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<document id="D22B48BB14A25B8DED7BBE7F9977B65F" ID-DOI="10.5281/zenodo.3673491" ID-GBIF-Dataset="d8f38b1a-d989-4559-8f58-95878de25602" ID-ISSN="1942-1354" ID-Zenodo-Dep="3673491" ID-ZooBank="B2EA9141-6419-4A1A-85CC-BCE190C17802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582027722378" checkinUser="felipe" docAuthor="Harris, Steven C. &amp; Armitage, Brian J." docDate="2019" docId="17184D65FFA8650952BEFED1FA187EB7" docLanguage="en" docName="InsectaMundi.0707.1-54.pdf.imf" docOrigin="Insecta Mundi 707 (707)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Alisotrichia bernali Harris and Armitage 2019, new species" docType="treatment" docVersion="11" lastPageNumber="8" masterDocId="EB21351DFFA16500527EFFFCFF967A60" masterDocTitle="The Trichoptera of Panama X. The Quebrada Rambala drainage, with description of 19 new species of microcaddisflies (Trichoptera: Hydroptilidae)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="8" updateTime="1698718896375" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="19A1C82D3450EE6E2FA03980FF116BDF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="582E45C38B6430DBF8DAB443AC4B8C42">The Trichoptera of Panama X. The Quebrada Rambala drainage, with description of 19 new species of microcaddisflies (Trichoptera: Hydroptilidae)</mods:title>
</mods:titleInfo>
<mods:name id="CFE7A2248BA4B83D2025949F86D93F47" type="personal">
<mods:role id="9D4772DA88E7550F8BC97507BECCCFCC">
<mods:roleTerm id="59FAA61112938EBA0D052741C9B6A6B6">Author</mods:roleTerm>
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<mods:namePart id="00F0417C23A14E90D7CEA39D394039AB">Harris, Steven C.</mods:namePart>
<mods:affiliation id="25D713C554111D5FFCEACFDEBB2ACC66">Department of Biology and Geosciences Clarion University Clarion, PA 16214 USA</mods:affiliation>
<mods:nameIdentifier id="358B02E9A4A52FBD6D16169BBAD07DF6" type="email">sharris@clarion.edu</mods:nameIdentifier>
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<mods:name id="936C6E2EF3C03D776DBDCDD868E4B368" type="personal">
<mods:role id="3ACC8C884FC67CD72F0EC1E62721D1C6">
<mods:roleTerm id="86D52A9AE854BEB7A28017C07724E16B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E694ADB05522948EC803206694B12506">Armitage, Brian J.</mods:namePart>
<mods:affiliation id="8E19CB5446869DEB6771A503D51AE6DB">Museo de Peces de Agua Dulce e Invertebrados (MUPADI) Universidad Autónoma de Chiriquí (UNACHI) David, Republic of Panama</mods:affiliation>
<mods:nameIdentifier id="A8CAF76078BF97129305A9D5BE7DBF13" type="email">tobikera89@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="2C5C24AFF0A086A95EDEA4258B1C5E38">text</mods:typeOfResource>
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<mods:titleInfo id="8F802D6A7793690B6BAA0EB2BFAEBBBC">
<mods:title id="CDDB43FD9E930CCC3DA1EC88D28C0130">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="353AFB81BAAD2C670E8D4BCB505D863A">
<mods:date id="D27384E82A0086BDDA9367D41AE4094D">2019</mods:date>
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<mods:number id="2549C6FB7FE14CAF036062C5A64514E4">2019-05-31</mods:number>
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<mods:number id="DF360F1A4CBE526961B2FB9431AC645B">707</mods:number>
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<mods:number id="B81E41BA065C90729555473D667C4D3A">707</mods:number>
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<mods:classification id="AF8F7582784189CF903799CEB8F91D8C">journal article</mods:classification>
<mods:identifier id="376FFCEE3D9966A4862A45888B8BEE61" type="DOI">10.5281/zenodo.3673491</mods:identifier>
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<mods:identifier id="3DDB7018A0F035DD6975210BC5FF6E2C" type="ISSN">1942-1354</mods:identifier>
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<treatment id="17184D65FFA8650952BEFED1FA187EB7" ID-DOI="http://doi.org/10.5281/zenodo.3679991" ID-GBIF-Taxon="162161895" ID-Zenodo-Dep="3679991" LSID="urn:lsid:plazi:treatment:17184D65FFA8650952BEFED1FA187EB7" httpUri="http://treatment.plazi.org/id/17184D65FFA8650952BEFED1FA187EB7" lastPageNumber="8" pageId="9" pageNumber="8">
<subSubSection id="D7ABAFF8FFA8650952BEFED1FC317B25" box="[192,935,301,325]" pageId="9" pageNumber="8" type="nomenclature">
<paragraph id="9F0EFC73FFA8650952BEFED1FC317B25" blockId="9.[192,935,301,358]" box="[192,935,301,325]" pageId="9" pageNumber="8">
<heading id="C4464B1FFFA8650952BEFED1FC317B25" bold="true" box="[192,935,301,325]" fontSize="10" level="2" pageId="9" pageNumber="8" reason="3">
<emphasis id="ADC52061FFA8650952BEFED1FC317B25" bold="true" box="[192,935,301,325]" pageId="9" pageNumber="8">
<taxonomicName id="58B187F0FFA8650952BEFED1FD607B25" authority="Harris and Armitage" authorityName="Harris and Armitage" authorityYear="2019" box="[192,758,301,325]" class="Insecta" family="Hydroptilidae" genus="Alisotrichia" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="bernali" status="sp. nov.">
<emphasis id="ADC52061FFA8650952BEFED1FE597B25" bold="true" box="[192,463,301,325]" italics="true" pageId="9" pageNumber="8">Alisotrichia bernali</emphasis>
Harris and Armitage
</taxonomicName>
,
<taxonomicNameLabel id="B6F69D1AFFA86509517DFED1FC317B25" box="[771,935,301,325]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFA8650952BEFEB2FE907B05" box="[192,262,333,358]" pageId="9" pageNumber="8" type="description">
<paragraph id="9F0EFC73FFA8650952BEFEB2FE907B05" blockId="9.[192,935,301,358]" box="[192,262,333,358]" pageId="9" pageNumber="8">
<figureCitation id="078AE0F6FFA8650952BEFEB2FE907B05" box="[192,262,333,358]" captionStart="Figure 5" captionStartId="29.[192,275,1848,1870]" captionTargetBox="[192,1440,230,1787]" captionTargetId="figure@29.[192,1440,230,1787]" captionTargetPageId="29" captionText="Figure 5. Alisotrichia bernali, sp. n., Male genitalia. A) Lateral view. B) Ventral view. C) Dorsal view. D) Phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3673501" httpUri="https://zenodo.org/record/3673501/files/figure.png" pageId="9" pageNumber="8">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFA8650952BEFE81FC277856" pageId="9" pageNumber="8" type="diagnosis">
<paragraph id="9F0EFC73FFA8650952BEFE81FC277856" blockId="9.[192,1440,381,566]" pageId="9" pageNumber="8">
<emphasis id="ADC52061FFA8650952BEFE81FED97BF5" bold="true" box="[192,335,381,405]" pageId="9" pageNumber="8">Diagnosis.</emphasis>
<taxonomicName id="58B187F0FFA86509532AFE81FDD67BF5" authorityName="Harris and Armitage" authorityYear="2019" box="[340,576,381,405]" class="Insecta" family="Hydroptilidae" genus="Alisotrichia" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="bernali">
<emphasis id="ADC52061FFA86509532AFE81FDD67BF5" box="[340,576,381,405]" italics="true" pageId="9" pageNumber="8">Alisotrichia bernali</emphasis>
</taxonomicName>
appears to be most similar to
<taxonomicName id="58B187F0FFA8650951D8FE82FBC37BF6" authority="Flint" authorityName="Flint" box="[934,1109,381,406]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="lobata">
<emphasis id="ADC52061FFA8650951D8FE82FB857BF5" box="[934,1043,381,405]" italics="true" pageId="9" pageNumber="8">A. lobata</emphasis>
Flint
</taxonomicName>
and
<taxonomicName id="58B187F0FFA8650956F0FE82FAC97BF6" authority="Flint" authorityName="Flint" authorityYear="1992" box="[1166,1375,382,406]" class="Insecta" family="Hydroptilidae" genus="Alisotrichia" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="circinata">
<emphasis id="ADC52061FFA8650956F0FE82FA8C7BF5" box="[1166,1306,382,405]" italics="true" pageId="9" pageNumber="8">A. circinata</emphasis>
Flint
</taxonomicName>
, both of which have abdominal segment VIII narrowing distally to an acute point and a tubular, thin phallus. The new species differs in that the acute apex of segment VIII is bifid and the phallus apex is widened. In
<taxonomicName id="58B187F0FFA86509529DFE22FDAF7B96" authority="Harris and Flint" authorityName="Harris and Flint" box="[227,569,477,502]" class="Insecta" family="Hydroptilidae" genus="Alisotrichia" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mathisi">
<emphasis id="ADC52061FFA86509529DFE22FEF27B95" box="[227,356,477,501]" italics="true" pageId="9" pageNumber="8">A. mathisi</emphasis>
Harris and Flint
</taxonomicName>
there are a pair of spines at the apex of segment VIII which resemble the structure of segment VIII in the new species, but the dorsal view of
<taxonomicName id="58B187F0FFA865095649FE02FB217876" box="[1079,1207,510,534]" class="Insecta" family="Hydroptilidae" genus="Alisotrichia" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="mathisi">
<emphasis id="ADC52061FFA865095649FE02FB217876" box="[1079,1207,510,534]" italics="true" pageId="9" pageNumber="8">A. mathisi</emphasis>
</taxonomicName>
shows clearly that these are spines and not a bifid apical portion of the segment.
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFA8650952BEFDB2FE257E77" pageId="9" pageNumber="8" type="description">
<paragraph id="9F0EFC73FFA8650952BEFDB2FB307986" blockId="9.[192,1441,590,999]" pageId="9" pageNumber="8">
<emphasis id="ADC52061FFA8650952BEFDB2FE9C7806" bold="true" box="[192,266,590,614]" pageId="9" pageNumber="8">Male.</emphasis>
Length
<quantity id="58495196FFA86509530CFDB2FE447806" box="[370,466,590,614]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="8" unit="mm" value="1.7">1.7 mm</quantity>
. Antenna 18 segments, scape slightly enlarged, remaining segments spatulate. Abdominal segment VI annular, with bifid, ventromesal process posteriorly. Segment VII thin ventrally, widening dorsally; emarginate posteriorly in dorsal view. Segment VIII elongate and narrowing distally to bifid apex, ventrally thin, narrowing anteriorly: dorsally and ventrally truncate, deeply incised posteriorly forming lateral bifid processes, mesally with pair of rounded humps; ventrally elongate with mesal humps posteriorly and bifid laterally. Segment IX entirely enclosed within VIII, anteriorly sharply narrowed into elongate, thin apodeme which extends into segment V, posteriorly divided into dorsal process bearing elongate, thickened seta, and a ventral process which is narrow and projects ventrad to an acute apex; ventrally with narrow anterior apodemes, posteriorly squarish, with sclerotized, slightly produced lateral margins, and truncate distally; dorsally narrow laterally with posterior inner lobes bearing elongate thickened seta, mesally with pair of thin sclerites. Segment X shelf-like in lateral view, rounded distally; in dorsal view generally rectanguloid, tapering to truncate apex. Phallus thin and tubular, widening basally, distally abruptly widening to oval, spatulate apex.
</paragraph>
<paragraph id="9F0EFC73FFA8650952BEFC03FE257E77" blockId="9.[192,435,1023,1047]" box="[192,435,1023,1047]" pageId="9" pageNumber="8">
<emphasis id="ADC52061FFA8650952BEFC03FEBA7E77" bold="true" box="[192,300,1023,1047]" pageId="9" pageNumber="8">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFA8650952BEFBD3FDC07EE7" pageId="9" pageNumber="8" type="materials_examined">
<paragraph id="9F0EFC73FFA8650952BEFBD3FDC07EE7" blockId="9.[192,1440,1071,1159]" pageId="9" pageNumber="8">
<emphasis id="ADC52061FFA8650952BEFBD3FE1D7E27" bold="true" box="[192,395,1071,1095]" pageId="9" pageNumber="8">Type material.</emphasis>
<materialsCitation id="2FD9F62EFFA8650953EAFBD3FDC07EE7" ID-GBIF-Occurrence="2565952432" collectingDate="2017-02-06" collectingDateMax="2017-02-12" collectingDateMin="2017-02-06" collectorName="E. Carlson." country="Panama" county="Quebrada Rambala" elevation="120" latitude="8.91543" location="Rambala Jungle Lodge" longLatPrecision="1" longitude="-82.15527" municipality="Rambala" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Bocas del Toro" typeStatus="holotype">
<emphasis id="ADC52061FFA8650953EAFBD3FBC07E27" bold="true" box="[404,1110,1071,1095]" pageId="9" pageNumber="8">
<typeStatus id="400A42D1FFA8650953EAFBD3FD857E27" box="[404,531,1071,1095]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
, male—
<collectingCountry id="E7A6BCE3FFA865095001FBD3FD677E27" box="[639,753,1071,1095]" name="Panama" pageId="9" pageNumber="8">Panama</collectingCountry>
,
<collectingRegion id="5D753291FFA86509517EFBD3FBC07E27" box="[768,1110,1071,1095]" country="Panama" name="Bocas del Toro" pageId="9" pageNumber="8">Bocas del Toro Province</collectingRegion>
</emphasis>
,
<collectingCounty id="766F84FFFFA865095619FBD3FACC7E27" box="[1127,1370,1071,1095]" pageId="9" pageNumber="8">
Quebrada
<collectingMunicipality id="7F6A6609FFA865095697FBD3FACC7E27" box="[1257,1370,1071,1095]" pageId="9" pageNumber="8">Rambala</collectingMunicipality>
</collectingCounty>
, near
<location id="9A6EAAA8FFA8650952BEFBB3FE707E07" LSID="urn:lsid:plazi:treatment:17184D65FFA8650952BEFED1FA187EB7:9A6EAAA8FFA8650952BEFBB3FE707E07" box="[192,486,1103,1127]" country="Panama" county="Quebrada Rambala" latitude="8.91543" longLatPrecision="1" longitude="-82.15527" municipality="Rambala" name="Rambala Jungle Lodge" pageId="9" pageNumber="8" stateProvince="Bocas del Toro">
<location id="9A6EAAA8FFA8650952BEFBB3FEB97E07" LSID="urn:lsid:plazi:treatment:17184D65FFA8650952BEFED1FA187EB7:9A6EAAA8FFA8650952BEFBB3FEB97E07" box="[192,303,1103,1127]" country="Panama" county="Quebrada Rambala" latitude="8.91543" longLatPrecision="1" longitude="-82.15527" municipality="Rambala" name="Rambala" pageId="9" pageNumber="8" stateProvince="Bocas del Toro">Rambala</location>
Jungle Lodge
</location>
,
<locationDeviation id="8EBCE551FFA865095388FBB3FC8C7E07" box="[502,794,1103,1127]" location="Rambala" pageId="9" pageNumber="8">
<quantity id="58495196FFA865095388FBB3FDCB7E07" box="[502,605,1103,1127]" metricMagnitude="3" metricUnit="m" metricValue="3.74" pageId="9" pageNumber="8" unit="km" value="3.74">3.74 km</quantity>
SSE Rambala
</locationDeviation>
,
<geoCoordinate id="FA859AB4FFA865095157FBB3FC3C7E07" box="[809,938,1103,1127]" degrees="8.91543" direction="north" orientation="latitude" pageId="9" pageNumber="8" precision="1" value="8.91543">8.91543°N</geoCoordinate>
and
<geoCoordinate id="FA859AB4FFA865095191FBB3FB107E07" box="[1007,1158,1103,1127]" degrees="82.15527" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="1" value="-82.15527">82.15527°W</geoCoordinate>
,
<quantity id="58495196FFA8650956E8FBB3FB717E07" box="[1174,1255,1103,1127]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="8" unit="m" value="120.0">
<elevation id="149C1B40FFA8650956E8FBB3FB717E07" box="[1174,1255,1103,1127]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="8" unit="m" value="120.0">120 m</elevation>
</quantity>
,
<collectingMethod id="46F08464FFA865095686FBB3FA0D7E07" box="[1272,1435,1103,1127]" pageId="9" pageNumber="8">Malaise trap</collectingMethod>
,
<date id="EB0FDAB3FFA8650952BEFB93FE577EE7" box="[192,449,1135,1159]" pageId="9" pageNumber="8" value="2017-02-06" valueMax="2017-02-12" valueMin="2017-02-06">
<collectingDate id="FB4B235BFFA8650952BEFB93FE577EE7" box="[192,449,1135,1159]" pageId="9" pageNumber="8" value="2017-02-06" valueMax="2017-02-12" valueMin="2017-02-06">February 612, 2017</collectingDate>
</date>
,
<collectorName id="324499A5FFA8650953B3FB8CFDC07EE7" box="[461,598,1135,1159]" pageId="9" pageNumber="8">E. Carlson.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFA8650952BEFB63FA187EB7" pageId="9" pageNumber="8" type="etymology">
<paragraph id="9F0EFC73FFA8650952BEFB63FA187EB7" blockId="9.[192,1439,1183,1240]" pageId="9" pageNumber="8">
<emphasis id="ADC52061FFA8650952BEFB63FECE7ED7" bold="true" box="[192,344,1183,1207]" pageId="9" pageNumber="8">Etymology.</emphasis>
With respect and sadness, we name this species after Dr. Juan Bernal Vega of the University of
<collectingRegion id="5D753291FFA8650952A3FB43FED27EB7" box="[221,324,1215,1239]" country="Panama" name="Chiriqui" pageId="9" pageNumber="8">Chiriqui</collectingRegion>
(UNACHI) who passed away very recently while on a field trip. He will be deeply missed.
</paragraph>
</subSubSection>
</treatment>
</document>