281 lines
38 KiB
XML
281 lines
38 KiB
XML
<document id="957092DA7B159DEEA5D7E0B68B5C9D83" ID-CLB-Dataset="21790" ID-DOI="10.11646/zootaxa.4786.1.10" ID-GBIF-Dataset="b8b4f8df-8f3b-41ac-aa3c-a465c8093450" ID-ISSN="1175-5326" ID-Zenodo-Dep="3864941" ID-ZooBank="2335111C-05A5-4CCD-9A50-4B10350FEB90" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590737384037" checkinUser="plazi" docAuthor="Yin, Zi-Wei & Lin, Ye-Jie" docDate="2020" docId="413E87CFF246FFC670BCFCC3FAABFE29" docLanguage="en" docName="zootaxa.4786.1.10.pdf" docOrigin="Zootaxa 4786 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tibetyrus formicarius Yin & Lin 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="136" masterDocId="BD07FFB7F244FFC3702BFFE2FFE1FFDE" masterDocTitle="Tibetyrus gen. nov., a new myrmecophilous Tyrini from Xizang, China (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="137" masterPageNumber="131" pageNumber="133" updateTime="1698841864291" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="C2A9BCDCB2F577F51C89439FE6D9B30F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E3EEC9C5D16C2AB8B92D1BAE45D4616D">Tibetyrus gen. nov., a new myrmecophilous Tyrini from Xizang, China (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="E4A00CCBB226D65288DF38A4F2A7B769" type="personal">
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<mods:roleTerm id="4FA4ECAF1C2D7AE74A20060E837B66A5">Author</mods:roleTerm>
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<mods:namePart id="B93AA48536A80843B14878CB4DF6D12E">Yin, Zi-Wei</mods:namePart>
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<mods:affiliation id="35C432B9DAEEA98453E07492BC25D213">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China. pselaphinae @ gmail. com; https: // orcid. org / 0000 - 0001 - 6659 - 9448</mods:affiliation>
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</mods:name>
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<mods:name id="45301E5BE04C925433264C3F88422307" type="personal">
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<mods:roleTerm id="4189EBDEDB5150C0FA02203F3FA433B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F313CB2899CFAB29C9CA44D91B80D376">Lin, Ye-Jie</mods:namePart>
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<mods:affiliation id="3DCF6666639BA39CFFAB2D38F21E330C">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:typeOfResource id="88849351CCD089BE91CA2943A0FABA22">text</mods:typeOfResource>
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<mods:title id="A16ED9C713D6651DDFBE3FCE6DD8C6C6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5AC1BD7838634AFBDF9CB3CF3999A4D1">
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<mods:date id="7B1C10E017053D29AD30D8D3338BC053">2020</mods:date>
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<mods:number id="788E88F52F27A4585D1FF837BBEB23AA">2020-05-29</mods:number>
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<mods:number id="E422CF4C292D4D9F6B92EEB74B6C8A9A">4786</mods:number>
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<mods:classification id="484CFDEBA0A5E21F6947F2576AD055D0">journal article</mods:classification>
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<mods:identifier id="8820A3517A66A1A3B7FC263281A47DAF" type="DOI">10.11646/zootaxa.4786.1.10</mods:identifier>
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<treatment id="413E87CFF246FFC670BCFCC3FAABFE29" ID-DOI="http://doi.org/10.5281/zenodo.3866572" ID-GBIF-Taxon="164392496" ID-Zenodo-Dep="3866572" LSID="urn:lsid:plazi:treatment:413E87CFF246FFC670BCFCC3FAABFE29" httpUri="http://treatment.plazi.org/id/413E87CFF246FFC670BCFCC3FAABFE29" lastPageId="5" lastPageNumber="136" pageId="2" pageNumber="133">
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<subSubSection id="818D6552F246FFC170BCFCC3FD6EFCE5" box="[151,655,801,828]" pageId="2" pageNumber="133" type="nomenclature">
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<paragraph id="C92836D9F246FFC170BCFCC3FD6EFCE5" blockId="2.[151,655,801,863]" box="[151,655,801,828]" pageId="2" pageNumber="133">
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<heading id="926081B5F246FFC170BCFCC3FD6EFCE5" bold="true" box="[151,655,801,828]" fontSize="11" level="1" pageId="2" pageNumber="133" reason="1">
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<emphasis id="FBE3EACBF246FFC170BCFCC3FD6EFCE5" bold="true" box="[151,655,801,828]" pageId="2" pageNumber="133">
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<taxonomicName id="0E974D5AF246FFC170BCFCC3FDC4FCE5" ID-CoL="8YBW3" authority="Yin & Lin" authorityName="Yin & Lin" authorityYear="2020" box="[151,549,801,828]" class="Insecta" family="Staphylinidae" genus="Tibetyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="formicarius" status="sp. nov.">
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<emphasis id="FBE3EACBF246FFC170BCFCC3FE7FFCE5" bold="true" box="[151,414,801,828]" italics="true" pageId="2" pageNumber="133">Tibetyrus formicarius</emphasis>
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Yin & Lin
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</taxonomicName>
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,
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<taxonomicNameLabel id="E0D057B0F246FFC17219FCC3FD6EFCE5" box="[562,655,801,827]" pageId="2" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="818D6552F246FFC170BCFCA7FEEAFC81" box="[151,267,837,863]" pageId="2" pageNumber="133" type="description">
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<paragraph id="C92836D9F246FFC170BCFCA7FEEAFC81" blockId="2.[151,655,801,863]" box="[151,267,837,863]" pageId="2" pageNumber="133">
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(
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<figureCitation id="51AC2A5CF246FFC170B4FCA7FEE3FC81" box="[159,258,837,863]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1443,1468]" captionStartId-1="4.[151,250,1836,1861]" captionStartId-2="5.[151,250,1552,1577]" captionTargetBox-0="[160,1436,189,1412]" captionTargetBox-1="[151,1436,181,1806]" captionTargetBox-2="[151,1435,536,1527]" captionTargetId-0="figure@3.[151,1436,179,1416]" captionTargetId-1="figure@4.[151,1436,181,1811]" captionTargetId-2="figure@5.[151,1435,525,1527]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." captionText-1="FIGURE 2. Tibetyrus formicarius, male. A. Head, pronotum, and elytral base. B. Meso- and metaventrite. C. Sternite 7 (IX). D–F. Aedeagus, in dorsal (D), lateral (E), and ventral (F) view. Scale bars: 0.5 mm in A, B; 0.1 mm in C; 0.2 mm in D–F." captionText-2="FIGURE 3. Host ant (A, B) and collection environment (C, D) of Tibetyrus formicarius. A. Habitus, lateral. B. Head, in fullface view. C, D. Jiarelongba Valley, where the ant nests were found under stones beside the pass." figureDoi-0="http://doi.org/10.5281/zenodo.3864943" figureDoi-1="http://doi.org/10.5281/zenodo.3864945" figureDoi-2="http://doi.org/10.5281/zenodo.3864947" httpUri-0="https://zenodo.org/record/3864943/files/figure.png" httpUri-1="https://zenodo.org/record/3864945/files/figure.png" httpUri-2="https://zenodo.org/record/3864947/files/figure.png" pageId="2" pageNumber="133">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="818D6552F246FFC170BCFC6FFBE4FC31" pageId="2" pageNumber="133" type="materials_examined">
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<paragraph id="C92836D9F246FFC170BCFC6FFBE4FC31" blockId="2.[151,1437,908,2015]" pageId="2" pageNumber="133">
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<emphasis id="FBE3EACBF246FFC170BCFC6FFEA9FC79" bold="true" box="[151,328,908,935]" pageId="2" pageNumber="133">Type material.</emphasis>
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<materialsCitation id="79FF3C84F246FFC17178FC6EFBE4FC31" ID-GBIF-Occurrence="2628389301" collectingDate="2019-10-18" collectionCode="SNUC" country="China" county="Gongbo'gyamda Co." elevation="3485" latitude="30.0075" location="Bahe Town" longLatPrecision="6" longitude="93.7818" municipality="Nyingchi" pageId="2" pageNumber="133" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
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<emphasis id="FBE3EACBF246FFC17178FC6EFD8DFC79" bold="true" box="[339,620,908,935]" pageId="2" pageNumber="133">
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<typeStatus id="162C887BF246FFC17178FC6EFE17FC79" box="[339,502,908,935]" pageId="2" pageNumber="133" type="holotype">HOLOTYPE</typeStatus>
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:
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<collectingCountry id="B1807649F246FFC1722EFC6EFD83FC79" box="[517,610,908,935]" name="China" pageId="2" pageNumber="133">CHINA</collectingCountry>
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:
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</emphasis>
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<specimenCount id="DF91FD50F246FFC1725DFC6FFD6CFC79" box="[630,653,909,935]" pageId="2" pageNumber="133" type="male">♂</specimenCount>
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, ‘China:
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<collectingRegion id="0B53F83BF246FFC172D0FC6FFCAEFC79" box="[763,847,909,935]" country="China" name="Xizang" pageId="2" pageNumber="133">Xizang</collectingRegion>
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,
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<collectingMunicipality id="294CACA3F246FFC17375FC6FFC28FC79" box="[862,969,909,935]" pageId="2" pageNumber="133">Nyingchi</collectingMunicipality>
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,
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<collectingCounty id="20494E55F246FFC173FCFC6FFB25FC79" box="[983,1220,909,935]" pageId="2" pageNumber="133">Gongbo'gyamda Co.</collectingCounty>
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,
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<location id="CC486002F246FFC174F8FC6FFAB9FC79" LSID="urn:lsid:plazi:treatment:413E87CFF246FFC670BCFCC3FAABFE29:CC486002F246FFC174F8FC6FFAB9FC79" box="[1235,1368,909,935]" country="China" county="Gongbo'gyamda Co." latitude="30.0075" longLatPrecision="6" longitude="93.7818" municipality="Nyingchi" name="Bahe Town" pageId="2" pageNumber="133" stateProvince="Xizang">Bahe Town</location>
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, Jiare Vill.,
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<location id="CC486002F246FFC170FDFC53FEBBFC15" LSID="urn:lsid:plazi:treatment:413E87CFF246FFC670BCFCC3FAABFE29:CC486002F246FFC170FDFC53FEBBFC15" box="[214,346,945,971]" country="China" county="Gongbo'gyamda Co." latitude="30.0075" longLatPrecision="6" longitude="93.7818" municipality="Nyingchi" name="Jiarelongba" pageId="2" pageNumber="133" stateProvince="Xizang">Jiarelongba</location>
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,
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<geoCoordinate id="ACA3501EF246FFC1714FFC52FE04FC15" box="[356,485,944,971]" degrees="30.0075" direction="north" orientation="latitude" pageId="2" pageNumber="133" precision="5" value="30.0075">30.0075º N</geoCoordinate>
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,
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<geoCoordinate id="ACA3501EF246FFC171DBFC53FD8CFC15" box="[496,621,945,971]" degrees="93.7818" direction="east" orientation="longitude" pageId="2" pageNumber="133" precision="5" value="93.7818">93.7818º E</geoCoordinate>
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,
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<quantity id="0E6F9B3CF246FFC17252FC52FD37FC15" box="[633,726,944,971]" metricMagnitude="3" metricUnit="m" metricValue="3.485" pageId="2" pageNumber="133" unit="m" value="3485.0">
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<elevation id="42BAD1EAF246FFC17252FC52FD37FC15" box="[633,726,944,971]" metricMagnitude="3" metricUnit="m" metricValue="3.485" pageId="2" pageNumber="133" unit="m" value="3485.0">3,485 m</elevation>
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</quantity>
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,
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<date id="BD291019F246FFC172C9FC53FCB5FC15" box="[738,852,945,971]" pageId="2" pageNumber="133" value="2019-10-18">
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<collectingDate id="AD6DE9F1F246FFC172C9FC53FCB5FC15" box="[738,852,945,971]" pageId="2" pageNumber="133" value="2019-10-18">18.x.2019</collectingDate>
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</date>
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, Lin Y-J, TŨỮḔDZOiDZAEÙDZAEȐDZ’ (
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<collectionCode id="AF86AE1CF246FFC1756BFC53FA70FC15" box="[1344,1425,945,971]" pageId="2" pageNumber="133">SNUC</collectionCode>
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).
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<emphasis id="FBE3EACBF246FFC170BCFC36FE38FC31" bold="true" box="[151,473,980,1007]" pageId="2" pageNumber="133">
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PARATYTYPES:
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<collectingCountry id="B1807649F246FFC17159FC36FE2EFC31" box="[370,463,980,1007]" name="China" pageId="2" pageNumber="133">CHINA</collectingCountry>
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:
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</emphasis>
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<specimenCount id="DF91FD50F246FFC171CBFC37FDC1FC31" box="[480,544,981,1007]" pageId="2" pageNumber="133" type="female">2 ♀♀</specimenCount>
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, same collection data as holotype (
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<collectionCode id="AF86AE1CF246FFC17383FC37FC18FC31" box="[936,1017,981,1007]" pageId="2" pageNumber="133">SNUC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="818D6552F246FFC070ECFC1AFE3EF8B2" lastPageId="3" lastPageNumber="134" pageId="2" pageNumber="133" type="description">
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<paragraph id="C92836D9F246FFC170ECFC1AFB86FB85" blockId="2.[151,1437,908,2015]" pageId="2" pageNumber="133">
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<emphasis id="FBE3EACBF246FFC170ECFC1AFEB8FBCD" bold="true" box="[199,345,1016,1043]" pageId="2" pageNumber="133">Description.</emphasis>
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Male (
|
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<figureCitation id="51AC2A5CF246FFC17182FC1BFDE3FBCD" box="[425,514,1017,1043]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="2" pageNumber="133">Fig. 1A</figureCitation>
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). Body generally elongate, length (combined length of head, pronotum, elytra and abdomen)
|
||
<quantity id="0E6F9B3CF246FFC1713BFBFFFE96FBE9" box="[272,375,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="3.71" pageId="2" pageNumber="133" unit="mm" value="3.71">3.71 mm</quantity>
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||
, color uniformly reddish-brown; dorsal surface of whole body finely punctate, with short decumbent setae of elytra and abdomen slightly denser than those on head and pronotum.
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||
</paragraph>
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<paragraph id="C92836D9F246FFC170ECFB87FF1AF98D" blockId="2.[151,1437,908,2015]" pageId="2" pageNumber="133">
|
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Head (
|
||
<figureCitation id="51AC2A5CF246FFC17139FB87FE8EFBA1" box="[274,367,1125,1151]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="2" pageNumber="133">Figs 1A</figureCitation>
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,
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||
<figureCitation id="51AC2A5CF246FFC17157FB87FE41FBA1" box="[380,416,1125,1151]" captionStart="FIGURE 2" captionStartId="4.[151,250,1836,1861]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure@4.[151,1436,181,1811]" captionTargetPageId="4" captionText="FIGURE 2. Tibetyrus formicarius, male. A. Head, pronotum, and elytral base. B. Meso- and metaventrite. C. Sternite 7 (IX). D–F. Aedeagus, in dorsal (D), lateral (E), and ventral (F) view. Scale bars: 0.5 mm in A, B; 0.1 mm in C; 0.2 mm in D–F." figureDoi="http://doi.org/10.5281/zenodo.3864945" httpUri="https://zenodo.org/record/3864945/files/figure.png" pageId="2" pageNumber="133">2A</figureCitation>
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) lengthily and bluntly triangular, length from anterior margin of clypeus to head base
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||
<quantity id="0E6F9B3CF246FFC17540FB87FF24FB7D" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="133" unit="mm" value="0.71">0.71 mm</quantity>
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||
, width across eyes
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||
<quantity id="0E6F9B3CF246FFC1718AFB6AFDE8FB7D" box="[417,521,1160,1187]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="2" pageNumber="133" unit="mm" value="0.57">0.57 mm</quantity>
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; vertexal foveae at level of midline of eyes; frons dilated apically, longitudinally and smoothly impressed between antennal tubercles; postocular margins rounded, with tufts of dense setae on each side; eyes moderately prominent, each composed of about 15 facets. Maxillary palpi (
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||
<figureCitation id="51AC2A5CF246FFC17443FB33FB5FFB35" box="[1128,1214,1233,1259]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="2" pageNumber="133">Fig. 1B</figureCitation>
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) with short, evenly distributed sensillar setae. Antennae elongate (
|
||
<figureCitation id="51AC2A5CF246FFC172B4FB17FD16FAD1" box="[671,759,1269,1295]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="2" pageNumber="133">Fig. 1A</figureCitation>
|
||
), length
|
||
<quantity id="0E6F9B3CF246FFC17372FB17FC21FAD1" box="[857,960,1269,1295]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="133" unit="mm" value="2.3">2.30 mm</quantity>
|
||
; antennomeres 1 (scapes) distinctly longer than 2 (pedicels) and 3 combined, but shorter than combined length of antennomeres 2 to 4, approximately 1.8× as long as wide, antennomeres 2 and 3 of similar size, antennomeres 4–6 longer than 2, successively wider, antennomeres 7 slightly wider than 6, transverse, antennomeres 8 slightly narrower and much shorter than 7, antennomeres 9–11 (
|
||
<figureCitation id="51AC2A5CF246FFC170F4FA67FED4FA41" box="[223,309,1413,1439]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="2" pageNumber="133">Fig. 1C</figureCitation>
|
||
) strongly enlarged to form distinct clubs, 9 about three times as long as and twice as wide as 8, 10 much larger than 9, broadly and deeply excavated on ventral surface, 11 much smaller than 10 and slightly larger than 9, their bases strongly elongate and constricted to form long projections, which extend downwards to exceed over half of antennomeres 10, with disk-like protuberance near bases of projections, basal portions of antennomeres 11 also circularly impressed, within impressions one fovea-like setose pit, apices much broader than bases and rounded.
|
||
</paragraph>
|
||
<paragraph id="C92836D9F246FFC170ECF9BFFEECF961" blockId="2.[151,1437,908,2015]" pageId="2" pageNumber="133">
|
||
Pronotum (
|
||
<figureCitation id="51AC2A5CF246FFC1716DF9BFFE44F9A9" box="[326,421,1629,1655]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="2" pageNumber="133">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="51AC2A5CF246FFC17198F9BFFE36F9A9" box="[435,471,1629,1655]" captionStart="FIGURE 2" captionStartId="4.[151,250,1836,1861]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure@4.[151,1436,181,1811]" captionTargetPageId="4" captionText="FIGURE 2. Tibetyrus formicarius, male. A. Head, pronotum, and elytral base. B. Meso- and metaventrite. C. Sternite 7 (IX). D–F. Aedeagus, in dorsal (D), lateral (E), and ventral (F) view. Scale bars: 0.5 mm in A, B; 0.1 mm in C; 0.2 mm in D–F." figureDoi="http://doi.org/10.5281/zenodo.3864945" httpUri="https://zenodo.org/record/3864945/files/figure.png" pageId="2" pageNumber="133">2A</figureCitation>
|
||
) elongate, mid-length
|
||
<quantity id="0E6F9B3CF246FFC172F1F9BFFCA3F9A9" box="[730,834,1629,1655]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="133" unit="mm" value="0.76">0.76 mm</quantity>
|
||
, maximum width
|
||
<quantity id="0E6F9B3CF246FFC17439F9BFFB9BF9A9" box="[1042,1146,1629,1655]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="133" unit="mm" value="0.66">0.66 mm</quantity>
|
||
, with slightly arcuate anterior and sinuate posterior margin, sides rounded at middle, then narrowing toward apex and subparallel in basal two-fifths.
|
||
</paragraph>
|
||
<paragraph id="C92836D9F246FFC170ECF92BFD1CF8AD" blockId="2.[151,1437,908,2015]" pageId="2" pageNumber="133">
|
||
Elytra (
|
||
<figureCitation id="51AC2A5CF246FFC17135F92BFE99F93D" box="[286,376,1737,1763]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="2" pageNumber="133">Fig. 1A</figureCitation>
|
||
) much wider than long, length along suture
|
||
<quantity id="0E6F9B3CF246FFC173A8F92BFC0DF93D" box="[899,1004,1737,1763]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="133" unit="mm" value="0.89">0.89 mm</quantity>
|
||
, maximum width
|
||
<quantity id="0E6F9B3CF246FFC174EAF92AFACBF93D" box="[1217,1322,1736,1763]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="133" unit="mm" value="1.15">1.15 mm</quantity>
|
||
, posterior margin with row of dense setae that are longer than those on disc. Mesoventrite (
|
||
<figureCitation id="51AC2A5CF246FFC17409F90FFB99F8D9" box="[1058,1144,1773,1799]" captionStart="FIGURE 2" captionStartId="4.[151,250,1836,1861]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure@4.[151,1436,181,1811]" captionTargetPageId="4" captionText="FIGURE 2. Tibetyrus formicarius, male. A. Head, pronotum, and elytral base. B. Meso- and metaventrite. C. Sternite 7 (IX). D–F. Aedeagus, in dorsal (D), lateral (E), and ventral (F) view. Scale bars: 0.5 mm in A, B; 0.1 mm in C; 0.2 mm in D–F." figureDoi="http://doi.org/10.5281/zenodo.3864945" httpUri="https://zenodo.org/record/3864945/files/figure.png" pageId="2" pageNumber="133">Fig. 2B</figureCitation>
|
||
) strongly transverse, with thin ridges lateral to middle; metaventrite also transverse, with large, lamina-like postero-submedial projections, posterior margin of each projection strongly curved ventrally to form large spine; setae of meso- and metaventrite much denser and longer in lateral areas than at middle.
|
||
</paragraph>
|
||
<paragraph id="C92836D9F246FFC170ECF89FFD8CF849" blockId="2.[151,1437,908,2015]" box="[199,621,1917,1943]" pageId="2" pageNumber="133">Legs elongate, lacking modifications.</paragraph>
|
||
<paragraph id="C92836D9F246FFC170ECF843FA8FF801" blockId="2.[151,1437,908,2015]" pageId="2" pageNumber="133">
|
||
Abdomen about as long as wide, widest at posterior margin of tergite 1 (IV), mid-length
|
||
<quantity id="0E6F9B3CF246FFC1749CF842FAFFF865" box="[1207,1310,1952,1979]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="2" pageNumber="133" unit="mm" value="1.35">1.35 mm</quantity>
|
||
, maximum width
|
||
<quantity id="0E6F9B3CF246FFC170F5F827FEA4F801" box="[222,325,1989,2015]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="2" pageNumber="133" unit="mm" value="1.28">1.28 mm</quantity>
|
||
. Sternite 7 (IX) (
|
||
<figureCitation id="51AC2A5CF246FFC17229F827FDB9F801" box="[514,600,1989,2015]" captionStart="FIGURE 2" captionStartId="4.[151,250,1836,1861]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure@4.[151,1436,181,1811]" captionTargetPageId="4" captionText="FIGURE 2. Tibetyrus formicarius, male. A. Head, pronotum, and elytral base. B. Meso- and metaventrite. C. Sternite 7 (IX). D–F. Aedeagus, in dorsal (D), lateral (E), and ventral (F) view. Scale bars: 0.5 mm in A, B; 0.1 mm in C; 0.2 mm in D–F." figureDoi="http://doi.org/10.5281/zenodo.3864945" httpUri="https://zenodo.org/record/3864945/files/figure.png" pageId="2" pageNumber="133">Fig. 2C</figureCitation>
|
||
) semi-membranous, elongate, with round apex and dense apical setae.
|
||
</paragraph>
|
||
<caption id="9DE86651F247FFC070BCFA41FD28FA01" ID-DOI="http://doi.org/10.5281/zenodo.3864943" ID-Zenodo-Dep="3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="3" pageNumber="134" startId="3.[151,250,1443,1468]" targetBox="[160,1436,189,1412]" targetPageId="3">
|
||
<paragraph id="C92836D9F247FFC070BCFA41FD28FA01" blockId="3.[151,1437,1443,1504]" pageId="3" pageNumber="134">
|
||
<emphasis id="FBE3EACBF247FFC070BCFA41FEF2FA62" bold="true" box="[151,275,1443,1468]" pageId="3" pageNumber="134">FIGURE 1.</emphasis>
|
||
<taxonomicName id="0E974D5AF247FFC07131FA46FE15FA62" authorityName="Yin & Lin" authorityYear="2020" box="[282,500,1444,1468]" class="Insecta" family="Staphylinidae" genus="Tibetyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="formicarius">
|
||
<emphasis id="FBE3EACBF247FFC07131FA46FE15FA62" box="[282,500,1444,1468]" italics="true" pageId="3" pageNumber="134">Tibetyrus formicarius</emphasis>
|
||
</taxonomicName>
|
||
(A–C. Male. D. Female).
|
||
<emphasis id="FBE3EACBF247FFC0732BFA41FCF9FA62" bold="true" box="[768,792,1443,1468]" pageId="3" pageNumber="134">A.</emphasis>
|
||
Habitus.
|
||
<emphasis id="FBE3EACBF247FFC07351FA46FC73FA62" bold="true" box="[890,914,1444,1468]" pageId="3" pageNumber="134">B.</emphasis>
|
||
Right maxillary palpus.
|
||
<emphasis id="FBE3EACBF247FFC074A6FA41FB24FA62" bold="true" box="[1165,1221,1443,1468]" pageId="3" pageNumber="134">C, D.</emphasis>
|
||
Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C92836D9F247FFC070ECF9ECFBB8F967" blockId="3.[151,1437,1550,2045]" pageId="3" pageNumber="134">
|
||
Aedeagus (
|
||
<figureCitation id="51AC2A5CF247FFC0716EF9ECFE67F9F6" box="[325,390,1550,1576]" captionStart="FIGURE 2" captionStartId="4.[151,250,1836,1861]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure@4.[151,1436,181,1811]" captionTargetPageId="4" captionText="FIGURE 2. Tibetyrus formicarius, male. A. Head, pronotum, and elytral base. B. Meso- and metaventrite. C. Sternite 7 (IX). D–F. Aedeagus, in dorsal (D), lateral (E), and ventral (F) view. Scale bars: 0.5 mm in A, B; 0.1 mm in C; 0.2 mm in D–F." figureDoi="http://doi.org/10.5281/zenodo.3864945" httpUri="https://zenodo.org/record/3864945/files/figure.png" pageId="3" pageNumber="134">Fig. 2</figureCitation>
|
||
D–F) elongate, length
|
||
<quantity id="0E6F9B3CF247FFC072A8F9ECFD08F9F7" box="[643,745,1550,1577]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="134" unit="mm" value="0.62">0.62 mm</quantity>
|
||
, median lobe with large, nearly symmetric basal capsule, apical half strongly twisted, narrowing apicad, forming pointed apex; endophallus complex in structure, composed of one elongate, broad, curved and weakly sclerotized plate-like sclerite abruptly narrowed and acute at apex, one relatively broad and bi-sinuate sclerite narrowing from basal two-thirds to apex, and one much shorter and thinner elongate sclerite near middle; parameres reduced and symmetric, lacking apical setae.
|
||
</paragraph>
|
||
<paragraph id="C92836D9F247FFC070ECF920FE3EF8B2" blockId="3.[151,1437,1550,2045]" pageId="3" pageNumber="134">
|
||
Female. General appearance similar to male; antennae shorter, clubs (
|
||
<figureCitation id="51AC2A5CF247FFC073F3F920FBCEF902" box="[984,1071,1730,1756]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="3" pageNumber="134">Fig. 1D</figureCitation>
|
||
) unmodified; metaventrite lacking projections. Measurements (as for male): body length
|
||
<quantity id="0E6F9B3CF247FFC07336F907FC23F8DF" box="[797,962,1765,1793]" metricMagnitude="-3" metricUnit="m" metricValue="3.605" metricValueMax="3.71" metricValueMin="3.5" pageId="3" pageNumber="134" unit="mm" value="3.605" valueMax="3.71" valueMin="3.5">3.50–3.71 mm</quantity>
|
||
, length/width of head 0.65–0.66/
|
||
<quantity id="0E6F9B3CF247FFC07519F907FA79F8DF" box="[1330,1432,1765,1793]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="134" unit="mm" value="0.56">0.56 mm</quantity>
|
||
, pronotum 0.72–0.76/
|
||
<quantity id="0E6F9B3CF247FFC071ACF8EBFDCFF8FB" box="[391,558,1801,1829]" metricMagnitude="-4" metricUnit="m" metricValue="6.45" metricValueMax="6.5" metricValueMin="6.4" pageId="3" pageNumber="134" unit="mm" value="0.645" valueMax="0.65" valueMin="0.64">0.64–0.65 mm</quantity>
|
||
, elytra 0.85–0.91/
|
||
<quantity id="0E6F9B3CF247FFC0732AF8EBFC46F8FB" box="[769,935,1801,1829]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" metricValueMax="1.19" metricValueMin="1.15" pageId="3" pageNumber="134" unit="mm" value="1.17" valueMax="1.19" valueMin="1.15">1.15–1.19 mm</quantity>
|
||
, abdomen 1.27–1.39/
|
||
<quantity id="0E6F9B3CF247FFC0748BF8E8FAA6F8FB" box="[1184,1351,1802,1829]" metricMagnitude="-3" metricUnit="m" metricValue="1.345" metricValueMax="1.36" metricValueMin="1.33" pageId="3" pageNumber="134" unit="mm" value="1.345" valueMax="1.36" valueMin="1.33">1.33–1.36 mm</quantity>
|
||
, length of antennae
|
||
<quantity id="0E6F9B3CF247FFC0710AF8CFFE69F897" box="[289,392,1837,1865]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="3" pageNumber="134" unit="mm" value="1.85">1.85 mm</quantity>
|
||
, antennomeres 1 approximately as long as 2–3 combined and 1.3× as long as wide; each eye composed of about 18 facets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="818D6552F247FFC670ECF897FBFBFE0D" lastPageId="5" lastPageNumber="136" pageId="3" pageNumber="134" type="distribution">
|
||
<paragraph id="C92836D9F247FFC070ECF897FD50F84E" blockId="3.[151,1437,1550,2045]" box="[199,689,1909,1936]" pageId="3" pageNumber="134">
|
||
<emphasis id="FBE3EACBF247FFC070ECF897FE80F84E" bold="true" box="[199,353,1909,1936]" pageId="3" pageNumber="134">Distribution.</emphasis>
|
||
Southwestern
|
||
<collectingCountry id="B1807649F247FFC0722CF894FDAFF84E" box="[519,590,1910,1936]" name="China" pageId="3" pageNumber="134">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="0B53F83BF247FFC07272F894FD4CF84E" box="[601,685,1910,1936]" country="China" name="Xizang" pageId="3" pageNumber="134">Xizang</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C92836D9F247FFC670ECF87BFBFBFE0D" blockId="3.[151,1437,1550,2045]" lastBlockId="5.[151,1436,153,503]" lastPageId="5" lastPageNumber="136" pageId="3" pageNumber="134">
|
||
<emphasis id="FBE3EACBF247FFC070ECF87BFDD4F86A" bold="true" box="[199,565,1945,1972]" pageId="3" pageNumber="134">Biological and collection notes.</emphasis>
|
||
The collection site is located near the entrance of a deep, north-south oriented valley named Jiarelongba, approximately
|
||
<quantity id="0E6F9B3CF247FFC0725CF85CFD53F806" box="[631,690,1982,2008]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="134" unit="km" value="2.0">2 km</quantity>
|
||
north of Jiare Village, and
|
||
<quantity id="0E6F9B3CF247FFC073D8F85CFBDAF806" box="[1011,1083,1982,2008]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="134" unit="km" value="11.0">11 km</quantity>
|
||
west of Basongcuo Lake. The valley has abundant vegetation, with a broad pass and a creek on the west side (
|
||
<figureCitation id="51AC2A5CF247FFC07400F800FB4BF823" box="[1067,1194,2018,2045]" captionStart="FIGURE 3" captionStartId="5.[151,250,1552,1577]" captionTargetBox="[151,1435,536,1527]" captionTargetId="figure@5.[151,1435,525,1527]" captionTargetPageId="5" captionText="FIGURE 3. Host ant (A, B) and collection environment (C, D) of Tibetyrus formicarius. A. Habitus, lateral. B. Head, in fullface view. C, D. Jiarelongba Valley, where the ant nests were found under stones beside the pass." figureDoi="http://doi.org/10.5281/zenodo.3864947" httpUri="https://zenodo.org/record/3864947/files/figure.png" pageId="3" pageNumber="134">Fig. 3C, D</figureCitation>
|
||
). The field work was conducted at around 9:00 a.m. Two individuals of the pselaphid were first discovered in tunnels of an ant nest found under an approximately
|
||
<quantity id="0E6F9B3CF241FFC6718CFF5FFE0DFF09" box="[423,492,189,215]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="136" unit="cm" value="30.0">30 cm</quantity>
|
||
<superScript id="3EE29B91F241FFC671C7FF5EFE15FF14" attach="left" box="[492,500,188,202]" fontSize="6" pageId="5" pageNumber="136">2</superScript>
|
||
stone on the ground just beside the pass. Later a third individual was collected from another nest in the same area less than
|
||
<quantity id="0E6F9B3CF241FFC67267FF02FD67FF25" box="[588,646,224,251]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="136" unit="m" value="50.0">50 m</quantity>
|
||
away. The host ant was later identified as a member of the genus
|
||
<taxonomicName id="0E974D5AF241FFC67542FF00FF2CFEC1" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FBE3EACBF241FFC67542FF00FF2CFEC1" italics="true" pageId="5" pageNumber="136">Formica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="51AC2A5CF241FFC670F5FEE7FE7BFEC1" box="[222,410,261,287]" captionStart="FIGURE 3" captionStartId="5.[151,250,1552,1577]" captionTargetBox="[151,1435,536,1527]" captionTargetId="figure@5.[151,1435,525,1527]" captionTargetPageId="5" captionText="FIGURE 3. Host ant (A, B) and collection environment (C, D) of Tibetyrus formicarius. A. Habitus, lateral. B. Head, in fullface view. C, D. Jiarelongba Valley, where the ant nests were found under stones beside the pass." figureDoi="http://doi.org/10.5281/zenodo.3864947" httpUri="https://zenodo.org/record/3864947/files/figure.png" pageId="5" pageNumber="136">Fig. 3A, B; S.-Y</figureCitation>
|
||
. Zhou & H. Ran, pers. comm.). The new species is apparently myrmecophilous because of the repeated circumstances of its collection, and it may possibly be a synoekete or a symphile due to the presence of what may be paired trichomes on the postocular margin. Similar trichomes of the head are commonly found in many myrmecophilous pselaphine genera, especially in the tribe
|
||
<taxonomicName id="0E974D5AF241FFC67354FE93FC02FE55" authorityName="Reitter" authorityYear="1882" box="[895,995,369,395]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
||
. The markedly constricted elytral base and loss of hind wings are typical for various myrmecophilous/termitophilous beetle groups. However, additional field observations are necessary to further reveal the biology of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="9DE86651F240FFC770BCF8CEFA85F8B7" ID-DOI="http://doi.org/10.5281/zenodo.3864945" ID-Zenodo-Dep="3864945" httpUri="https://zenodo.org/record/3864945/files/figure.png" pageId="4" pageNumber="135" startId="4.[151,250,1836,1861]" targetBox="[151,1436,181,1806]" targetPageId="4">
|
||
<paragraph id="C92836D9F240FFC770BCF8CEFA85F8B7" blockId="4.[151,1437,1836,1897]" pageId="4" pageNumber="135">
|
||
<emphasis id="FBE3EACBF240FFC770BCF8CEFEF5F89B" bold="true" box="[151,276,1836,1861]" pageId="4" pageNumber="135">FIGURE 2.</emphasis>
|
||
<taxonomicName id="0E974D5AF240FFC77130F8CFFE17F89B" authorityName="Yin & Lin" authorityYear="2020" box="[283,502,1837,1861]" class="Insecta" family="Staphylinidae" genus="Tibetyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="formicarius">
|
||
<emphasis id="FBE3EACBF240FFC77130F8CFFE17F89B" box="[283,502,1837,1861]" italics="true" pageId="4" pageNumber="135">Tibetyrus formicarius</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="FBE3EACBF240FFC77268F8CEFDBAF89B" bold="true" box="[579,603,1836,1861]" pageId="4" pageNumber="135">A.</emphasis>
|
||
Head, pronotum, and elytral base.
|
||
<emphasis id="FBE3EACBF240FFC773E8F8CFFC3BF89B" bold="true" box="[963,986,1837,1861]" pageId="4" pageNumber="135">B.</emphasis>
|
||
Meso- and metaventrite.
|
||
<emphasis id="FBE3EACBF240FFC774CAF8CEFB18F89B" bold="true" box="[1249,1273,1836,1861]" pageId="4" pageNumber="135">C.</emphasis>
|
||
Sternite 7 (IX).
|
||
<emphasis id="FBE3EACBF240FFC770BCF8B3FF28F8B7" bold="true" box="[151,201,1873,1897]" pageId="4" pageNumber="135">D–F.</emphasis>
|
||
Aedeagus, in dorsal (D), lateral (E), and ventral (F) view. Scale bars: 0.5 mm in A, B; 0.1 mm in C; 0.2 mm in D–F.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="818D6552F241FFC670ECFE3FFAABFE29" box="[199,1354,477,503]" pageId="5" pageNumber="136" type="etymology">
|
||
<paragraph id="C92836D9F241FFC670ECFE3FFAABFE29" blockId="5.[151,1436,153,503]" box="[199,1354,477,503]" pageId="5" pageNumber="136">
|
||
<emphasis id="FBE3EACBF241FFC670ECFE3FFEAFFE29" bold="true" box="[199,334,477,503]" pageId="5" pageNumber="136">Etymology.</emphasis>
|
||
The specific epithet recognizes the association between the new species and
|
||
<taxonomicName id="0E974D5AF241FFC67485FE3CFAEFFE29" box="[1198,1294,478,503]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FBE3EACBF241FFC67485FE3CFAEFFE29" box="[1198,1294,478,503]" italics="true" pageId="5" pageNumber="136">Formica</emphasis>
|
||
</taxonomicName>
|
||
ants.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |