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<document id="95F2C05B4AAD1A0375117C5F19A41B3E" ID-DOI="10.11646/zootaxa.3722.2.1" ID-GBIF-Dataset="f2c46514-f165-43d3-a574-47c55b89dcd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="218998" ID-ZooBank="29D77A16-096D-4FC1-A5B4-9EEDF2E761A7" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460690754500" checkinUser="plazi" docAuthor="Vallejo, Luís Fernando &amp; Wolff, Martha" docDate="2013" docId="3009D414FFF6FFA339B6F993FAC5A3EF" docLanguage="en" docName="zt03722p142.pdf" docOrigin="Zootaxa 3722 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllophaga (Phyllophaga) schizorhina Bates 1888" docType="treatment" docVersion="8" lastPageNumber="118" masterDocId="CC30AC6CFFF9FFB23921FFF1FF91A242" masterDocTitle="The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains" masterLastPageNumber="142" masterPageNumber="101" pageNumber="116" updateTime="1698351836216" updateUser="plazi">
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<mods:title id="DC7545576041E50BA6EF3F2176332989">The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains</mods:title>
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<mods:namePart id="D05F0CC42ADE479E77B62A7797181BFD">Vallejo, Luís Fernando</mods:namePart>
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<mods:namePart id="AB932B9BBD2F08BA7D62750B4EB5DBAF">Wolff, Martha</mods:namePart>
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<mods:date id="9D9362CAC509EECA62B3FF03DD028AFB">2013</mods:date>
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<mods:number id="AA353B5C595383ED41945B893C69B445">3722</mods:number>
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<treatment id="3009D414FFF6FFA339B6F993FAC5A3EF" ID-DOI="http://doi.org/10.5281/zenodo.6151933" ID-Zenodo-Dep="6151933" LSID="urn:lsid:plazi:treatment:3009D414FFF6FFA339B6F993FAC5A3EF" httpUri="http://treatment.plazi.org/id/3009D414FFF6FFA339B6F993FAC5A3EF" lastPageId="17" lastPageNumber="118" pageId="15" pageNumber="116">
<subSubSection id="F0BA3689FFF6FFA339B6F993FF6DA2B5" lastPageId="17" lastPageNumber="118" pageId="15" pageNumber="116" type="key">
<paragraph id="B81F6502FFF6FFBD39B6F993FCDCA43E" blockId="15.[151,845,1634,1660]" box="[151,845,1634,1660]" pageId="15" pageNumber="116">
<heading id="E357D26EFFF6FFBD39B6F993FCDCA43E" bold="true" box="[151,845,1634,1660]" fontSize="11" level="1" pageId="15" pageNumber="116" reason="1">
<keyStep id="03547DA7FFF6FFBD39B6F993FCDCA43E" box="[151,845,1634,1660]" pageId="15" pageNumber="116">
<key id="CC313233FFF6FFBD39B6F993FCDCA43E" box="[151,845,1634,1660]" pageId="15" pageNumber="116">
<emphasis id="8AD4B910FFF6FFBD39B6F993FCDCA43E" bold="true" box="[151,845,1634,1660]" pageId="15" pageNumber="116">
<keyLead id="0351C637FFF6FFBD39B6F993FCDCA43E" box="[151,845,1634,1660]" pageId="15" pageNumber="116">
10.
<taxonomicName id="7FA01E81FFF6FFBD39E2F993FCDCA43E" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[195,845,1634,1660]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="schizorhina" subGenus="Phyllophaga">
<emphasis id="8AD4B910FFF6FFBD39E2F993FECDA43E" bold="true" box="[195,348,1634,1660]" italics="true" pageId="15" pageNumber="116">Phyllophaga</emphasis>
(
<emphasis id="8AD4B910FFF6FFBD384CF993FD97A43E" bold="true" box="[365,518,1634,1660]" italics="true" pageId="15" pageNumber="116">Phyllophaga</emphasis>
)
<emphasis id="8AD4B910FFF6FFBD3B36F993FD34A43E" bold="true" box="[535,677,1634,1660]" italics="true" pageId="15" pageNumber="116">schizorhina</emphasis>
(Bates, 1888)
</taxonomicName>
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</emphasis>
</key>
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</heading>
</paragraph>
<paragraph id="B81F6502FFF6FFA339B6F959FF6DA2B5" blockId="15.[151,1436,1704,2017]" lastBlockId="17.[151,1436,151,429]" lastPageId="17" lastPageNumber="118" pageId="15" pageNumber="116">
<emphasis id="8AD4B910FFF6FFBD39B6F959FED5A483" bold="true" box="[151,324,1704,1729]" pageId="15" pageNumber="116">Redescription.</emphasis>
Length
<quantity id="7F58C8E7FFF6FFBD3885F958FDB3A482" box="[420,546,1705,1729]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="15" pageNumber="116" unit="mm" value="22.5" valueMax="24.0" valueMin="21.0">2124 mm</quantity>
. Body oval, elongate; dorsal surface dark chestnut-brown, punctate. Pronotum (
<figureCitation id="209B7987FFF6FFBD39BEF93DFF64A4A7" box="[159,245,1740,1765]" captionStart="FIGURES 41 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="15" pageNumber="116">Fig. 41</figureCitation>
) is wider than long the dorsal half, lateral margins irregular, crenulate. Clypeus (
<figureCitation id="209B7987FFF6FFBD3D98F93DFA9EA4A7" box="[1209,1295,1740,1765]" captionStart="FIGURES 41 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="15" pageNumber="116">Fig. 41</figureCitation>
) is bilobed, concave, with a raised margin. Antennae (
<figureCitation id="209B7987FFF6FFBD3B53F901FD55A54B" box="[626,708,1776,1801]" captionStart="FIGURES 41 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="15" pageNumber="116">Fig. 42</figureCitation>
) have 10 antennomeres with the antennal club as long as the six preceding antennomeres. Tarsal claws (
<figureCitation id="209B7987FFF6FFBD3B6FF8E4FD0EA56F" box="[590,671,1813,1838]" captionStart="FIGURES 41 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="15" pageNumber="116">Fig. 43</figureCitation>
) have a ventral tooth located next to the basal expansion. Metatibiae are long, wide, and flat (
<figureCitation id="209B7987FFF6FFBD3886F8C9FE68A513" box="[423,505,1848,1873]" captionStart="FIGURES 41 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="15" pageNumber="116">Fig. 44</figureCitation>
) with two typical spurs, of which the inner spur is shorter than the outer spur and is fused with the tibia; similarly, two toothed carinae in the mid and basal portion are present. The pygidial plate has minute setae on its ventral and frontal extremes (
<figureCitation id="209B7987FFF6FFBD3B99F871FCD3A5DB" box="[696,834,1920,1945]" captionStart="FIGURES 41 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="15" pageNumber="116">Figs. 4547</figureCitation>
); also, anterior-distal punctures in the ventral position are present. The male genital structure (
<figureCitation id="209B7987FFF6FFBD3B72F854FD48A5FC" box="[595,729,1957,1982]" captionStart="FIGURES 41 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="15" pageNumber="116">Figs. 4850</figureCitation>
) is easily differentiable, the base of genital capsule has a short, sinuous distal margin and the dorsal basal margin is projected. Parameres are 1.3 times longer than the width of the base, finishing in a rounded apex. The ventral membrane of the genital capsule does not have sclerotized plates (
<figureCitation id="209B7987FFE8FFA339BEFF4DFEBAA296" box="[159,299,188,213]" captionStart="FIGURES 41 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="17" pageNumber="118">Figs. 4849</figureCitation>
); the aedeagus is slightly asymmetric with a narrow, somewhat sclerotized, hook-shaped support at the apex.
</paragraph>
</subSubSection>
<caption id="ECDF358AFFE9FFA239B6F850FB0CA5B6" httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="16" pageNumber="117" targetBox="[232,1365,244,1877]" targetPageId="16">
<paragraph id="B81F6502FFE9FFA239B6F850FB0CA5B6" blockId="16.[151,1436,1953,2036]" pageId="16" pageNumber="117">
<emphasis id="8AD4B910FFE9FFA239B6F850FEC7A5F4" bold="true" box="[151,342,1953,1976]" pageId="16" pageNumber="117">FIGURES 4150.</emphasis>
<taxonomicName id="7FA01E81FFE9FFA2387EF850FDF1A5F5" box="[351,608,1953,1975]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="schizorhina">
<emphasis id="8AD4B910FFE9FFA2387EF850FDF1A5F5" box="[351,608,1953,1975]" italics="true" pageId="16" pageNumber="117">Phyllophaga schizorhina</emphasis>
</taxonomicName>
adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 4850 after Morón 2003) Scale: 1 mm.
</paragraph>
</caption>
<subSubSection id="F0BA3689FFE8FFA339E6FEF5FAC5A3EF" pageId="17" pageNumber="118" type="discussion">
<paragraph id="B81F6502FFE8FFA339E6FEF5FEADA302" blockId="17.[151,1436,151,429]" pageId="17" pageNumber="118">
<emphasis id="8AD4B910FFE8FFA339E6FEF5FEDEA35F" bold="true" box="[199,335,260,285]" pageId="17" pageNumber="118">Comments.</emphasis>
Originally this species was collected in tropical forests of
<collectingCountry id="C0B72592FFE8FFA33ADFFEF5FBE5A35F" box="[1022,1140,260,285]" name="Nicaragua" pageId="17" pageNumber="118">Nicaragua</collectingCountry>
,
<collectingCountry id="C0B72592FFE8FFA33DA3FEF5FA93A35F" box="[1154,1282,260,285]" name="Costa Rica" pageId="17" pageNumber="118">Costa Rica</collectingCountry>
, and
<collectingCountry id="C0B72592FFE8FFA33C62FEF4FA0DA35E" box="[1347,1436,261,284]" name="Panama" pageId="17" pageNumber="118">Panama</collectingCountry>
(Morón 2003).
</paragraph>
<paragraph id="B81F6502FFE8FFA339E6FEBDFAC5A3EF" blockId="17.[151,1436,151,429]" pageId="17" pageNumber="118">
<emphasis id="8AD4B910FFE8FFA339E6FEBDFCF8A327" bold="true" box="[199,873,332,357]" pageId="17" pageNumber="118">New geographic records. Departamento de Risaralda:</emphasis>
Municipio de Quinchía, (
<quantity id="7F58C8E7FFE8FFA33D83FEBDFA96A326" box="[1186,1287,332,357]" metricMagnitude="0" metricUnit="m" metricValue="1.825" pageId="17" pageNumber="118" unit="m" value="1.825">1,825 m</quantity>
)
<date id="CC1E43C2FFE8FFA33C3DFEBCFA06A326" box="[1308,1431,332,357]" pageId="17" pageNumber="118" value="2007-05">May 2007</date>
, Octavio Útima, cafetal,
<specimenCount id="AEA6AE8BFFE8FFA33889FE81FD92A3CA" box="[424,515,367,392]" pageId="17" pageNumber="118" type="male">2 males</specimenCount>
,
<specimenCount id="AEA6AE8BFFE8FFA33B31FE81FDE6A3CA" box="[528,631,367,392]" pageId="17" pageNumber="118" type="female">1 female</specimenCount>
.
<emphasis id="8AD4B910FFE8FFA33BA2FE9EFC29A3CA" bold="true" box="[643,952,367,392]" pageId="17" pageNumber="118">Departamento de Caldas:</emphasis>
Municipio de Manizales, Granja
<taxonomicName id="7FA01E81FFE8FFA33C1BFE9EFA09A3CA" box="[1338,1432,367,392]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="131" phylum="Arthropoda" rank="species" species="tesorito">Tesorito</taxonomicName>
, (
<quantity id="7F58C8E7FFE8FFA339BFFE65FE91A3EE" box="[158,256,404,428]" metricMagnitude="0" metricUnit="m" metricValue="2.28" pageId="17" pageNumber="118" unit="m" value="2.28">2,280 m</quantity>
).
<date id="CC1E43C2FFE8FFA33832FE65FE50A3EF" box="[275,449,404,429]" pageId="17" pageNumber="118" value="2006-03-17">17 March 2006</date>
, en arboloco (
<taxonomicName id="7FA01E81FFE8FFA33B41FE64FC19A3EF" box="[608,904,404,429]" class="Magnoliopsida" family="Asteraceae" genus="Montanoa" kingdom="Plantae" order="Asterales" pageId="17" pageNumber="118" phylum="Tracheophyta" rank="species" species="quadrangularis">
<emphasis id="8AD4B910FFE8FFA33B41FE64FC19A3EF" box="[608,904,404,429]" italics="true" pageId="17" pageNumber="118">Montanoa quadrangularis</emphasis>
</taxonomicName>
), Fernando Vallejo,
<specimenCount id="AEA6AE8BFFE8FFA33D4FFE65FB44A3EF" box="[1134,1237,404,429]" pageId="17" pageNumber="118" type="male">12 males</specimenCount>
,
<specimenCount id="AEA6AE8BFFE8FFA33DC1FE64FAC1A3EF" box="[1248,1360,404,429]" pageId="17" pageNumber="118" type="female">7 females</specimenCount>
.
</paragraph>
</subSubSection>
</treatment>
</document>