pageNumber="34">Fig. 2G</figureCitation>
). It can be easily distinguished from
<taxonomicName box="[618,738,1504,1531]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="aridum">
<emphasis box="[618,738,1504,1531]" italics="true" pageId="3" pageNumber="34">C. aridum</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Hopp &amp; Caterino C" authorityYear="2009" box="[754,891,1504,1531]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="celsifrons">
<emphasis box="[754,891,1504,1531]" italics="true" pageId="3" pageNumber="34">C. celsifrons</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[958,1107,1505,1531]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="mariposae">
<emphasis box="[958,1107,1505,1531]" italics="true" pageId="3" pageNumber="34">C. mariposae</emphasis>
</taxonomicName>
by the presence of eyes in these species (
<figureCitation box="[613,705,1539,1566]" captionStart="Figure 4" captionStartId="8.[140,212,1040,1062]" captionTargetBox="[140,1108,165,1021]" captionTargetId="figure@8.[140,1108,165,1021]" captionTargetPageId="8" captionText="Figure 4. A–D Characters on the head, including number of ommatidia and male frons A–B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="3" pageNumber="34">Fig. 4A</figureCitation>
, C-D), and can be separated from
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[140,350,1575,1601]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[140,350,1575,1601]" italics="true" pageId="3" pageNumber="34">C. grandarboreum</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Hopp &amp; Caterino F" authorityYear="2009" box="[365,500,1575,1601]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[365,500,1575,1601]" italics="true" pageId="3" pageNumber="34">C. canestroi</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Hopp &amp; Caterino G" authorityYear="2009" box="[558,676,1575,1601]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis box="[558,676,1575,1601]" italics="true" pageId="3" pageNumber="34">C. gilberti</emphasis>
</taxonomicName>
by the humeral angle of the elytron, which is raised, dorsally flattened and apically rounded in these three species (
<figureCitation captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="3" pageNumber="34">Fig. 3</figureCitation>
D-F).
</paragraph>
<paragraph blockId="4.[140,1108,1558,1676]" pageId="4" pageNumber="35">
<emphasis bold="true" box="[140,299,1558,1580]" pageId="4" pageNumber="35">Figure |. A–H</emphasis>
Dorsal habitus SEMs, all to same scale
<emphasis bold="true" box="[673,692,1558,1580]" pageId="4" pageNumber="35">A</emphasis>
<taxonomicName authority="Brendel B" authorityName="Brendel B" authorityYear="1889" box="[698,1070,1558,1580]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis box="[698,968,1558,1579]" italics="true" pageId="4" pageNumber="35">Cephennium anophthalmicum</emphasis>
Brendel
<emphasis bold="true" box="[1053,1070,1558,1580]" pageId="4" pageNumber="35">B</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino C" authorityName="Hopp &amp; Caterino C" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="celsifrons">
<emphasis italics="true" pageId="4" pageNumber="35">Cephennium celsifrons</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[486,505,1590,1612]" pageId="4" pageNumber="35">C</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino D" authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[509,908,1590,1612]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="mariposae">
<emphasis box="[509,714,1590,1612]" italics="true" pageId="4" pageNumber="35">Cephennium mariposae</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[889,908,1590,1612]" pageId="4" pageNumber="35">D</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino E" authorityName="Hopp &amp; Caterino E" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis italics="true" pageId="4" pageNumber="35">Cephennium grandarboreum</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[396,411,1622,1644]" pageId="4" pageNumber="35">E</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino F" authorityName="Hopp &amp; Caterino F" authorityYear="2009" box="[420,822,1622,1644]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[420,617,1622,1644]" italics="true" pageId="4" pageNumber="35">Cephennium canestroi</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[808,822,1622,1644]" pageId="4" pageNumber="35">F</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino G" authorityName="Hopp &amp; Caterino G" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis box="[831,1015,1622,1643]" italics="true" pageId="4" pageNumber="35">Cephennium gilberti</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[226,246,1654,1676]" pageId="4" pageNumber="35">G</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino H" authorityName="Hopp &amp; Caterino H" authorityYear="2009" box="[251,650,1654,1676]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="urbanum">
<emphasis box="[251,452,1654,1675]" italics="true" pageId="4" pageNumber="35">Cephennium urbanum</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[630,650,1654,1676]" pageId="4" pageNumber="35">H</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino." authorityName="Hopp &amp; Caterino." box="[655,1018,1654,1676]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="aridum">
<emphasis box="[655,839,1654,1675]" italics="true" pageId="4" pageNumber="35">Cephennium aridum</emphasis>
Hopp &amp; Caterino.
</taxonomicName>
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770027" ID-Zenodo-Dep="3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="5" pageNumber="36" startId="5.[140,212,1290,1312]" targetBox="[140,1108,165,1270]" targetPageId="5">
<paragraph blockId="5.[140,1108,1247,1408]" pageId="5" pageNumber="36">
<emphasis bold="true" box="[140,306,1290,1312]" pageId="5" pageNumber="36">Figure 2. A–H</emphasis>
Mesosternal keel SEMs
<emphasis bold="true" box="[542,561,1290,1312]" pageId="5" pageNumber="36">A</emphasis>
<taxonomicName authority="Brendel B" authorityName="Brendel B" authorityYear="1889" box="[569,947,1290,1312]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis box="[569,842,1290,1311]" italics="true" pageId="5" pageNumber="36">Cephennium anophthalmicum</emphasis>
Brendel
<emphasis bold="true" box="[930,947,1290,1312]" pageId="5" pageNumber="36">B</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino C" authorityName="Hopp &amp; Caterino C" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="celsifrons">
<emphasis italics="true" pageId="5" pageNumber="36">Cephennium celsifrons</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[388,407,1322,1344]" pageId="5" pageNumber="36">C</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino D" authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[415,839,1322,1344]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="mariposae">
<emphasis box="[415,628,1322,1344]" italics="true" pageId="5" pageNumber="36">Cephennium mariposae</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[820,839,1322,1344]" pageId="5" pageNumber="36">D</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino E" authorityName="Hopp &amp; Caterino E" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[848,1108,1322,1343]" italics="true" pageId="5" pageNumber="36">Cephennium grandarboreum</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[312,327,1354,1376]" pageId="5" pageNumber="36">E</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino F" authorityName="Hopp &amp; Caterino F" authorityYear="2009" box="[333,721,1354,1376]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[333,530,1354,1376]" italics="true" pageId="5" pageNumber="36">Cephennium canestroi</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[707,721,1354,1376]" pageId="5" pageNumber="36">F</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino G" authorityName="Hopp &amp; Caterino G" authorityYear="2009" box="[727,1108,1354,1376]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis box="[727,910,1354,1375]" italics="true" pageId="5" pageNumber="36">Cephennium gilberti</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[1088,1108,1354,1376]" pageId="5" pageNumber="36">G</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino H" authorityName="Hopp &amp; Caterino H" authorityYear="2009" box="[140,544,1386,1408]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="urbanum">
<emphasis box="[140,343,1386,1407]" italics="true" pageId="5" pageNumber="36">Cephennium urbanum</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[524,544,1386,1408]" pageId="5" pageNumber="36">H</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino." box="[549,917,1386,1408]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="aridum">
<emphasis box="[549,736,1386,1407]" italics="true" pageId="5" pageNumber="36">Cephennium aridum</emphasis>
Hopp &amp; Caterino.
</taxonomicName>
</paragraph>
</caption>
<subSubSection lastPageId="7" lastPageNumber="38" pageId="5" pageNumber="36" type="description">
<paragraph blockId="5.[140,1108,1469,1672]" lastBlockId="7.[140,1108,167,686]" lastPageId="7" lastPageNumber="38" pageId="5" pageNumber="36">
<emphasis bold="true" box="[185,439,1469,1495]" pageId="5" pageNumber="36">
<emphasis box="[185,354,1469,1495]" inLineHeading="true" pageId="5" pageNumber="36" reason="1">Redescription</emphasis>
. Male.
</emphasis>
Length:
<quantity box="[552,680,1469,1496]" metricMagnitude="-4" metricUnit="m" metricValue="8.74" pageId="5" pageNumber="36" unit="mm" value="0.874">0.874 mm</quantity>
; pronotal width:
<quantity box="[893,1021,1469,1496]" metricMagnitude="-4" metricUnit="m" metricValue="4.18" pageId="5" pageNumber="36" unit="mm" value="0.418">0.418 mm</quantity>
; elytral width:
<quantity box="[220,342,1504,1531]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" pageId="5" pageNumber="36" unit="mm" value="0.475">0.475 mm</quantity>
. Body elongate, slender, weakly convex; testaceous; evenly and moderately pubescent; pubescence golden, slender, moderately long, weakly decumbent (Fig. 1A). Dorsal surface of head smooth, weakly pubescent, narrowing anteriorly from antennal insertions. Eyes absent. Antenna setose, antennomere I and II longer than broad, antennomeres III-VI quadrate and smaller than antennomeres II and VII, an- tennomere VIII smaller than antennomeres VII and IX, antennomeres IX-XI gradually clavate forming a loose club. Pronotum moderately pubescent, broadest between middle and anterior third, very convex in disc and moderately flattened near each posterior angle; anterior margin not visible from above; anterior and posterior margin lacking marginal bead; lateral marginal bead complete, gradually widening towards base; lateral edge broadly rounded to posterior third, then weakly sinuate to base (Fig. 1A). Hypomeron smooth, sparsely setose towards anterior quarter and along outside (lateral) edge. Prosternum lacking protuberant nodules anterolaterad procoxal cavities (
<figureCitation box="[148,236,1504,1530]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="6" pageNumber="37">Fig. 2A</figureCitation>
). Elytra smooth, as pubescent as pronotum, covering all abdominal segments; elytral suture flat; basomedial fovea present on each elytron; fovea moderate in size, moderately pubescent (Figs. 1A, 3A). Humeral angle of elytron projecting laterally to blunt point, dorsally raised and flattened (
<figureCitation box="[666,757,1610,1636]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="6" pageNumber="37">Fig. 3A</figureCitation>
). Scutellum weakly triangular, lacking setae (
<figureCitation box="[297,386,1645,1671]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="6" pageNumber="37">Fig. 3A</figureCitation>
). Mesosternal keel sparsely setose, lacking scale-like microsculp- ture, posterior quarter impunctate, apex weakly bifid (divergent), divergent projections short, triangulate (
<figureCitation box="[408,496,202,228]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="7" pageNumber="38">Fig. 2A</figureCitation>
). Metathoracic wings vestigial. Femora strongly clavate in distal half, tibiae expanded and becoming more densely setose towards distal half. Five visible abdominal sternites, ventrites V and VI partially fused. Aedeagus strongly sclerotized, with median lobe basally rounded, pill-shaped; parameres thin, sinuate, bisetose apically, extending to apex of rather narrow, bluntly triangular median dorsal process; apical digiform process curving ventrad at apex, extending just beyond apical collar; membranous apical collar with sclerotized clasper-like processes extending from apex; membranous lateral flaps present at base of apical collar (
<figureCitation box="[848,936,448,475]" captionStart="Figure 5" captionStartId="9.[140,212,1408,1430]" captionTargetBox="[156,1106,208,1373]" captionTargetId="figure@9.[140,1108,165,1389]" captionTargetPageId="9" captionText="Figure 5. A–F Dorsal and lateral view of the aedeagus A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium grandarboreum Hopp &amp; Caterino D Cephennium canestroi Hopp &amp; Caterino E Cephennium urbanum Hopp &amp; Caterino F Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770035" httpUri="https://zenodo.org/record/3770035/files/figure.png" pageId="7" pageNumber="38">Fig. 5A</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770029" ID-Zenodo-Dep="3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="6" pageNumber="37" startId="6.[140,212,1057,1079]" targetBox="[155,1108,208,1037]" targetPageId="6">
<paragraph blockId="6.[140,1108,1057,1175]" pageId="6" pageNumber="37">
<emphasis bold="true" box="[140,296,1057,1079]" pageId="6" pageNumber="37">Figure 3. A–H</emphasis>
Base of left elytron
<emphasis bold="true" box="[493,512,1057,1079]" pageId="6" pageNumber="37">A</emphasis>
<taxonomicName authority="Brendel B" authorityName="Brendel B" authorityYear="1889" box="[520,899,1057,1079]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis box="[520,794,1057,1078]" italics="true" pageId="6" pageNumber="37">Cephennium anophthalmicum</emphasis>
Brendel
<emphasis bold="true" box="[882,899,1057,1079]" pageId="6" pageNumber="37">B</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino C" authorityName="Hopp &amp; Caterino C" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="celsifrons">
<emphasis box="[907,1108,1057,1078]" italics="true" pageId="6" pageNumber="37">Cephennium celsifrons</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[317,336,1089,1111]" pageId="6" pageNumber="37">C</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino D" authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[342,755,1089,1111]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="mariposae">
<emphasis box="[342,552,1089,1111]" italics="true" pageId="6" pageNumber="37">Cephennium mariposae</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[736,755,1089,1111]" pageId="6" pageNumber="37">D</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino E" authorityName="Hopp &amp; Caterino E" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[762,1019,1089,1110]" italics="true" pageId="6" pageNumber="37">Cephennium grandarboreum</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[227,242,1121,1143]" pageId="6" pageNumber="37">E</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino F" authorityName="Hopp &amp; Caterino F" authorityYear="2009" box="[248,639,1121,1143]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[248,446,1121,1143]" italics="true" pageId="6" pageNumber="37">Cephennium canestroi</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[625,639,1121,1143]" pageId="6" pageNumber="37">F</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino G" authorityName="Hopp &amp; Caterino G" authorityYear="2009" box="[645,1028,1121,1143]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis box="[645,828,1121,1142]" italics="true" pageId="6" pageNumber="37">Cephennium gilberti</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[1008,1028,1121,1143]" pageId="6" pageNumber="37">G</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino H" authorityName="Hopp &amp; Caterino H" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="urbanum">
<emphasis italics="true" pageId="6" pageNumber="37">Cephennium urbanum</emphasis>
Hopp &amp; Caterino
<emphasis bold="true" box="[458,478,1153,1175]" pageId="6" pageNumber="37">H</emphasis>
</taxonomicName>
<taxonomicName authority="Hopp &amp; Caterino." authorityName="Hopp &amp; Caterino." box="[483,851,1153,1175]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="aridum">
<emphasis box="[483,670,1153,1174]" italics="true" pageId="6" pageNumber="37">Cephennium aridum</emphasis>
Hopp &amp; Caterino.
</taxonomicName>
</paragraph>
</caption>
<paragraph blockId="7.[140,1108,167,686]" box="[185,483,483,510]" pageId="7" pageNumber="38">
<emphasis bold="true" box="[185,277,483,509]" pageId="7" pageNumber="38">
<emphasis box="[185,272,483,509]" inLineHeading="true" pageId="7" pageNumber="38" reason="1">Female</emphasis>
.
</emphasis>
Identical to male.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="38" type="biology_ecology">
<paragraph blockId="7.[140,1108,167,686]" pageId="7" pageNumber="38">
<emphasis bold="true" box="[185,283,518,544]" pageId="7" pageNumber="38">
<emphasis box="[185,278,518,544]" inLineHeading="true" pageId="7" pageNumber="38" reason="1">Biology</emphasis>
.
</emphasis>
This species was first described from a single specimen that was sifted from vegetable debris. An additional specimen was sifted from forest duff. Beyond this, there is little known about the biology of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="38" type="distribution">
<paragraph blockId="7.[140,1108,167,686]" pageId="7" pageNumber="38">
<emphasis bold="true" box="[185,344,624,650]" pageId="7" pageNumber="38">
<emphasis box="[185,339,624,650]" inLineHeading="true" pageId="7" pageNumber="38" reason="1">Distribution</emphasis>
.
</emphasis>
This species has been collected around the San Francisco Bay Area in central coastal
<collectingRegion box="[337,451,659,685]" country="United States of America" name="California" pageId="7" pageNumber="38">California</collectingRegion>
(Fig. 6).
</paragraph>
</subSubSection>
</treatment>
</document>
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