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<document id="55E7E8B369B297254E7084825065456A" ID-CLB-Dataset="21790" ID-DOI="10.11646/zootaxa.4786.1.10" ID-GBIF-Dataset="b8b4f8df-8f3b-41ac-aa3c-a465c8093450" ID-ISSN="1175-5326" ID-Zenodo-Dep="3864941" ID-ZooBank="2335111C-05A5-4CCD-9A50-4B10350FEB90" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590737384037" checkinUser="plazi" docAuthor="Yin, Zi-Wei &amp; Lin, Ye-Jie" docDate="2020" docId="413E87CFF245FFC170BCFD57FE9CFD11" docLanguage="en" docName="zootaxa.4786.1.10.pdf" docOrigin="Zootaxa 4786 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tibetyrus Yin &amp; Lin 2020, gen. nov." docType="treatment" docVersion="6" lastPageNumber="133" masterDocId="BD07FFB7F244FFC3702BFFE2FFE1FFDE" masterDocTitle="Tibetyrus gen. nov., a new myrmecophilous Tyrini from Xizang, China (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="137" masterPageNumber="131" pageNumber="132" updateTime="1698841864291" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5820EE68BDDCEB06E279D51BB6E580B8">Tibetyrus gen. nov., a new myrmecophilous Tyrini from Xizang, China (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:name id="7DC439830318E191A894FEBBF858D1F9" type="personal">
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<mods:namePart id="D4A1A86127A0334B66179337BE7F9A15">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="B1C83414ECAB72858EBDA9547402C51B">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China. pselaphinae @ gmail. com; https: // orcid. org / 0000 - 0001 - 6659 - 9448</mods:affiliation>
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<mods:namePart id="0998A506CC3489EBD9F0ABD25AB0B7C3">Lin, Ye-Jie</mods:namePart>
<mods:affiliation id="2A09EE24567FEDA9F53D6E952413C667">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title id="AC00DE5D4A0F6E132462AC208CF17461">Zootaxa</mods:title>
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<mods:date id="23437D46BF6904695805400FAAF3E1FC">2020</mods:date>
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<mods:number id="601E1903979DFE1A78E77E407093A348">2020-05-29</mods:number>
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<mods:number id="D3731C4B54439207A269F6FAFA2D47F4">4786</mods:number>
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<mods:classification id="3A647501F4219D4AEFC8B05EE2282219">journal article</mods:classification>
<mods:identifier id="E8F4CF68D389A5B13172BA1AFE4BE6FB" type="CLB-Dataset">21790</mods:identifier>
<mods:identifier id="7FA2C6975198FC5314CB9435DDC962BA" type="DOI">10.11646/zootaxa.4786.1.10</mods:identifier>
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<treatment id="413E87CFF245FFC170BCFD57FE9CFD11" ID-DOI="http://doi.org/10.5281/zenodo.3866570" ID-GBIF-Taxon="164392495" ID-Zenodo-Dep="3866570" LSID="urn:lsid:plazi:treatment:413E87CFF245FFC170BCFD57FE9CFD11" httpUri="http://treatment.plazi.org/id/413E87CFF245FFC170BCFD57FE9CFD11" lastPageId="2" lastPageNumber="133" pageId="1" pageNumber="132">
<subSubSection id="818D6552F245FFC270BCFD57FDEAFD11" box="[151,523,693,720]" pageId="1" pageNumber="132" type="nomenclature">
<paragraph id="C92836D9F245FFC270BCFD57FDEAFD11" blockId="1.[151,523,693,720]" box="[151,523,693,720]" pageId="1" pageNumber="132">
<heading id="926081B5F245FFC270BCFD57FDEAFD11" bold="true" box="[151,523,693,720]" fontSize="11" level="1" pageId="1" pageNumber="132" reason="1">
<emphasis id="FBE3EACBF245FFC270BCFD57FDEAFD11" bold="true" box="[151,523,693,720]" pageId="1" pageNumber="132">
<taxonomicName id="0E974D5AF245FFC270BCFD57FE71FD11" ID-CoL="8XQDD" authority="Yin &amp; Lin" authorityName="Yin &amp; Lin" authorityYear="2020" box="[151,400,693,720]" class="Insecta" family="Staphylinidae" genus="Tibetyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FBE3EACBF245FFC270BCFD57FEE9FD0E" bold="true" box="[151,264,693,720]" italics="true" pageId="1" pageNumber="132">Tibetyrus</emphasis>
Yin &amp; Lin
</taxonomicName>
,
<taxonomicNameLabel id="E0D057B0F245FFC271B6FD57FDEAFD11" box="[413,523,693,719]" pageId="1" pageNumber="132" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="818D6552F245FFC270BCFD1BFCF7FCCF" box="[151,790,760,785]" pageId="1" pageNumber="132" type="type_taxon">
<paragraph id="C92836D9F245FFC270BCFD1BFCF7FCCF" blockId="1.[151,790,760,785]" box="[151,790,760,785]" pageId="1" pageNumber="132">
<emphasis id="FBE3EACBF245FFC270BCFD1BFEC2FCCF" bold="true" box="[151,291,760,785]" pageId="1" pageNumber="132">
<typeStatus id="162C887BF245FFC270BCFD1BFF2DFCCF" box="[151,204,761,785]" pageId="1" pageNumber="132">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="0E974D5AF245FFC27101FD1BFDE5FCCF" ID-CoL="8YBW3" authority="Yin &amp; Lin, 2020" authorityName="Yin &amp; Lin" authorityYear="2020" box="[298,516,761,785]" class="Insecta" family="Staphylinidae" genus="Tibetyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="formicarius" status="sp. nov.">
<emphasis id="FBE3EACBF245FFC27101FD1BFDE5FCCF" box="[298,516,761,785]" italics="true" pageId="1" pageNumber="132">Tibetyrus formicarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0D057B0F245FFC27220FD18FDBBFCCF" box="[523,602,762,785]" pageId="1" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
(here designated).
</paragraph>
</subSubSection>
<subSubSection id="818D6552F245FFC270BCFCDDFC04FC34" pageId="1" pageNumber="132" type="diagnosis">
<paragraph id="C92836D9F245FFC270BCFCDDFC04FC34" blockId="1.[151,1437,831,1974]" pageId="1" pageNumber="132">
<emphasis id="FBE3EACBF245FFC270BCFCDDFEF3FC84" bold="true" box="[151,274,831,858]" pageId="1" pageNumber="132">Diagnosis.</emphasis>
Head with relatively small vertexal and frontal foveae; maxillary palpi elongate and basally pedunculate, length of pedunculate portion successively shorter from palpomeres 2 to 4 (
<figureCitation id="51AC2A5CF245FFC273D6FC81FBB4FCA0" box="[1021,1109,867,894]" captionStart="FIGURE 1" captionStartId="3.[151,250,1443,1468]" captionTargetBox="[160,1436,189,1412]" captionTargetId="figure@3.[151,1436,179,1416]" captionTargetPageId="3" captionText="FIGURE 1. Tibetyrus formicarius (A–C. Male. D. Female). A. Habitus. B. Right maxillary palpus. C, D. Antennomeres 9–11. Scale bars: 1.0 mm in A; 0.2 mm in B; 0.5 mm in C, D." figureDoi="http://doi.org/10.5281/zenodo.3864943" httpUri="https://zenodo.org/record/3864943/files/figure.png" pageId="1" pageNumber="132">Fig. 1B</figureCitation>
). Pronotum with punctiform median- and small lateral antebasal foveae, with thin and shallow antebasal sulcus connecting foveae. Elytra short and strongly constricted at base, each with two small basal foveae. Visible tergites 1–4 (IV–VII) subequal at midlength. Legs with very short tarsomeres 1, and elongate tarsomeres 2 and 3.
</paragraph>
</subSubSection>
<subSubSection id="818D6552F245FFC170ECFC11FDD8FD59" lastPageId="2" lastPageNumber="133" pageId="1" pageNumber="132" type="description">
<paragraph id="C92836D9F245FFC270ECFC11FE3FFAF3" blockId="1.[151,1437,831,1974]" pageId="1" pageNumber="132">
<emphasis id="FBE3EACBF245FFC270ECFC11FEB8FBD0" bold="true" box="[199,345,1011,1038]" pageId="1" pageNumber="132">Description.</emphasis>
Relatively large-sized for pselaphines, body length
<quantity id="0E6F9B3CF245FFC2738AFC11FBCAFBD0" box="[929,1067,1011,1038]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="1" pageNumber="132" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.5–3.7 mm</quantity>
. Head with frontal rostrum moderately long, relatively narrow, formed largely by slightly prominent antennal tubercles; with setose frontal and vertexal foveae; with 11 antennomeres, clubs formed by three apical antennomeres; lacking postantennal notches at lateral margins; lacking lateral postantennal pits, at position where pits occur there are five short and thickened semi-transparent setae; maxillary palpi with small, subtriangular palpomeres 1, palpomeres 2 pedunculate for basal three-fifths, gradually widening and then narrowing again to apices, palpomeres 3 pedunculate for short distance at bases, widening to and truncate at apices, palpomeres 4 enlarged and elongate, suboval, constricted at bases and broadest at basal two-fifths, apices round, lacking apical palpal cones; gula moderately swollen, gular foveae close in shared transverse opening.
</paragraph>
<paragraph id="C92836D9F245FFC270ECFADAFECDFA47" blockId="1.[151,1437,831,1974]" pageId="1" pageNumber="132">Pronotum elongate, broadest at apical third, with disc moderately convex; median antebasal fovea punctiform and nude, lateral antebasal foveae relatively small and setose; lateral procoxal foveae present, lacking paranotal carinae/sulci.</paragraph>
<paragraph id="C92836D9F245FFC270ECFA41FC93FA3F" blockId="1.[151,1437,831,1974]" pageId="1" pageNumber="132">Elytra subtrapezoidal, each elytron with two small but distinct setose basal foveae; broad, shallow discal striae extending posteriorly from outer basal fovea to apical four-fifths.</paragraph>
<paragraph id="C92836D9F245FFC270ECFA09FB4CF993" blockId="1.[151,1437,831,1974]" pageId="1" pageNumber="132">Mesoventrite lacking median mesosventral fovea, with large lateral mesosventral foveae deeply forked internally, anterolateral mesosventral foveae absent; with large lateral mesocoxal foveae; single median metaventral fovea large and densely setose, posterior margin of metaventrite with looped notch at middle.</paragraph>
<paragraph id="C92836D9F245FFC270ECF9BAFDF7F903" blockId="1.[151,1437,831,1974]" pageId="1" pageNumber="132">Legs noticeably elongate, lacking distinct cuticular features; femora moderately thickened, broadest at middle; pro- and mesotibiae slightly curved medially, metatibiae slightly sinuate, all tibiae narrowed at bases and thickened to apices; tarsomeres 1 very short, tarsomeres 2 and 3 elongate, 2 moderately to slightly shorter than 3; pretarsal claws paired and curved at apices.</paragraph>
<paragraph id="C92836D9F245FFC270ECF905FE35F8B3" blockId="1.[151,1437,831,1974]" pageId="1" pageNumber="132">Abdomen with tergites moderately convex, visible tergites 1–4 (IV–VII) subequal at mid-length, tergites 1–3 (IV–VI) each with transverse basal impression, 1–5 (IV–VIII) lacking mediobasal foveae, and each with pair of small basolateral foveae. Visible sternites 2–5 (IV–VII) each with deep basal sulcus, lacking mediobasal foveae, with two basolateral foveae.</paragraph>
<paragraph id="C92836D9F245FFC270ECF895FC54F86B" blockId="1.[151,1437,831,1974]" pageId="1" pageNumber="132">Males with modified antennae and metaventrite. Aedeagus with apical half of median lobe strongly asymmetric; parameres reduced and symmetric; dorsal diaphragm opening oval.</paragraph>
<paragraph id="C92836D9F246FFC170ECFF7AFDD8FD59" blockId="2.[151,1437,152,719]" pageId="2" pageNumber="133">
<emphasis id="FBE3EACBF246FFC170ECFF7AFE4EFF6D" bold="true" box="[199,431,152,179]" pageId="2" pageNumber="133">Comparative notes.</emphasis>
Among the Asian genera of
<taxonomicName id="0E974D5AF246FFC172C3FF7BFCCEFF6D" authorityName="Reitter" authorityYear="1882" box="[744,815,153,179]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="tribe" tribe="Tyrini">Tyrini</taxonomicName>
,
<taxonomicName id="0E974D5AF246FFC17312FF7BFC40FF6D" authority="Yin &amp; Lin, 2020" authorityName="Yin &amp; Lin" authorityYear="2020" box="[825,929,153,179]" class="Insecta" family="Staphylinidae" genus="Tibetyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FBE3EACBF246FFC17312FF7BFC40FF6D" box="[825,929,153,179]" italics="true" pageId="2" pageNumber="133">Tibetyrus</emphasis>
</taxonomicName>
<emphasis id="FBE3EACBF246FFC1738CFF7BFBEEFF6D" bold="true" box="[935,1039,153,179]" pageId="2" pageNumber="133">
<taxonomicNameLabel id="E0D057B0F246FFC1738CFF7BFBEEFF6D" box="[935,1039,153,179]" pageId="2" pageNumber="133" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
shares only with
<taxonomicName id="0E974D5AF246FFC174E4FF78FEEEFF09" authority="Hlavac &amp; Nomura" authorityName="Hlavac &amp; Nomura" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Megatyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="FBE3EACBF246FFC174E4FF78FAA6FF6D" box="[1231,1351,154,179]" italics="true" pageId="2" pageNumber="133">Megatyrus</emphasis>
Hlaváč &amp; Nomura
</taxonomicName>
the basally pedunculate maxillary palpi that lack apical cones on palpomeres 4. However,
<taxonomicName id="0E974D5AF246FFC174D6FF5FFA84FF09" authority="Yin &amp; Lin, 2020" authorityName="Yin &amp; Lin" authorityYear="2020" box="[1277,1381,189,215]" class="Insecta" family="Staphylinidae" genus="Tibetyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FBE3EACBF246FFC174D6FF5FFA84FF09" box="[1277,1381,189,215]" italics="true" pageId="2" pageNumber="133">Tibetyrus</emphasis>
</taxonomicName>
<emphasis id="FBE3EACBF246FFC17540FF5FFF29FF25" bold="true" pageId="2" pageNumber="133">
<taxonomicNameLabel id="E0D057B0F246FFC17540FF5FFF29FF25" pageId="2" pageNumber="133" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="0E974D5AF246FFC17176FF00FE34FF25" authorityName="Hlavac &amp; Nomura" authorityYear="2003" box="[349,469,226,251]" class="Insecta" family="Staphylinidae" genus="Megatyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="FBE3EACBF246FFC17176FF00FE34FF25" box="[349,469,226,251]" italics="true" pageId="2" pageNumber="133">Megatyrus</emphasis>
</taxonomicName>
by numerous characters: 1) area between antennal tubercles much less impressed, not deeply sulcate, 2) much shorter pedunculate portion of maxillary palpomeres 3–4, 3) scapes longer than antennomeres 2–3 combined in male, subequal in length in female, 4) a different shape of the pronotum, 5) strongly reduced elytra and total loss of the hindwings, 6) tergites 1–4 (IV–VII) being subequal at their mid-length, 7) lack of a median carina on tergite 1 (IV), and 8) male sexual characters located on antennal clubs and metaventrite. In contrast, all
<taxonomicName id="0E974D5AF246FFC17096FE74FED4FE71" authorityName="Hlavac &amp; Nomura" authorityYear="2003" box="[189,309,406,431]" class="Insecta" family="Staphylinidae" genus="Megatyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="FBE3EACBF246FFC17096FE74FED4FE71" box="[189,309,406,431]" italics="true" pageId="2" pageNumber="133">Megatyrus</emphasis>
</taxonomicName>
species possess a deep to moderately deep median sulcus on the frons that separates the antennal tubercles, the pedunculate portions of maxillary palpomeres 3–4 are much longer, the scapes are shorter than the following two antennomeres combined in both sexes, the pronotum is broadest at the middle, the elytra have a very broad base, tergite 1 (IV) is as long as 2–3 (V–VI) combined, and with a median carina extends through the entire tergal length, and the male characters are located on the scapes and legs.
<taxonomicName id="0E974D5AF246FFC173E9FDC7FBCBFDE1" authority="Yin &amp; Lin, 2020" authorityName="Yin &amp; Lin" authorityYear="2020" box="[962,1066,549,575]" class="Insecta" family="Staphylinidae" genus="Tibetyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FBE3EACBF246FFC173E9FDC7FBCBFDE1" box="[962,1066,549,575]" italics="true" pageId="2" pageNumber="133">Tibetyrus</emphasis>
</taxonomicName>
<emphasis id="FBE3EACBF246FFC1741BFDC7FB78FDE1" bold="true" box="[1072,1177,549,575]" pageId="2" pageNumber="133">
<taxonomicNameLabel id="E0D057B0F246FFC1741BFDC7FB78FDE1" box="[1072,1177,549,575]" pageId="2" pageNumber="133" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be otherwise separated from all known Asian genera by the form of the maxillary palpi, together with the other external characters as well as the highly distinctive habitus.
</paragraph>
</subSubSection>
<subSubSection id="818D6552F246FFC170ECFD73FE9CFD11" pageId="2" pageNumber="133" type="etymology">
<paragraph id="C92836D9F246FFC170ECFD73FE9CFD11" blockId="2.[151,1437,152,719]" pageId="2" pageNumber="133">
<emphasis id="FBE3EACBF246FFC170ECFD73FEAFFD75" bold="true" box="[199,334,657,683]" pageId="2" pageNumber="133">Etymology.</emphasis>
The new generic name is a combination of ‘Tibet’ and ‘
<taxonomicName id="0E974D5AF246FFC173FDFD70FBF5FD75" authorityName="Aube" authorityYear="1833" box="[982,1044,658,683]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="FBE3EACBF246FFC173FDFD70FBF5FD75" box="[982,1044,658,683]" italics="true" pageId="2" pageNumber="133">Tyrus</emphasis>
</taxonomicName>
Aubé’ (
<typeStatus id="162C887BF246FFC1745BFD73FB43FD75" box="[1136,1186,657,683]" pageId="2" pageNumber="133">type</typeStatus>
genus of
<taxonomicName id="0E974D5AF246FFC1753FFD73FABCFD72" authorityName="Reitter" authorityYear="1882" box="[1300,1373,657,684]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="tribe" tribe="Tyrini">Tyrini</taxonomicName>
). The gender is masculine.
</paragraph>
</subSubSection>
</treatment>
</document>
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