treatments-xml/data/41/57/F9/4157F97AFF97FFF7FF7EFC3EFECA16FF.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.3790610" ID-GBIF-Dataset="ab7d8c5f-f732-4f73-9e1f-5fe2bae6c9a7" ID-GBIF-Taxon="163754309" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790610" ID-ZooBank="BB883B1E-E58F-4074-92EB-7E814E78F678" LSID="urn:lsid:zoobank.org:act:2E36C8F3-E54C-4CD3-AAF4-A5D366014F5E" approvalRequired="19" approvalRequired_for_textStreams="19" checkinTime="1587996592918" checkinUser="plazi" docAuthor="Hopp, Katie &amp; Caterino, Michael" docDate="2009" docId="4157F97AFF97FFF7FF7EFC3EFECA16FF" docLanguage="en" docName="ZK_article_2083.pdf" docOrigin="ZooKeys 24 (24)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5c452e69" docStyleName="zookeys.2008.journal_article" docTitle="Cephennium mariposae Hopp &amp; Caterino D 2009, sp. n." docType="treatment" docUuid="2E36C8F3-E54C-4CD3-AAF4-A5D366014F5E" docUuidSource="ZooBank" docVersion="6" lastPageId="13" lastPageNumber="44" masterDocId="BD6E8102FF9DFFFAFFF2FF92FFBD1451" masterDocTitle="Seven new species of Cephennium Müller &amp; Kunze (Coleoptera, Staphylinidae, Scydmaeninae, Cephenniini) from California with a key to native North American species" masterLastPageNumber="54" masterPageNumber="31" pageId="10" pageNumber="41" updateTime="1643563761986" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title>Seven new species of Cephennium Müller &amp; Kunze (Coleoptera, Staphylinidae, Scydmaeninae, Cephenniini) from California with a key to native North American species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hopp, Katie</mods:namePart>
<mods:affiliation>,,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael</mods:namePart>
<mods:affiliation>Clemson University, Clemson, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-10-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>24</mods:number>
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<mods:detail type="issue">
<mods:number>24</mods:number>
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<mods:identifier type="DOI">10.3897/zookeys.24.247</mods:identifier>
<mods:identifier type="GBIF-Dataset">ab7d8c5f-f732-4f73-9e1f-5fe2bae6c9a7</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576549</mods:identifier>
<mods:identifier type="ZooBank">BB883B1E-E58F-4074-92EB-7E814E78F678</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790610" ID-GBIF-Taxon="163754309" ID-Zenodo-Dep="3790610" LSID="urn:lsid:zoobank.org:act:2E36C8F3-E54C-4CD3-AAF4-A5D366014F5E" httpUri="http://treatment.plazi.org/id/4157F97AFF97FFF7FF7EFC3EFECA16FF" lastPageId="13" lastPageNumber="44" pageId="10" pageNumber="41">
<subSubSection pageId="10" pageNumber="41" type="nomenclature">
<paragraph blockId="10.[140,983,940,1038]" box="[140,722,940,967]" pageId="10" pageNumber="41">
<heading box="[140,722,940,967]" fontSize="11" level="2" pageId="10" pageNumber="41" reason="3">
<emphasis bold="true" box="[140,722,940,967]" pageId="10" pageNumber="41">
<taxonomicName authority="Hopp &amp; Caterino" authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[140,644,940,967]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="mariposae" status="sp. n.">
<emphasis bold="true" box="[140,422,940,967]" italics="true" pageId="10" pageNumber="41">
<emphasis box="[140,292,940,967]" inLineHeading="true" pageId="10" pageNumber="41" reason="1">Cephennium</emphasis>
mariposae
</emphasis>
Hopp &amp; Caterino
</taxonomicName>
, 
<taxonomicNameLabel box="[657,722,941,967]" pageId="10" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act: 
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</paragraph>
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<subSubSection box="[140,406,1011,1038]" pageId="10" pageNumber="41" type="description">
<paragraph blockId="10.[140,983,940,1038]" box="[140,406,1011,1038]" pageId="10" pageNumber="41">Figs 1C, 2C, 3C, 4C, 6</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="materials_examined">
<paragraph blockId="10.[140,1108,1081,1636]" pageId="10" pageNumber="41">
<emphasis bold="true" box="[140,318,1081,1108]" pageId="10" pageNumber="41">
<emphasis box="[140,198,1082,1108]" inLineHeading="true" pageId="10" pageNumber="41" reason="1">Type</emphasis>
Material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608963385" collectingDate="1984-11-15" collectionCode="CSCA" collectorName="A. J. Gilbert" county="Mariposa Co." latitude="37.4766" location="Mariposa" longLatPrecision="6" longitude="-119.9584" pageId="10" pageNumber="41" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[328,441,1082,1108]" pageId="10" pageNumber="41">
<typeStatus box="[328,441,1082,1108]" pageId="10" pageNumber="41">Holotype</typeStatus>
</emphasis>
(sex not determined): “CALIF: 
<collectingCounty box="[824,985,1082,1108]" pageId="10" pageNumber="41">Mariposa Co.</collectingCounty>
; 
<location LSID="urn:lsid:plazi:treatment:4157F97AFF97FFF7FF7EFC3EFECA16FF:CC211EB7FF97FFF0FC1BFBA8FBEE1005" box="[1001,1107,1082,1108]" county="Mariposa Co." latitude="37.4766" longLatPrecision="6" longitude="-119.9584" name="Mariposa" pageId="10" pageNumber="41">Mariposa</location>
[̴ 
<geoCoordinate box="[157,294,1117,1143]" degrees="37.4766" direction="north" orientation="latitude" pageId="10" pageNumber="41" precision="5" value="37.4766">37.4766°N</geoCoordinate>
, 
<geoCoordinate box="[306,456,1116,1143]" degrees="119.9584" direction="west" orientation="longitude" pageId="10" pageNumber="41" precision="5" value="-119.9584">119.9584°W</geoCoordinate>
] 
<date box="[470,609,1117,1144]" pageId="10" pageNumber="41" value="1984-11-15">
<collectingDate box="[470,609,1117,1144]" pageId="10" pageNumber="41" value="1984-11-15">XI-15-1984</collectingDate>
</date>
; Berlese rotten log; 
<collectorName box="[832,973,1117,1144]" pageId="10" pageNumber="41">A. J. Gilbert</collectorName>
” (
<collectionCode box="[998,1078,1117,1143]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="10" pageNumber="41" type="Museum">CSCA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="etymology">
<paragraph blockId="10.[140,1108,1081,1636]" pageId="10" pageNumber="41">
<emphasis bold="true" box="[185,320,1152,1178]" pageId="10" pageNumber="41">
<emphasis box="[185,315,1152,1178]" inLineHeading="true" pageId="10" pageNumber="41" reason="1">Etymology</emphasis>
.
</emphasis>
This species is named for the county and town from which it was collected, Mariposa.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="diagnosis">
<paragraph blockId="10.[140,1108,1081,1636]" pageId="10" pageNumber="41">
<emphasis bold="true" box="[185,313,1222,1248]" pageId="10" pageNumber="41">
<emphasis box="[185,308,1222,1248]" inLineHeading="true" pageId="10" pageNumber="41" reason="1">Diagnosis</emphasis>
.
</emphasis>
This species can be distinguished from its 
<collectingRegion box="[794,910,1223,1249]" country="United States of America" name="California" pageId="10" pageNumber="41">California</collectingRegion>
congeners by the character combination of the presence of four ommatidia on each side of the head (
<figureCitation box="[148,241,1293,1319]" captionStart="Figure 4" captionStartId="8.[140,212,1040,1062]" captionTargetBox="[140,1108,165,1021]" captionTargetId="figure@8.[140,1108,165,1021]" captionTargetPageId="8" captionText="Figure 4. A–D Characters on the head, including number of ommatidia and male frons A–B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770031" httpUri="https://zenodo.org/record/3770031/files/figure.png" pageId="10" pageNumber="41">Fig. 4C</figureCitation>
), the mesosternal process truncate at the apex (
<figureCitation box="[797,890,1293,1319]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="10" pageNumber="41">Fig. 2C</figureCitation>
), and the humeral angle of the elytron weakly raised and dorsally flattened (
<figureCitation box="[786,875,1328,1354]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="10" pageNumber="41">Fig. 3C</figureCitation>
). 
<taxonomicName authorityName="Hopp &amp; Caterino D" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="mariposae">
<emphasis italics="true" pageId="10" pageNumber="41">Cephennium mariposae</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName authorityName="Hopp &amp; Caterino C" authorityYear="2009" box="[405,535,1363,1390]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="celsifrons">
<emphasis box="[405,535,1363,1390]" italics="true" pageId="10" pageNumber="41">C. celsifrons</emphasis>
</taxonomicName>
but can be distinguished from it by the latter having only two ommatidia on each side of the head (
<figureCitation box="[720,808,1399,1425]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="10" pageNumber="41">Fig. 2A</figureCitation>
) and a more elongate and slender body form (Fig. 1B). Th is species can be easily separated from 
<taxonomicName authorityName="Hopp &amp; Caterino F" authorityYear="2009" box="[961,1100,1434,1460]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="urbanum">
<emphasis box="[961,1100,1434,1460]" italics="true" pageId="10" pageNumber="41">C. urbanum</emphasis>
</taxonomicName>
, 
<taxonomicName box="[140,349,1469,1496]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="anopthalmicum">
<emphasis box="[140,349,1469,1496]" italics="true" pageId="10" pageNumber="41">C. anopthalmicum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[366,497,1470,1496]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[366,497,1470,1496]" italics="true" pageId="10" pageNumber="41">C. canestroi</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Hopp &amp; Caterino D" authorityYear="2009" box="[514,718,1469,1496]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[514,718,1469,1496]" italics="true" pageId="10" pageNumber="41">C. grandarboreum</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Hopp &amp; Caterino G" authorityYear="2009" box="[780,894,1469,1496]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis box="[780,894,1469,1496]" italics="true" pageId="10" pageNumber="41">C. gilberti</emphasis>
</taxonomicName>
by the absence of ommatidia and the apex of the mesosternal keel being divergent in these species. It can be further distinguished from 
<taxonomicName authorityName="Brendel B" authorityYear="1889" box="[529,750,1539,1566]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="anophthalmicum">
<emphasis box="[529,750,1539,1566]" italics="true" pageId="10" pageNumber="41">C. anophthalmicum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Hopp &amp; Caterino F" authorityYear="2009" box="[764,901,1539,1566]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="urbanum">
<emphasis box="[764,901,1539,1566]" italics="true" pageId="10" pageNumber="41">C. urbanum</emphasis>
</taxonomicName>
and 
<taxonomicName box="[957,1073,1539,1566]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="aridum">
<emphasis box="[957,1073,1539,1566]" italics="true" pageId="10" pageNumber="41">C. aridum</emphasis>
</taxonomicName>
by the humeral angle of the elytron projecting laterally to a point in these three species (
<figureCitation box="[148,236,1610,1636]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="10" pageNumber="41">Fig. 3A</figureCitation>
, G-H).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="42" type="description">
<paragraph blockId="11.[140,1108,166,1179]" pageId="11" pageNumber="42">
<emphasis bold="true" box="[185,339,166,192]" pageId="11" pageNumber="42">
<emphasis box="[185,333,166,192]" inLineHeading="true" pageId="11" pageNumber="42" reason="1">Description</emphasis>
.
</emphasis>
Length: 
<quantity box="[454,584,167,193]" metricMagnitude="-4" metricUnit="m" metricValue="8.36" pageId="11" pageNumber="42" unit="mm" value="0.836">0.836 mm</quantity>
; pronotal width: 
<quantity box="[801,931,167,193]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="42" unit="mm" value="0.38">0.380 mm</quantity>
; elytral width: 
<quantity box="[140,265,202,228]" metricMagnitude="-4" metricUnit="m" metricValue="4.37" pageId="11" pageNumber="42" unit="mm" value="0.437">0.437 mm</quantity>
. Body broad, ovate, slightly convex, testaceous, evenly densely pubescent, pubescence golden, slender, moderate in length (Fig. 1C). Head small, deflexed, sparsely pubescent, not narrowing anteriorly from antennal insertions; frons flat; four ommatidia present on each side of head. Antenna setose, antennomere I and II longer than broad, antennomeres III-VI quadrate and smaller than antennomeres II and VII, antennomere VIII smaller than antennomeres VII and IX, antennomeres IX-XI gradually clavate forming a loose club. Pronotum densely pubescent, broadest between middle and anterior third, disc convex medially and weakly flattened near each posterior angle; anterior margin not visible from above; anterior and posterior margin lacking marginal bead; marginal bead complete laterally, gradually widening towards base; lateral edge broadly rounded to posterior third, then evenly to base (Fig. 1C). Hypomeron smooth, sparsely setose towards upper quarter and along outside (lateral) edge; hypomeral bead anterolaterad procoxae sinuate. Prosternum without nodules anterolaterad procoxal cavities (
<figureCitation box="[600,691,659,686]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="11" pageNumber="42">Fig. 2C</figureCitation>
). Elytra impunctate, as pubescent as pronotum, covering all abdominal segments, weakly truncate at apex; elytral suture flat; elytral striae absent; basomedial fovea present on each elytron, fovea small, with moderately dense inwardly directed setae (Figs. 1C, 3C). Humeral angles of elytron raised, dorsally flattened (plateau-like), apically rounded, slender, moderately curving posterad around anterolateral angles (
<figureCitation box="[637,732,835,862]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="11" pageNumber="42">Fig. 3C</figureCitation>
). Scutellum roundly triangular, without setae (
<figureCitation box="[311,404,871,897]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="11" pageNumber="42">Fig. 3C</figureCitation>
). Mesosternal keel setose with indentation around each seta, apex not divergent (
<figureCitation box="[374,465,906,932]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="11" pageNumber="42">Fig. 2B</figureCitation>
). Metathoracic wings vestigial. Femora strongly clavate in distal half, tibiae expanded and becoming more densely setose towards distal half. Six visible abdominal sternites (fusion between ventrites V and VI). Aedeagus not studied.
</paragraph>
<paragraph blockId="11.[140,1108,166,1179]" box="[185,483,1047,1073]" pageId="11" pageNumber="42">
<emphasis bold="true" box="[185,277,1047,1073]" pageId="11" pageNumber="42">
<emphasis box="[185,272,1047,1073]" inLineHeading="true" pageId="11" pageNumber="42" reason="1">Female</emphasis>
.
</emphasis>
Identical to male.
</paragraph>
</subSubSection>
<subSubSection box="[185,1027,1081,1108]" pageId="11" pageNumber="42" type="biology_ecology">
<paragraph blockId="11.[140,1108,166,1179]" box="[185,1027,1081,1108]" pageId="11" pageNumber="42">
<emphasis bold="true" box="[185,283,1081,1107]" pageId="11" pageNumber="42">
<emphasis box="[185,278,1081,1107]" inLineHeading="true" pageId="11" pageNumber="42" reason="1">Biology</emphasis>
.
</emphasis>
Th e lone specimen of this species was collected from a rotten log.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="42" type="distribution">
<paragraph blockId="11.[140,1108,166,1179]" pageId="11" pageNumber="42">
<emphasis bold="true" box="[185,344,1117,1143]" pageId="11" pageNumber="42">
<emphasis box="[185,339,1117,1143]" inLineHeading="true" pageId="11" pageNumber="42" reason="1">Distribution</emphasis>
.
</emphasis>
This species is only known from Mariposa in Mariposa County, CA (Fig. 6).
</paragraph>
<paragraph blockId="11.[140,981,1257,1355]" pageId="11" pageNumber="42">
<emphasis bold="true" box="[140,786,1257,1284]" pageId="11" pageNumber="42">
<taxonomicName authority="Hopp &amp; Caterino" authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[140,708,1257,1284]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="grandarboreum" status="sp. n.">
<emphasis bold="true" box="[140,485,1257,1284]" italics="true" pageId="11" pageNumber="42">
<emphasis box="[140,292,1257,1284]" inLineHeading="true" pageId="11" pageNumber="42" reason="1">Cephennium</emphasis>
grandarboreum
</emphasis>
Hopp &amp; Caterino
</taxonomicName>
, 
<taxonomicNameLabel box="[721,786,1258,1284]" pageId="11" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
<uri box="[140,981,1293,1320]" pageId="11" pageNumber="42">
urn:lsid:zoobank.org:act: 
<uuid box="[420,981,1293,1320]" pageId="11" pageNumber="42">8CD00FEF-266D-42A1-8C2F-0031C5EAAF06</uuid>
</uri>
</paragraph>
<paragraph blockId="11.[140,981,1257,1355]" box="[140,413,1328,1355]" pageId="11" pageNumber="42">Figs 1D, 2D, 3D, 5C, 6</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42" type="materials_examined">
<paragraph blockId="11.[140,1108,1398,1672]" pageId="11" pageNumber="42">
<emphasis bold="true" box="[140,320,1398,1425]" pageId="11" pageNumber="42">
<emphasis box="[140,198,1399,1425]" inLineHeading="true" pageId="11" pageNumber="42" reason="1">Type</emphasis>
Material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608963382" collectionCode="SBMN" county="Monterey Co." latitude="36.2403" location="Monterey Co." longLatPrecision="6" longitude="-121.7781" pageId="11" pageNumber="42" specimenCode="CBP0041534" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[333,535,1399,1425]" pageId="11" pageNumber="42">
<typeStatus box="[333,449,1399,1425]" pageId="11" pageNumber="42">Holotype</typeStatus>
. Male.
</emphasis>
“CA: 
<collectingCounty box="[620,788,1399,1425]" pageId="11" pageNumber="42">Monterey Co.</collectingCounty>
; 
<geoCoordinate box="[807,937,1399,1425]" degrees="36.2403" direction="north" orientation="latitude" pageId="11" pageNumber="42" precision="5" value="36.2403">36.2403°N</geoCoordinate>
, 
<geoCoordinate box="[955,1103,1399,1425]" degrees="121.7781" direction="west" orientation="longitude" pageId="11" pageNumber="42" precision="5" value="-121.7781">121.7781°W</geoCoordinate>
; LPNF: Sycamore Cyn.; II.14.2006; Caterino&amp;Chatzimanolis; maple/redwood litter”/ “CA BEETLE PROJ; 
<specimenCode box="[392,551,1469,1496]" collectionCode="CBP" pageId="11" pageNumber="42">CBP0041534</specimenCode>
” (
<collectionCode box="[576,664,1469,1495]" pageId="11" pageNumber="42">SBMN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="11.[140,1108,1398,1672]" pageId="11" pageNumber="42">
<materialsCitation ID-GBIF-Occurrence="2608963383" collectionCode="SBMN, FMNH, CASC" county="Monterey Co." latitude="36.2403" location="Monterey Co." longLatPrecision="6" longitude="-121.7781" pageId="11" pageNumber="42" specimenCode="CBP0041525, CBP0041544, CBP0041516" specimenCount="3" typeStatus="paratype">
<emphasis bold="true" box="[185,300,1504,1530]" inLineHeading="true" pageId="11" pageNumber="42" reason="1">
<typeStatus box="[185,300,1504,1530]" pageId="11" pageNumber="42">Paratypes</typeStatus>
</emphasis>
(7): 
<specimenCount box="[358,496,1504,1530]" pageId="11" pageNumber="42" type="generic">3 specimens</specimenCount>
with same data as holotype (1 
<collectionCode box="[853,938,1504,1530]" pageId="11" pageNumber="42">SBMN</collectionCode>
, 
<specimenCode box="[950,1108,1504,1531]" collectionCode="CBP" pageId="11" pageNumber="42">CBP0041525</specimenCode>
[disarticulated]; 1 
<collectionCode box="[352,444,1540,1566]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="42" type="Museum">FMNH</collectionCode>
, 
<specimenCode box="[458,618,1539,1566]" collectionCode="CBP" pageId="11" pageNumber="42">CBP0041544</specimenCode>
; 1 
<collectionCode box="[656,733,1540,1566]" pageId="11" pageNumber="42">CASC</collectionCode>
, 
<specimenCode box="[746,908,1539,1566]" collectionCode="CBP" pageId="11" pageNumber="42">CBP0041516</specimenCode>
)
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608963396" collectionCode="CSCA, LACM, SBMN" collectorName="M. Caterino" county="Monterey Co." latitude="36.0772" location="Monterey Co." longLatPrecision="6" longitude="-121.5923" pageId="11" pageNumber="42" specimenCode="CBP0018437, CBP0018450" specimenCount="1" typeStatus="paratype">
“CA: 
<collectingCounty pageId="11" pageNumber="42">Monterey Co.</collectingCounty>
; 
<geoCoordinate box="[197,327,1575,1601]" degrees="36.0772" direction="north" orientation="latitude" pageId="11" pageNumber="42" precision="5" value="36.0772">36.0772°N</geoCoordinate>
, 
<geoCoordinate box="[340,488,1574,1601]" degrees="121.5923" direction="west" orientation="longitude" pageId="11" pageNumber="42" precision="5" value="-121.5923">121.5923°W</geoCoordinate>
; UC Big Creek Reserve; Redwood Camp, iii.28.2004. 
<collectorName box="[140,288,1610,1636]" pageId="11" pageNumber="42">M. Caterino</collectorName>
, redwood litter” (1 
<collectionCode box="[525,602,1610,1636]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="11" pageNumber="42" type="Museum">CSCA</collectionCode>
, 
<specimenCode box="[616,777,1610,1636]" collectionCode="CBP" pageId="11" pageNumber="42">CBP0018437</specimenCode>
; 1 
<collectionCode box="[817,902,1610,1636]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="11" pageNumber="42" type="Museum">LACM</collectionCode>
, 
<specimenCode box="[917,1077,1610,1637]" collectionCode="CBP" pageId="11" pageNumber="42">CBP0018450</specimenCode>
; 1 
<collectionCode box="[140,222,1645,1671]" pageId="11" pageNumber="42">SBMN</collectionCode>
[in freezer for DNA])
</materialsCitation>
; “CA: Monterey Co.; 
<geoCoordinate box="[720,851,1645,1671]" degrees="36.0812" direction="north" orientation="latitude" pageId="11" pageNumber="42" precision="5" value="36.0812">36.0812°N</geoCoordinate>
, 
<geoCoordinate box="[861,1010,1645,1672]" degrees="121.5974" direction="west" orientation="longitude" pageId="11" pageNumber="42" precision="5" value="-121.5974">121.5974°W</geoCoordinate>
; UC Big
</paragraph>
<paragraph blockId="12.[140,1108,167,1672]" pageId="12" pageNumber="43">
Creek Reserve; BigCk/BrunetteCk.confl.; ii.7.2003, M. Caterino; redwood litter” (1 
<collectionCode box="[140,225,202,228]" pageId="12" pageNumber="43">SBMN</collectionCode>
, 
<collectionCode box="[236,290,202,228]" pageId="12" pageNumber="43">CBP</collectionCode>
0006070 [gold coated for SEM]).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="43" type="etymology">
<paragraph blockId="12.[140,1108,167,1672]" pageId="12" pageNumber="43">
<emphasis bold="true" box="[185,320,237,263]" pageId="12" pageNumber="43">
<emphasis box="[185,315,237,263]" inLineHeading="true" pageId="12" pageNumber="43" reason="1">Etymology</emphasis>
.
</emphasis>
This species name is the combination of the Latin words 
<emphasis box="[979,1057,237,263]" italics="true" pageId="12" pageNumber="43">grandis</emphasis>
and 
<emphasis box="[140,230,272,298]" italics="true" pageId="12" pageNumber="43">arboreus</emphasis>
, meaning very large tree, as this species is associated with coast redwoods (
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Sequoia" kingdom="Plantae" order="Pinales" pageId="12" pageNumber="43" phylum="Tracheophyta" rank="species" species="sempervirens">
<emphasis italics="true" pageId="12" pageNumber="43">Sequoia sempervirens</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="43" type="diagnosis">
<paragraph blockId="12.[140,1108,167,1672]" pageId="12" pageNumber="43">
<emphasis bold="true" box="[185,312,342,368]" pageId="12" pageNumber="43">
<emphasis box="[185,307,342,368]" inLineHeading="true" pageId="12" pageNumber="43" reason="1">Diagnosis</emphasis>
.
</emphasis>
This species is most similar to 
<taxonomicName authorityName="Hopp &amp; Caterino G" authorityYear="2009" box="[670,895,343,369]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis box="[670,895,343,369]" italics="true" pageId="12" pageNumber="43">Cephennium gilberti</emphasis>
</taxonomicName>
but can be distinguished from it by its body size and shape. 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[632,945,378,404]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[632,945,378,404]" italics="true" pageId="12" pageNumber="43">Cephennium grandarboreum</emphasis>
</taxonomicName>
is represented by the largest specimens (
<quantity box="[439,643,413,440]" metricMagnitude="-3" metricUnit="m" metricValue="1.0545" metricValueMax="1.121" metricValueMin="0.988" pageId="12" pageNumber="43" unit="mm" value="1.0545" valueMax="1.121" valueMin="0.988">
<geoCoordinate box="[439,507,413,439]" degrees="0.988" direction="north" orientation="latitude" pageId="12" pageNumber="43" precision="55" value="0.988">0.988</geoCoordinate>
<geoCoordinate box="[507,587,413,439]" degrees="1.121" direction="west" orientation="longitude" pageId="12" pageNumber="43" precision="55" value="-1.121">–1.121</geoCoordinate>
mm
</quantity>
long) of this genus currently known to occur in 
<collectingRegion box="[238,356,448,474]" country="United States of America" name="California" pageId="12" pageNumber="43">California</collectingRegion>
, and is very robust. 
<taxonomicName authorityName="Hopp &amp; Caterino G" authorityYear="2009" box="[578,688,448,475]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis box="[578,688,448,475]" italics="true" pageId="12" pageNumber="43">C. gilberti</emphasis>
</taxonomicName>
is known from the smallest specimen (
<quantity box="[148,267,483,510]" metricMagnitude="-4" metricUnit="m" metricValue="6.46" pageId="12" pageNumber="43" unit="mm" value="0.646">0.646 mm</quantity>
long) of this genus known to occur in 
<collectingRegion box="[699,813,483,509]" country="United States of America" name="California" pageId="12" pageNumber="43">California</collectingRegion>
and is much more slender and elongate. These two species can also be separated by the mesosternal keel, which is strongly divergent in 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[382,582,554,580]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[382,582,554,580]" italics="true" pageId="12" pageNumber="43">C. grandarboreum</emphasis>
</taxonomicName>
, which has the apex ̴2.4× as wide as the widest anterior point (
<figureCitation box="[348,440,589,615]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="12" pageNumber="43">Fig. 2D</figureCitation>
). The mesosternal keel of 
<taxonomicName authorityName="Hopp &amp; Caterino G" authorityYear="2009" box="[729,839,589,616]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="gilberti">
<emphasis box="[729,839,589,616]" italics="true" pageId="12" pageNumber="43">C. gilberti</emphasis>
</taxonomicName>
is weakly divergent and ̴2× as wide as the widest anterior point (
<figureCitation box="[607,693,624,650]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="12" pageNumber="43">Fig. 2F</figureCitation>
). 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[713,1027,624,650]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[713,1027,624,650]" italics="true" pageId="12" pageNumber="43">Cephennium grandarboreum</emphasis>
</taxonomicName>
can be separated from all other species by having two scutellar setae (
<figureCitation box="[843,924,659,686]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="12" pageNumber="43">Fig.3D</figureCitation>
) instead of four (
<figureCitation box="[148,233,695,721]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="12" pageNumber="43">Fig. 3E</figureCitation>
) or zero (
<figureCitation box="[344,411,695,721]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="12" pageNumber="43">Figs 3</figureCitation>
A-C, G-H). Apart from the scutellar setae, 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[907,1107,695,721]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[907,1107,695,721]" italics="true" pageId="12" pageNumber="43">C. grandarboreum</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[190,318,730,756]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[190,318,730,756]" italics="true" pageId="12" pageNumber="43">C. canestroi</emphasis>
</taxonomicName>
are quite similar, with rounded elytral humeral angles, but the elytral foveae are large and densely setose in 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[568,768,765,792]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[568,768,765,792]" italics="true" pageId="12" pageNumber="43">C. grandarboreum</emphasis>
</taxonomicName>
(Figs. 1D, 2D), whereas they are smaller and moderately setose in 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[561,690,801,827]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[561,690,801,827]" italics="true" pageId="12" pageNumber="43">C. canestroi</emphasis>
</taxonomicName>
(Fig. 1E, 2E). Th e humeral angle of the elytron is also more slender and posterorly curved in 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[791,992,835,862]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="grandarboreum">
<emphasis box="[791,992,835,862]" italics="true" pageId="12" pageNumber="43">C. grandarboreum</emphasis>
</taxonomicName>
(
<figureCitation box="[1008,1098,835,862]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="12" pageNumber="43">Fig. 3D</figureCitation>
) than in 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[228,355,871,897]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[228,355,871,897]" italics="true" pageId="12" pageNumber="43">C. canestroi</emphasis>
</taxonomicName>
(
<figureCitation box="[370,456,871,897]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="12" pageNumber="43">Fig. 3E</figureCitation>
), and has a more strongly divergent mesosternal keel (
<figureCitation captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="12" pageNumber="43">Fig. 2D</figureCitation>
) than 
<taxonomicName authorityName="Hopp &amp; Caterino E" authorityYear="2009" box="[253,380,906,932]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="canestroi">
<emphasis box="[253,380,906,932]" italics="true" pageId="12" pageNumber="43">C. canestroi</emphasis>
</taxonomicName>
(
<figureCitation box="[395,482,906,932]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="12" pageNumber="43">Fig. 2E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="44" pageId="12" pageNumber="43" type="description">
<paragraph blockId="12.[140,1108,167,1672]" lastBlockId="13.[140,1108,167,686]" lastPageId="13" lastPageNumber="44" pageId="12" pageNumber="43">
<emphasis bold="true" box="[185,407,941,967]" pageId="12" pageNumber="43">
<emphasis box="[185,328,941,967]" inLineHeading="true" pageId="12" pageNumber="43" reason="1">Description</emphasis>
. Male.
</emphasis>
Length: 
<quantity box="[508,709,941,968]" metricMagnitude="-3" metricUnit="m" metricValue="1.0545" metricValueMax="1.121" metricValueMin="0.988" pageId="12" pageNumber="43" unit="mm" value="1.0545" valueMax="1.121" valueMin="0.988">
<geoCoordinate box="[508,575,941,967]" degrees="0.988" direction="north" orientation="latitude" pageId="12" pageNumber="43" precision="55" value="0.988">0.988</geoCoordinate>
<geoCoordinate box="[575,653,941,967]" degrees="1.121" direction="west" orientation="longitude" pageId="12" pageNumber="43" precision="55" value="-1.121">–1.121</geoCoordinate>
mm
</quantity>
; pronotal width: 
<quantity box="[902,1103,941,968]" metricMagnitude="-4" metricUnit="m" metricValue="4.275" metricValueMax="4.37" metricValueMin="4.18" pageId="12" pageNumber="43" unit="mm" value="0.4275" valueMax="0.437" valueMin="0.418">
<geoCoordinate box="[902,967,941,967]" degrees="0.418" direction="north" orientation="latitude" pageId="12" pageNumber="43" precision="55" value="0.418">0.418</geoCoordinate>
<geoCoordinate box="[967,1047,941,967]" degrees="0.437" direction="west" orientation="longitude" pageId="12" pageNumber="43" precision="55" value="-0.437">–0.437</geoCoordinate>
mm
</quantity>
; elytral width: 
<quantity box="[296,499,976,1003]" metricMagnitude="-4" metricUnit="m" metricValue="4.655" metricValueMax="4.75" metricValueMin="4.56" pageId="12" pageNumber="43" unit="mm" value="0.4655" valueMax="0.475" valueMin="0.456">
<geoCoordinate box="[296,362,976,1003]" degrees="0.456" direction="north" orientation="latitude" pageId="12" pageNumber="43" precision="55" value="0.456">0.456</geoCoordinate>
<geoCoordinate box="[362,442,976,1003]" degrees="0.475" direction="west" orientation="longitude" pageId="12" pageNumber="43" precision="55" value="-0.475">–0.475</geoCoordinate>
mm
</quantity>
. Body broad, ovate, slightly convex, rufo-testaceous to amber yellow, evenly densely pubescent, pubescence golden, slender, long (Fig. 1D). Head small, deflexed moderately pubescent, weakly narrowing anteriorly from antennal insertions; eyes absent. Antenna setose, antennomere I and II longer than broad, antennomeres III-VI quadrate and smaller than antennomeres II and VII, antennomere VIII smaller than antennomeres VII and IX, antennomeres IX-XI gradually clavate forming a loose club. Pronotum densely pubescent, broadest between middle and anterior third, disc very convex medially and weakly flattened near each posterior angle; anterior margin not visible from above; anterior and posterior margin lacking marginal bead; marginal bead complete laterally, gradually widening towards base; lateral edge broadly rounded to posterior third, then weakly sinuate at base (Fig. 1D). Hypomeron smooth, sparsely setose towards upper quarter and along outside (lateral) edge, hypomeral bead anterolaterad procoxae with small knob at apex. Prosternum with small, weakly produced egg-shaped nodules anterolaterad procoxal cavities (
<figureCitation box="[788,879,1434,1460]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="12" pageNumber="43">Fig. 2D</figureCitation>
). Elytra impunctate, as pubescent as pronotum, covering all abdominal segments, weakly truncate at apex; elytral suture flat; elytral striae absent; basomedial fovea present on each elytron, fovea large, deep, with dense inwardly directed setae (Fig. 1D, 3D). Humeral angle of elytron raised laterad scutellum to humeral angle, humeral plateau slender, dorsally flattened, apically rounded; apex reaching posterad past midline of elytral fovea (
<figureCitation box="[919,1009,1610,1636]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="12" pageNumber="43">Fig. 3D</figureCitation>
). Scutellum roundly triangular, with two setae (
<figureCitation box="[594,686,1645,1671]" captionStart="Figure 3" captionStartId="6.[140,212,1057,1079]" captionTargetBox="[155,1108,208,1037]" captionTargetId="figure@6.[140,1108,165,1038]" captionTargetPageId="6" captionText="Figure 3. A–H Base of left elytron A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770029" httpUri="https://zenodo.org/record/3770029/files/figure.png" pageId="12" pageNumber="43">Fig. 3D</figureCitation>
). Mesosternal keel setose, texture ap- pearing as fish scales anteriorly, abruptly smooth and impunctate near mesometasternal suture, apex strongly bifid and divergent, posterior projections long (
<figureCitation box="[893,983,202,228]" captionStart="Figure 2" captionStartId="5.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@5.[140,1108,165,1271]" captionTargetPageId="5" captionText="Figure 2. A–H Mesosternal keel SEMs A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium mariposae Hopp &amp; Caterino D Cephennium grandarboreum Hopp &amp; Caterino E Cephennium canestroi Hopp &amp; Caterino F Cephennium gilberti Hopp &amp; Caterino G Cephennium urbanum Hopp &amp; Caterino H Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770027" httpUri="https://zenodo.org/record/3770027/files/figure.png" pageId="13" pageNumber="44">Fig. 2D</figureCitation>
). Metathoracic wings vestigial. Femora strongly clavate in distal half, tibiae expanded and becoming more densely setose towards distal half. Five visible abdominal sternites (sometimes appearing as six due to weak fusion between ventrites V and VI). Aedeagus with median lobe bulbous, heavily sclerotized, median dorsal projection coming to point at apex (triangular), reaching past apical collar; apical collar with dense setae surrounding opening; dorsal parameres slender, longer than median lobe but not extending past apical collar, with lateral subapical setae on each side; apical digiform process extending past apical collar, bent ventrad near apex (
<figureCitation box="[483,572,483,510]" captionStart="Figure 5" captionStartId="9.[140,212,1408,1430]" captionTargetBox="[156,1106,208,1373]" captionTargetId="figure@9.[140,1108,165,1389]" captionTargetPageId="9" captionText="Figure 5. A–F Dorsal and lateral view of the aedeagus A Cephennium anophthalmicum Brendel B Cephennium celsifrons Hopp &amp; Caterino C Cephennium grandarboreum Hopp &amp; Caterino D Cephennium canestroi Hopp &amp; Caterino E Cephennium urbanum Hopp &amp; Caterino F Cephennium aridum Hopp &amp; Caterino." figureDoi="http://doi.org/10.5281/zenodo.3770035" httpUri="https://zenodo.org/record/3770035/files/figure.png" pageId="13" pageNumber="44">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph blockId="13.[140,1108,167,686]" box="[185,483,519,545]" pageId="13" pageNumber="44">
<emphasis bold="true" box="[185,277,519,545]" pageId="13" pageNumber="44">
<emphasis box="[185,272,519,545]" inLineHeading="true" pageId="13" pageNumber="44" reason="1">Female</emphasis>
.
</emphasis>
Identical to male.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="44" type="biology_ecology">
<paragraph blockId="13.[140,1108,167,686]" pageId="13" pageNumber="44">
<emphasis bold="true" box="[185,283,553,579]" pageId="13" pageNumber="44">
<emphasis box="[185,278,553,579]" inLineHeading="true" pageId="13" pageNumber="44" reason="1">Biology</emphasis>
.
</emphasis>
Specimens were extracted from redwood litter and a combination of maple and redwood litter with the use of Berlese funnels.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="44" type="distribution">
<paragraph blockId="13.[140,1108,167,686]" pageId="13" pageNumber="44">
<emphasis bold="true" box="[185,344,624,650]" pageId="13" pageNumber="44">
<emphasis box="[185,339,624,650]" inLineHeading="true" pageId="13" pageNumber="44" reason="1">Distribution</emphasis>
.
</emphasis>
This species is known from a few localities near Big Sur, Monterey County, CA (Fig. 6).
</paragraph>
</subSubSection>
</treatment>
</document>