581 lines
64 KiB
XML
581 lines
64 KiB
XML
<document id="678A304B44ABF250147DECFEF9D10EBA" ID-CLB-Dataset="46017" ID-DOI="10.11646/zootaxa.3786.2.4" ID-GBIF-Dataset="a4d5783e-d852-4210-b8b0-27972739baf4" ID-ISSN="1175-5326" ID-Zenodo-Dep="253006" ID-ZooBank="06D44DCE-0816-459A-A201-171F273BC210" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460408271099" checkinUser="plazi" docAuthor="Thomas, Travis M., Granatosky, Michael C., Bourque, Jason R., Krysko, Kenneth L., Moler, Paul E., Gamble, Tony, Suarez, Eric, Leone, Erin, Enge, Kevin M. & Roman, Joe" docDate="2014" docId="03D787A6FFC6FFCDFF34F89292AFC682" docLanguage="en" docName="zt03786p165.pdf" docOrigin="Zootaxa 3786 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrochelys apalachicolae Thomas, Granatosky, Bourque, Krysko, Moler, Gamble, Suarez, Leone, Enge & Roman, 2014, sp. nov." docType="treatment" docUuid="3C044843-F221-4384-A480-6D6CE3975FEC" docUuidSource="ZooBank" docVersion="10" lastPageNumber="160" masterDocId="FFEEFFDEFFCCFFDEFFA3FFB79258C062" masterDocTitle="Taxonomic assessment of Alligator Snapping Turtles (Chelydridae: Macrochelys), with the description of two new species from the southeastern United States" masterLastPageNumber="165" masterPageNumber="141" pageNumber="151" updateTime="1698300110313" updateUser="ExternalLinkService">
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<mods:title id="419AEEC6B15066394CE798DF693CD1E3">Taxonomic assessment of Alligator Snapping Turtles (Chelydridae: Macrochelys), with the description of two new species from the southeastern United States</mods:title>
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<mods:namePart id="1611823BA1E32A541E31B24716A48E06">Thomas, Travis M.</mods:namePart>
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<mods:namePart id="1E73BA86C553DB731EA63BFEAC50F506">Granatosky, Michael C.</mods:namePart>
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<mods:namePart id="376891E2AA1329110911248685656194">Bourque, Jason R.</mods:namePart>
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<mods:namePart id="C5437728AD550F935778910C6C7D8CD1">Krysko, Kenneth L.</mods:namePart>
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<mods:namePart id="E0DC4296E474766D23A00D4E71B1AC91">Moler, Paul E.</mods:namePart>
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<mods:namePart id="F969FBAC035C873FCC7E9C978DA5DA9E">Gamble, Tony</mods:namePart>
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<mods:namePart id="6B59EFF6BFF83DFCE0B52BAE919044F8">Suarez, Eric</mods:namePart>
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<mods:namePart id="8C2E9AE23298B209CE13E3C3E448A718">Leone, Erin</mods:namePart>
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<mods:namePart id="AB55DE92B35D985DD605DE6A8C4C8C32">Enge, Kevin M.</mods:namePart>
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<mods:namePart id="88040DABB4A47EABC714BA8506C9DA76">Roman, Joe</mods:namePart>
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<mods:title id="BC10462A1A6D3F79FB60923B6BAC2756">Zootaxa</mods:title>
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<mods:date id="0B1431992EDB9B5E7B53BFC94C1AC941">2014</mods:date>
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<mods:number id="9BC99D378EC72F4B1C19D1E85EA754C0">3786</mods:number>
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<mods:identifier id="C5FF8D460F326DB94AB5064B79C00609" type="DOI">10.11646/zootaxa.3786.2.4</mods:identifier>
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<treatment id="03D787A6FFC6FFCDFF34F89292AFC682" ID-DOI="http://doi.org/10.5281/zenodo.6141536" ID-GBIF-Taxon="119422505" ID-Zenodo-Dep="6141536" LSID="urn:lsid:zoobank.org:act:3C044843-F221-4384-A480-6D6CE3975FEC" httpUri="http://treatment.plazi.org/id/03D787A6FFC6FFCDFF34F89292AFC682" lastPageId="19" lastPageNumber="160" pageId="10" pageNumber="151">
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<subSubSection id="C364653BFFC6FFD4FF34F892914EC7E7" pageId="10" pageNumber="151" type="nomenclature">
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<paragraph id="8BC136B0FFC6FFD4FF34F892901EC722" blockId="10.[151,582,1829,1856]" box="[151,582,1829,1856]" pageId="10" pageNumber="151">
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<heading id="D08981DCFFC6FFD4FF34F892901EC722" bold="true" box="[151,582,1829,1856]" fontSize="11" level="1" pageId="10" pageNumber="151" reason="1">
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<emphasis id="B90AEAA2FFC6FFD4FF34F892901EC722" bold="true" box="[151,582,1829,1856]" pageId="10" pageNumber="151">
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<taxonomicName id="4C7E4D33FFC6FFD4FF34F89293B8C75D" box="[151,480,1829,1855]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="151" phylum="Chordata" rank="species" species="apalachicolae" status="sp. nov.">
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<emphasis id="B90AEAA2FFC6FFD4FF34F89293B8C75D" bold="true" box="[151,480,1829,1855]" italics="true" pageId="10" pageNumber="151">Macrochelys apalachicolae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23957D9FFC6FFD4FE4BF891901EC722" box="[488,582,1830,1856]" pageId="10" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC136B0FFC6FFD4FF34F8DB914EC7E7" blockId="10.[151,1436,1900,1998]" box="[151,790,1900,1925]" pageId="10" pageNumber="151">
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<emphasis id="B90AEAA2FFC6FFD4FF34F8DB930DC7E7" bold="true" box="[151,341,1900,1925]" pageId="10" pageNumber="151">Common name.</emphasis>
|
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Apalachicola Alligator Snapping Turtle
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</paragraph>
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</subSubSection>
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<subSubSection id="C364653BFFC6FFD0FF64F827961CC629" lastPageId="14" lastPageNumber="155" pageId="10" pageNumber="151" type="materials_examined">
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<paragraph id="8BC136B0FFC6FFD4FF64F82796CDC7AF" blockId="10.[151,1436,1900,1998]" pageId="10" pageNumber="151">
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<emphasis id="B90AEAA2FFC6FFD4FF64F8279363C7CB" bold="true" box="[199,315,1936,1961]" pageId="10" pageNumber="151">
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<typeStatus id="54C58812FFC6FFD4FF64F827936EC7CB" box="[199,310,1936,1961]" pageId="10" pageNumber="151" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
UF 3998, partial skeleton from the Apalachicola River, Gadsden County, Florida, on
|
||
<date id="FFC01070FFC6FFD4FAA8F82697C4C7CB" box="[1291,1436,1936,1961]" pageId="10" pageNumber="151" value="1953-04-04">4 April 1953</date>
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||
by the Florida Museum of Natural History (see
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<figureCitation id="13452A35FFC6FFD4FD0FF802917DC7AF" box="[684,805,1973,1998]" captionStart="FIGURE 13" captionStartId="16.[151,250,950,972]" captionTargetBox="[169,1420,208,913]" captionTargetId="figure@16.[151,1436,193,928]" captionTargetPageId="16" captionText="FIGURE 13. Photograph of Macrochelys apalachicolae holotype (UF 3998) demonstrating external (A) and internal (B) carapace morphology." httpUri="https://zenodo.org/record/253021/files/figure.png" pageId="10" pageNumber="151">Figures 13</figureCitation>
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,
|
||
<figureCitation id="13452A35FFC6FFD4FC92F8029108C7AF" box="[817,848,1973,1997]" captionStart="FIGURE 14" captionStartId="17.[151,250,1874,1896]" captionTargetBox="[166,1413,278,1845]" captionTargetId="figure@17.[151,1436,265,1852]" captionTargetPageId="17" captionText="FIGURE 14. Photograph of Macrochelys apalachicolae holotype (UF 3998) demonstrating a superior (A), inferior (B), cranial (C), caudal (D), and left (E) and right (F) lateral view of skull morphology." httpUri="https://zenodo.org/record/253022/files/figure.png" pageId="10" pageNumber="151">14</figureCitation>
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). (Central lineage;
|
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<figureCitation id="13452A35FFC6FFD4FB84F80296DFC7AF" box="[1063,1159,1973,1998]" captionStart="FIGURE 9" captionStartId="11.[151,250,1743,1765]" captionTargetBox="[159,1435,206,1703]" captionTargetId="figure@11.[151,1436,193,1725]" captionTargetPageId="11" captionText="FIGURE 9. Bayesian inference phylogeny for extant chelydrids (Chelydra and Macrochelys). Note that representative skeletal synapomorphies in skull and carapace (above and below, respectively, next to lineage name) are provided for each lineage of Macrochelys; values above major nodes represent posterior probabilities (≥ 95 %); values below major nodes represent the mean divergence time estimation of the most recent common ancestor (MRCA); and bars at major nodes represent 95 % Highest Posterior Density (HPD)." httpUri="https://zenodo.org/record/253015/files/figure.png" pageId="10" pageNumber="151">Figure 9</figureCitation>
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).
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</paragraph>
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<caption id="DF016638FFC7FFD5FF34F97893B5C702" httpUri="https://zenodo.org/record/253015/files/figure.png" pageId="11" pageNumber="152" targetBox="[159,1435,206,1703]" targetPageId="11">
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<paragraph id="8BC136B0FFC7FFD5FF34F97893B5C702" blockId="11.[151,1435,1743,1888]" pageId="11" pageNumber="152">
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<emphasis id="B90AEAA2FFC7FFD5FF34F978934BC686" bold="true" box="[151,275,1743,1765]" pageId="11" pageNumber="152">FIGURE 9.</emphasis>
|
||
Bayesian inference phylogeny for extant chelydrids (
|
||
<taxonomicName id="4C7E4D33FFC7FFD5FC92F97891D7C687" box="[817,911,1743,1765]" class="Reptilia" family="Chelydridae" genus="Chelydra" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="152" phylum="Chordata" rank="genus">
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<emphasis id="B90AEAA2FFC7FFD5FC92F97891D7C687" box="[817,911,1743,1765]" italics="true" pageId="11" pageNumber="152">Chelydra</emphasis>
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</taxonomicName>
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and
|
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<taxonomicName id="4C7E4D33FFC7FFD5FC63F978961AC687" box="[960,1090,1743,1765]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="152" phylum="Chordata" rank="genus">
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<emphasis id="B90AEAA2FFC7FFD5FC63F978961AC687" box="[960,1090,1743,1765]" italics="true" pageId="11" pageNumber="152">Macrochelys</emphasis>
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||
</taxonomicName>
|
||
). Note that representative skeletal synapomorphies in skull and carapace (above and below, respectively, next to lineage name) are provided for each lineage of
|
||
<taxonomicName id="4C7E4D33FFC7FFD5FF34F8BB9341C740" box="[151,281,1804,1826]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="152" phylum="Chordata" rank="genus">
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<emphasis id="B90AEAA2FFC7FFD5FF34F8BB9341C740" box="[151,281,1804,1826]" italics="true" pageId="11" pageNumber="152">Macrochelys</emphasis>
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</taxonomicName>
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||
; values above major nodes represent posterior probabilities (≥ 95%); values below major nodes represent the mean divergence time estimation of the most recent common ancestor (MRCA); and bars at major nodes represent 95% Highest Posterior Density (HPD).
|
||
</paragraph>
|
||
</caption>
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<caption id="DF016638FFC0FFD2FF34F8AE93CBC74D" box="[151,403,1817,1839]" httpUri="https://zenodo.org/record/253016/files/figure.png" pageId="12" pageNumber="153" targetBox="[160,1436,205,1754]" targetPageId="12">
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<paragraph id="8BC136B0FFC0FFD2FF34F8AE93CBC74D" blockId="12.[151,403,1817,1839]" box="[151,403,1817,1839]" pageId="12" pageNumber="153">
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<emphasis id="B90AEAA2FFC0FFD2FF34F8AE934BC74C" bold="true" box="[151,275,1817,1839]" pageId="12" pageNumber="153">FIGURE 9.</emphasis>
|
||
(Continued)
|
||
</paragraph>
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||
</caption>
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||
<caption id="DF016638FFC1FFD3FF34FC2C93CBC3D3" box="[151,403,923,945]" httpUri="https://zenodo.org/record/253017/files/figure.png" pageId="13" pageNumber="154" targetBox="[162,1436,212,865]" targetPageId="13">
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<paragraph id="8BC136B0FFC1FFD3FF34FC2C93CBC3D3" blockId="13.[151,403,923,945]" box="[151,403,923,945]" pageId="13" pageNumber="154">
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||
<emphasis id="B90AEAA2FFC1FFD3FF34FC2C934BC3D2" bold="true" box="[151,275,923,945]" pageId="13" pageNumber="154">FIGURE 9.</emphasis>
|
||
(Continued)
|
||
</paragraph>
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</caption>
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<caption id="DF016638FFC1FFD3FF34F94C932FC752" httpUri="https://zenodo.org/record/253018/files/figure.png" pageId="13" pageNumber="154" targetBox="[159,1429,999,1761]" targetPageId="13">
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<paragraph id="8BC136B0FFC1FFD3FF34F94C932FC752" blockId="13.[151,1436,1787,1840]" pageId="13" pageNumber="154">
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<emphasis id="B90AEAA2FFC1FFD3FF34F94C937BC772" bold="true" box="[151,291,1787,1809]" pageId="13" pageNumber="154">FIGURE 10.</emphasis>
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||
Photograph of
|
||
<taxonomicName id="4C7E4D33FFC1FFD3FE6BF94C9082C773" box="[456,730,1787,1809]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="154" phylum="Chordata" rank="species" species="suwanniensis">
|
||
<emphasis id="B90AEAA2FFC1FFD3FE6BF94C9082C773" box="[456,730,1787,1809]" italics="true" pageId="13" pageNumber="154">Macrochelys suwanniensis</emphasis>
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||
</taxonomicName>
|
||
holotype (UF 166146) demonstrating external (A) and internal (B) carapace morphology.
|
||
</paragraph>
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</caption>
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<caption id="DF016638FFC2FFD0FF34FA3C9340C5A2" httpUri="https://zenodo.org/record/253019/files/figure.png" pageId="14" pageNumber="155" targetBox="[151,1436,176,1397]" targetPageId="14">
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<paragraph id="8BC136B0FFC2FFD0FF34FA3C9340C5A2" blockId="14.[151,1436,1419,1472]" pageId="14" pageNumber="155">
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||
<emphasis id="B90AEAA2FFC2FFD0FF34FA3C937BC5C2" bold="true" box="[151,291,1419,1441]" pageId="14" pageNumber="155">FIGURE 11.</emphasis>
|
||
Photograph of
|
||
<taxonomicName id="4C7E4D33FFC2FFD0FE6FFA3C9087C5C3" box="[460,735,1419,1441]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="155" phylum="Chordata" rank="species" species="suwanniensis">
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<emphasis id="B90AEAA2FFC2FFD0FE6FFA3C9087C5C3" box="[460,735,1419,1441]" italics="true" pageId="14" pageNumber="155">Macrochelys suwanniensis</emphasis>
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||
</taxonomicName>
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||
holotype (UF 166146) demonstrating a superior view of plastron morphology.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136B0FFC2FFD0FF64FA50961CC629" blockId="14.[151,1436,1511,2007]" pageId="14" pageNumber="155">
|
||
<emphasis id="B90AEAA2FFC2FFD0FF64FA50931EC662" bold="true" box="[199,326,1511,1536]" pageId="14" pageNumber="155">
|
||
<typeStatus id="54C58812FFC2FFD0FF64FA509319C662" box="[199,321,1511,1536]" pageId="14" pageNumber="155" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
UF 52676, partial skeleton from Waddells Mill Creek, Jackson County, Florida, on
|
||
<date id="FFC01070FFC2FFD0FB5EFA5F97C4C662" box="[1277,1436,1511,1536]" pageId="14" pageNumber="155" value="1978-04-10">10 April 1978</date>
|
||
by L. Richard Franz
|
||
<emphasis id="B90AEAA2FFC2FFD0FE23F9B893E8C644" box="[384,432,1550,1574]" italics="true" pageId="14" pageNumber="155">et al</emphasis>
|
||
.; UF 152479 skull from Econfina Creek, Bay County, Florida (
|
||
<geoCoordinate id="EE4A5077FFC2FFD0FBD4F9B99759C644" box="[1143,1281,1547,1575]" direction="north" orientation="latitude" pageId="14" pageNumber="155" precision="1" value="30.15274">30.15274o N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5077FFC2FFD0FAAFF9B997CEC644" box="[1292,1430,1547,1575]" direction="west" orientation="longitude" pageId="14" pageNumber="155" precision="1" value="-85.55748">85.55748o W</geoCoordinate>
|
||
, elev.
|
||
<quantity id="4C869B55FFC2FFD0FF71F98492A7C628" box="[210,255,1587,1611]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="155" unit="m" value="2.0">2 m</quantity>
|
||
,
|
||
<quantity id="4C869B55FFC2FFD0FEA9F9849300C628" box="[266,344,1587,1611]" metricMagnitude="1" metricUnit="m" metricValue="1.31" pageId="14" pageNumber="155" unit="m" value="13.1">13.1 m</quantity>
|
||
depth), on
|
||
<date id="FFC01070FFC2FFD0FE7BF98490D3C629" box="[472,651,1587,1611]" pageId="14" pageNumber="155" value="1982-08-21">21 August 1982</date>
|
||
by Joseph P. Ward and Joseph J. Ward.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364653BFFC2FFD0FF64F9E196BCC740" pageId="14" pageNumber="155" type="diagnosis">
|
||
<paragraph id="8BC136B0FFC2FFD0FF64F9E196BCC740" blockId="14.[151,1436,1511,2007]" pageId="14" pageNumber="155">
|
||
<emphasis id="B90AEAA2FFC2FFD0FF64F9E19319C60D" bold="true" box="[199,321,1622,1647]" pageId="14" pageNumber="155">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C7E4D33FFC2FFD0FEEAF9E190D8C60C" box="[329,640,1622,1646]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="155" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFC2FFD0FEEAF9E190D8C60C" box="[329,640,1622,1646]" italics="true" pageId="14" pageNumber="155">Macrochelys apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by the following: carapacial caudal notch narrow and triangular or narrow and U-shaped (
|
||
<figureCitation id="13452A35FFC2FFD0FDE8F9CD9099C6F1" box="[587,705,1658,1683]" captionStart="FIGURE 13" captionStartId="16.[151,250,950,972]" captionTargetBox="[169,1420,208,913]" captionTargetId="figure@16.[151,1436,193,928]" captionTargetPageId="16" captionText="FIGURE 13. Photograph of Macrochelys apalachicolae holotype (UF 3998) demonstrating external (A) and internal (B) carapace morphology." httpUri="https://zenodo.org/record/253021/files/figure.png" pageId="14" pageNumber="155">Figure 13</figureCitation>
|
||
), relatively shallow, and reduced; posterior projection of the squamosal globular and obtusely angled in lateral aspect (Figure 5,14); pygal with two serrations, with medial suture; peripheral 11 with one serration; distal rib end of costal 1 enters posterior third of peripheral 3; pleural scute
|
||
<date id="FFC01070FFC2FFD0FF34F9509294C69D" box="[151,204,1767,1791]" pageId="14" pageNumber="155">set 1</date>
|
||
with slight to no overlap onto the nuchal; processus trochlearis oticum relatively straight with a single distal protuberance; posterior margin of squamosal-opisthotic contact relatively straight in dorsal aspect.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364653BFFC2FFD1FF64F89990EAC17E" lastPageId="15" lastPageNumber="156" pageId="14" pageNumber="155" type="discussion">
|
||
<paragraph id="8BC136B0FFC2FFD1FF64F89990EAC17E" blockId="14.[151,1436,1511,2007]" lastBlockId="15.[151,1436,151,285]" lastPageId="15" lastPageNumber="156" pageId="14" pageNumber="155">
|
||
<emphasis id="B90AEAA2FFC2FFD0FF64F8999317C725" bold="true" box="[199,335,1838,1863]" pageId="14" pageNumber="155">Comments.</emphasis>
|
||
Although there is a general pattern of small triangular pygal regions of the carapace, there is observable variation within the species. All cranial specimens are characterized by large, globular squamosal projections that are intermediate between those of
|
||
<taxonomicName id="4C7E4D33FFC2FFD0FD6DF8C091DAC7EC" box="[718,898,1911,1934]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="155" phylum="Chordata" rank="species" species="suwannensis">
|
||
<emphasis id="B90AEAA2FFC2FFD0FD6DF8C091DAC7EC" box="[718,898,1911,1934]" italics="true" pageId="14" pageNumber="155">M. suwannensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFC2FFD0FC1BF8C09600C7EC" box="[952,1112,1910,1934]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="155" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFC2FFD0FC1BF8C09600C7EC" box="[952,1112,1910,1934]" italics="true" pageId="14" pageNumber="155">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
. Although
|
||
<taxonomicName id="4C7E4D33FFC2FFD0FB7BF8C097C4C7EC" box="[1240,1436,1910,1934]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="155" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFC2FFD0FB7BF8C097C4C7EC" box="[1240,1436,1910,1934]" italics="true" pageId="14" pageNumber="155">M. apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
is genetically most similar to
|
||
<taxonomicName id="4C7E4D33FFC2FFD0FE47F82C90DCC7D0" box="[484,644,1946,1970]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="155" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFC2FFD0FE47F82C90DCC7D0" box="[484,644,1946,1970]" italics="true" pageId="14" pageNumber="155">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
, in some ways it is morphologically more similar to
|
||
<taxonomicName id="4C7E4D33FFC2FFD0FB7AF82C97CCC7D0" box="[1241,1428,1947,1970]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="155" phylum="Chordata" rank="species" species="suwanniensis">
|
||
<emphasis id="B90AEAA2FFC2FFD0FB7AF82C97CCC7D0" box="[1241,1428,1947,1970]" italics="true" pageId="14" pageNumber="155">M. suwanniensis</emphasis>
|
||
</taxonomicName>
|
||
; they share the unique synapomorphy of a sutured pygal.
|
||
<taxonomicName id="4C7E4D33FFC2FFD0FCB7F8099611C7B4" box="[788,1097,1982,2006]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="155" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFC2FFD0FCB7F8099611C7B4" box="[788,1097,1982,2006]" italics="true" pageId="14" pageNumber="155">Macrochelys apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
is somewhat morphologically intermediate between
|
||
<taxonomicName id="4C7E4D33FFC3FFD1FE33FF2E906AC0D2" box="[400,562,152,176]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="156" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFC3FFD1FE33FF2E906AC0D2" box="[400,562,152,176]" italics="true" pageId="15" pageNumber="156">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFC3FFD1FDC8FF2E9147C0CD" box="[619,799,152,176]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="156" phylum="Chordata" rank="species" species="suwannensis">
|
||
<emphasis id="B90AEAA2FFC3FFD1FDC8FF2E9147C0CD" box="[619,799,152,176]" italics="true" pageId="15" pageNumber="156">M. suwannensis</emphasis>
|
||
</taxonomicName>
|
||
with regard to carapacial caudal notch proportions. The degree of overlap of pleural 1 onto the nuchal also suggests this (usually lying on or just anterior to the nuchalcostal 1 suture), as does a pygal that possesses two serrations (a western character) that is typically sutured medially (a character found in
|
||
<taxonomicName id="4C7E4D33FFC3FFD1FE4CFEB290F9C17E" box="[495,673,261,284]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="156" phylum="Chordata" rank="species" species="suwannensis">
|
||
<emphasis id="B90AEAA2FFC3FFD1FE4CFEB290F9C17E" box="[495,673,261,284]" italics="true" pageId="15" pageNumber="156">M. suwannensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF016638FFC3FFD1FF34F8129182C7BB" httpUri="https://zenodo.org/record/253020/files/figure.png" pageId="15" pageNumber="156" targetBox="[151,1434,337,1933]" targetPageId="15">
|
||
<paragraph id="8BC136B0FFC3FFD1FF34F8129182C7BB" blockId="15.[151,1436,1956,2009]" pageId="15" pageNumber="156">
|
||
<emphasis id="B90AEAA2FFC3FFD1FF34F812937CC7D8" bold="true" box="[151,292,1957,1979]" pageId="15" pageNumber="156">FIGURE 12.</emphasis>
|
||
Photograph of
|
||
<taxonomicName id="4C7E4D33FFC3FFD1FE68F8139086C7D9" box="[459,734,1956,1979]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="156" phylum="Chordata" rank="species" species="suwanniensis">
|
||
<emphasis id="B90AEAA2FFC3FFD1FE68F8139086C7D9" box="[459,734,1956,1979]" italics="true" pageId="15" pageNumber="156">Macrochelys suwanniensis</emphasis>
|
||
</taxonomicName>
|
||
holotype (UF 166146) demonstrating a superior (A), inferior (B), cranial (C), caudal (D), and left (E) and right (F) lateral view of skull morphology.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF016638FFDCFFCEFF34FC01932FC388" httpUri="https://zenodo.org/record/253021/files/figure.png" pageId="16" pageNumber="157" targetBox="[169,1420,208,913]" targetPageId="16">
|
||
<paragraph id="8BC136B0FFDCFFCEFF34FC01932FC388" blockId="16.[151,1436,950,1002]" pageId="16" pageNumber="157">
|
||
<emphasis id="B90AEAA2FFDCFFCEFF34FC01937CC3A9" bold="true" box="[151,292,950,972]" pageId="16" pageNumber="157">FIGURE 13.</emphasis>
|
||
Photograph of
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFE6FFC0190B0C3AE" box="[460,744,950,972]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFDCFFCEFE6FFC0190B0C3AE" box="[460,744,950,972]" italics="true" pageId="16" pageNumber="157">Macrochelys apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
holotype (UF 3998) demonstrating external (A) and internal (B) carapace morphology.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C364653BFFDCFFCEFF64FBA093C3C436" pageId="16" pageNumber="157" type="distribution">
|
||
<paragraph id="8BC136B0FFDCFFCEFF64FBA093C3C436" blockId="16.[151,1436,1047,2008]" pageId="16" pageNumber="157">
|
||
<emphasis id="B90AEAA2FFDCFFCEFF64FBA09338C452" bold="true" box="[199,352,1047,1072]" pageId="16" pageNumber="157">Distribution.</emphasis>
|
||
Restricted to river drainages bounded by the Choctawhatchee and Ochlockonee rivers in Florida,
|
||
<collectingCountry id="F3697620FFDCFFCEFF34FB8C92ACC436" box="[151,244,1083,1108]" name="Georgia" pageId="16" pageNumber="157">Georgia</collectingCountry>
|
||
, and Alabama.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364653BFFDCFFCEFF64FBE89350C516" pageId="16" pageNumber="157" type="etymology">
|
||
<paragraph id="8BC136B0FFDCFFCEFF64FBE89004C4DD" blockId="16.[151,1436,1047,2008]" pageId="16" pageNumber="157">
|
||
<emphasis id="B90AEAA2FFDCFFCEFF64FBE89316C41A" bold="true" box="[199,334,1119,1144]" pageId="16" pageNumber="157">Etymology.</emphasis>
|
||
Specific epithet refers to the new Latin
|
||
<emphasis id="B90AEAA2FFDCFFCEFCACFBE891FAC415" box="[783,930,1119,1143]" italics="true" pageId="16" pageNumber="157">apalachicol–</emphasis>
|
||
(referring to the Apalachicola River) and the Latin
|
||
<emphasis id="B90AEAA2FFDCFFCEFF78FB33935CC4F9" box="[219,260,1156,1179]" italics="true" pageId="16" pageNumber="157">–ae</emphasis>
|
||
(treating the name of the river as a Latin cognate in the First Declension, genitive case), combined to form the composite noun
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFE15FB10900DC4DD" box="[438,597,1191,1215]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFDCFFCEFE15FB10900DC4DD" box="[438,597,1191,1215]" italics="true" pageId="16" pageNumber="157">apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDCFFCEFF64FB7C9350C516" blockId="16.[151,1436,1047,2008]" pageId="16" pageNumber="157">
|
||
<emphasis id="B90AEAA2FFDCFFCEFF64FB7C9331C486" bold="true" box="[199,361,1227,1252]" pageId="16" pageNumber="157">Fossil record.</emphasis>
|
||
The earliest fossil representatives of
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFCB1FB7C91FAC481" box="[786,930,1227,1251]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFCB1FB7C91FAC481" box="[786,930,1227,1251]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
in Florida are from the early Miocene, early Barstovian NALMA, ca. 15–16 Ma. These fossils are fragmentary and consist of a partial costal 8 (UF-Vertebrate Paleontology [VP] 259076) and partial hyo- and hypoplastron (UF-VP 259077). Although difficult to ascribe to the species level, they are contemporaneous with
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFD0BFA809612C52D" authority="Matthew 1924" authorityName="Matthew" authorityYear="1924" box="[680,1098,1335,1360]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="stricta">
|
||
<emphasis id="B90AEAA2FFDCFFCEFD0BFA8091D2C52D" box="[680,906,1335,1359]" italics="true" pageId="16" pageNumber="157">Macrochelys stricta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEF4B41FFDCFFCEFC39FA80961AC52D" author="Matthew" box="[922,1090,1335,1360]" pageId="16" pageNumber="157" refString="Matthews, W. D. (1924) Third contribution to the Snake Creek fauna. Bulletin of American Natural Hist ory, 50, 59 - 210." type="journal article" year="1924">Matthew 1924</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the early Barstovian of Nebraska.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364653BFFDCFFCDFF64FAC892AFC682" lastPageId="19" lastPageNumber="160" pageId="16" pageNumber="157" type="reference_group">
|
||
<paragraph id="8BC136B0FFDCFFCFFF64FAC89109C0B6" blockId="16.[151,1436,1047,2008]" lastBlockId="17.[151,1436,151,213]" lastPageId="17" lastPageNumber="158" pageId="16" pageNumber="157">
|
||
Additional
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFEE8FAC89383C5F5" box="[331,475,1407,1431]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFEE8FAC89383C5F5" box="[331,475,1407,1431]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
specimens are not observed in Florida until the late Miocene, early Hemphillian NALMA, ca. 8–9 Ma, with the occurrence of
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFD14FA149620C5DE" authority="Dobie 1968" authorityName="Dobie" authorityYear="1968" box="[695,1144,1443,1468]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDCFFCEFD14FA149189C5D9" box="[695,977,1443,1467]" italics="true" pageId="16" pageNumber="157">Macrochelys auffenbergi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEF4B41FFDCFFCEFC40FA149637C5DE" author="Dobie" box="[995,1135,1443,1468]" pageId="16" pageNumber="157" refString="Dobie, J. L. (1968) A new turtle species of the genus Macrochelys (Chelydridae) from the Florida Pliocene. Tulane Studies in Zoology, 15, 59 - 63." type="journal article" year="1968">Dobie 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which is represented by fairly complete material from the McGehee Farm locality in Alachua County. We reexamined these
|
||
<typeStatus id="54C58812FFDCFFCEFAC8FA7F97C3C582" box="[1387,1435,1480,1504]" pageId="16" pageNumber="157">type</typeStatus>
|
||
specimens, as well as previously undescribed specimens of
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFC99FA5B91B8C661" box="[826,992,1515,1539]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDCFFCEFC99FA5B91B8C661" box="[826,992,1515,1539]" italics="true" pageId="16" pageNumber="157">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
, to diagnose the species based on shell and skull characters and to distinguish it from extant
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFD55F9B891DEC645" box="[758,902,1551,1575]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFD55F9B891DEC645" box="[758,902,1551,1575]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFC17F9A79603C645" box="[948,1115,1551,1575]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDCFFCEFC17F9A79603C645" box="[948,1115,1551,1575]" italics="true" pageId="16" pageNumber="157">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
, the nuchal and cervical are relatively narrow; pleural 1 does not contact the nuchal (shared with western
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFC53F98396C8C629" box="[1008,1168,1587,1611]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFDCFFCEFC53F98396C8C629" box="[1008,1168,1587,1611]" italics="true" pageId="16" pageNumber="157">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFB66F9849775C629" box="[1221,1325,1587,1611]" class="Reptilia" family="Chelydridae" genus="Chelydra" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFB66F9849775C629" box="[1221,1325,1587,1611]" italics="true" pageId="16" pageNumber="157">Chelydra</emphasis>
|
||
</taxonomicName>
|
||
); pygal is much longer than wide, with two serrations and a very narrow caudal notch; pygal lacks medial suturing and is keeled along vertebral 5; epiplastra are relatively wide and lobate (these are long and slender in extant
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFF34F928937FC6D5" box="[151,295,1695,1719]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFF34F928937FC6D5" box="[151,295,1695,1719]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
); and sulcal impressions for scutes on the plastron are distinct and deeply incised (these scales are very thin and their impressions faint to lacking in extant
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFC96F974919CC6B9" box="[821,964,1731,1755]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFC96F974919CC6B9" box="[821,964,1731,1755]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
). In extant
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFBF3F97496B8C6B9" box="[1104,1248,1731,1755]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFBF3F97496B8C6B9" box="[1104,1248,1731,1755]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
, the nuchal and cervical are wide; Pleural 1 does (in both
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFDCEF95F916AC69D" box="[621,818,1767,1791]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFDCFFCEFDCEF95F916AC69D" box="[621,818,1767,1791]" italics="true" pageId="16" pageNumber="157">M. apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFCCAF95F967CC69D" box="[873,1060,1768,1791]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="suwanniensis">
|
||
<emphasis id="B90AEAA2FFDCFFCEFCCAF95F967CC69D" box="[873,1060,1768,1791]" italics="true" pageId="16" pageNumber="157">M. suwanniensis</emphasis>
|
||
</taxonomicName>
|
||
) and does not (in
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFB51F95F97CBC69D" box="[1266,1427,1767,1791]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFDCFFCEFB51F95F97CBC69D" box="[1266,1427,1767,1791]" italics="true" pageId="16" pageNumber="157">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
) contact the nuchal; the pygal is much wider than long (in both
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFC30F8BB9604C741" box="[915,1116,1803,1827]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFDCFFCEFC30F8BB9604C741" box="[915,1116,1803,1827]" italics="true" pageId="16" pageNumber="157">M. apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFB3FF8BB9704C741" box="[1180,1372,1804,1827]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="suwanniensis">
|
||
<emphasis id="B90AEAA2FFDCFFCEFB3FF8BB9704C741" box="[1180,1372,1804,1827]" italics="true" pageId="16" pageNumber="157">M. suwanniensis</emphasis>
|
||
</taxonomicName>
|
||
); the epiplastra are very narrow; and the plastron lacks well-defined scute sulcal impressions. The skull of
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFADEF887934FC709" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDCFFCEFADEF887934FC709" italics="true" pageId="16" pageNumber="157">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
, although relatively large, does not exhibit the extraordinary megacephaly expressed in extant
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFF34F8C0937FC7ED" box="[151,295,1911,1935]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFF34F8C0937FC7ED" box="[151,295,1911,1935]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
. The relative head size is much smaller in
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFCBAF8CF9199C7ED" box="[793,961,1911,1935]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDCFFCEFCBAF8CF9199C7ED" box="[793,961,1911,1935]" italics="true" pageId="16" pageNumber="157">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
than in modern
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFB22F8C09749C7ED" box="[1153,1297,1911,1935]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFB22F8C09749C7ED" box="[1153,1297,1911,1935]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
, and in that way the fossil taxon is plesiomorphic. The triturating surfaces of the skull and mandible are slender and not as expanded as seen in extant
|
||
<taxonomicName id="4C7E4D33FFDCFFCEFE6DF8089006C7B5" box="[462,606,1983,2007]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDCFFCEFE6DF8089006C7B5" box="[462,606,1983,2007]" italics="true" pageId="16" pageNumber="157">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
, perhaps an indication that the fossil taxon was less durophagous (eating fewer hard-shelled organisms) than the extant
|
||
<taxonomicName id="4C7E4D33FFDDFFCFFD3EFF2F9174C0D2" box="[669,812,152,176]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="17" pageNumber="158" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDDFFCFFD3EFF2F9174C0D2" box="[669,812,152,176]" italics="true" pageId="17" pageNumber="158">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
. Increase in head size through time appears to correlate with a decrease in plastral forelobe width within
|
||
<taxonomicName id="4C7E4D33FFDDFFCFFD19FF0B9111C0B6" box="[698,841,188,212]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="17" pageNumber="158" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDDFFCFFD19FF0B9111C0B6" box="[698,841,188,212]" italics="true" pageId="17" pageNumber="158">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF016638FFDDFFCFFF34F8E591D7C7E4" httpUri="https://zenodo.org/record/253022/files/figure.png" pageId="17" pageNumber="158" targetBox="[166,1413,278,1845]" targetPageId="17">
|
||
<paragraph id="8BC136B0FFDDFFCFFF34F8E591D7C7E4" blockId="17.[151,1436,1874,1926]" pageId="17" pageNumber="158">
|
||
<emphasis id="B90AEAA2FFDDFFCFFF34F8E59347C705" bold="true" box="[151,287,1874,1896]" pageId="17" pageNumber="158">FIGURE 14.</emphasis>
|
||
Photograph of
|
||
<taxonomicName id="4C7E4D33FFDDFFCFFE18F8E59089C70A" box="[443,721,1874,1896]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="17" pageNumber="158" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFDDFFCFFE18F8E59089C70A" box="[443,721,1874,1896]" italics="true" pageId="17" pageNumber="158">Macrochelys apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
holotype (UF 3998) demonstrating a superior (A), inferior (B), cranial (C), caudal (D), and left (E) and right (F) lateral view of skull morphology.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FE96969FC113" blockId="18.[1200,1260,289,1918]" box="[1201,1223,289,369]" pageId="18" pageNumber="159">, locality</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FECF969FC1BE" blockId="18.[1200,1260,289,1918]" box="[1201,1223,376,476]" pageId="18" pageNumber="159">collection</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FE53969FC22E" blockId="18.[1200,1260,289,1918]" box="[1201,1223,484,588]" pageId="18" pageNumber="159">associated</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FDE4969FC2E3" blockId="18.[1200,1260,289,1918]" box="[1201,1223,595,641]" pageId="18" pageNumber="159">their</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FD3E969FC36F" blockId="18.[1200,1260,289,1918]" box="[1201,1223,649,781]" pageId="18" pageNumber="159">summarizing</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FCA3969FC31C" blockId="18.[1200,1260,289,1918]" box="[1201,1223,788,894]" pageId="18" pageNumber="159">specimens</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FC32969FC3DE" blockId="18.[1200,1260,289,1918]" box="[1201,1223,901,956]" pageId="18" pageNumber="159">fossil</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FC73969EC424" blockId="18.[1200,1260,289,1918]" box="[1201,1222,964,1094]" pageId="18" pageNumber="159">
|
||
<taxonomicName id="4C7E4D33FFDEFFCCFB12FC73969EC424" box="[1201,1222,964,1094]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="18" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDEFFCCFB12FC73969EC424" box="[1201,1222,964,1094]" italics="true" pageId="18" pageNumber="159">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FBFA969FC4AB" blockId="18.[1200,1260,289,1918]" box="[1201,1223,1101,1225]" pageId="18" pageNumber="159">unpublished</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FB66969FC55E" blockId="18.[1200,1260,289,1918]" box="[1201,1223,1233,1340]" pageId="18" pageNumber="159">previously</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCCFB12FAF4969FC505" blockId="18.[1200,1260,289,1918]" box="[1201,1223,1347,1383]" pageId="18" pageNumber="159">and</paragraph>
|
||
<paragraph id="8BC136B0FFDEFFCDFB12FAD89141C1B2" blockId="18.[1200,1260,289,1918]" lastBlockId="19.[151,1436,151,536]" lastPageId="19" lastPageNumber="160" pageId="18" pageNumber="159">
|
||
published. timeline of characteristics Paleontological identifying and,
|
||
<emphasis id="B90AEAA2FFDEFFCCFB13F94D969DC69B" bold="true" box="[1200,1221,1780,1813]" pageId="18" pageNumber="159">15.</emphasis>
|
||
age
|
||
<emphasis id="B90AEAA2FFDEFFCCFB13F8AB969EC71C" bold="true" box="[1200,1222,1820,1918]" pageId="18" pageNumber="159">FIGURE</emphasis>
|
||
geological
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFF64FF2F930FC0D2" box="[199,343,152,176]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFF64FF2F930FC0D2" box="[199,343,152,176]" italics="true" pageId="19" pageNumber="160">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
next appears in Florida from Polk County, from the Widden Creek and Palmetto faunas of the Bone Valley Formation, late Hemphillian, latest Miocene-earliest Pliocene (
|
||
<bibRefCitation id="EFEF4B41FFDFFFCDFBB9FF0B96E3C0B7" author="Meylan" box="[1050,1211,188,213]" pageId="19" pageNumber="160" refString="Meylan, P. A. (1995) Pleistocene amphibians and reptiles from the Leisey Shell Pit, Hillsborough, County, Florida. Bulletin of the Florida Museum of Natural History, 37, 273 - 297." type="journal article" year="1995">Meylan 1995</bibRefCitation>
|
||
). These fossils are fragmentary and occur well south of the current range of
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFCBFFF5791F3C09A" box="[796,939,224,248]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFCBFFF5791F3C09A" box="[796,939,224,248]" italics="true" pageId="19" pageNumber="160">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
. The isolated fossil elements are difficult to identify at the species level; however, the Bone Valley taxon appears larger than
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFBD0FEB29747C17E" box="[1139,1311,260,284]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDFFFCDFBD0FEB29747C17E" box="[1139,1311,260,284]" italics="true" pageId="19" pageNumber="160">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
and more comparable in size to extant specimens. A few features indicate that the fossil taxon is somewhat intermediate in morphology between
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFE28FEFA9073C106" box="[395,555,332,356]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFDFFFCDFE28FEFA9073C106" box="[395,555,332,356]" italics="true" pageId="19" pageNumber="160">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFDC1FEFA915FC106" box="[610,775,332,356]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDFFFCDFDC1FEFA915FC106" box="[610,775,332,356]" italics="true" pageId="19" pageNumber="160">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
. These features include having a pygal that is only slightly longer than wide or almost as wide as long (in
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFD3EFEC6911AC1EA" box="[669,834,368,392]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDFFFCDFD3EFEC6911AC1EA" box="[669,834,368,392]" italics="true" pageId="19" pageNumber="160">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
, the pygal is much longer than wide, and in the extant clade, it is much wider than long), and relatively long slender dentaries without overlying expanded triturating surfaces (also seen in
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFE2FFE0E9075C1B2" box="[396,557,440,464]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFDFFFCDFE2FFE0E9075C1B2" box="[396,557,440,464]" italics="true" pageId="19" pageNumber="160">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFDC0FE0E9150C1B2" box="[611,776,440,464]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDFFFCDFDC0FE0E9150C1B2" box="[611,776,440,464]" italics="true" pageId="19" pageNumber="160">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDFFFCDFF64FE6B9187C27A" blockId="19.[151,1436,151,536]" pageId="19" pageNumber="160">
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFF64FE6B930FC196" box="[199,343,476,500]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFF64FE6B930FC196" box="[199,343,476,500]" italics="true" pageId="19" pageNumber="160">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
fossils are relatively common in late Blancan to Recent (from ca. 2.5 Ma) fluvial and estuarine deposits in Florida. Most of these fossils are fragmentary. Records include:
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDFFFCDFF64FDFF9653C2C5" blockId="19.[151,1437,583,1760]" pageId="19" pageNumber="160">
|
||
1) Late Blancan (ca. 2.5 Ma)
|
||
<collectingCountry id="F3697620FFDFFFCDFDB4FDFF9063C202" box="[535,571,584,608]" name="United States of America" pageId="19" pageNumber="160">US</collectingCountry>
|
||
19 bridge site from the Suwannee River, Gilchrist County. A pygal (UF-VP 247166) from this locality is wider than long with two serrations and unsutured medially. The dentaries (UF-VP 247163–247165) are generally slender as in
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFD2BFD269176C2CA" box="[648,814,656,680]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDFFFCDFD2BFD269176C2CA" box="[648,814,656,680]" italics="true" pageId="19" pageNumber="160">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFCC7FD26965DC2CA" box="[868,1029,656,680]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFDFFFCDFCC7FD26965DC2CA" box="[868,1029,656,680]" italics="true" pageId="19" pageNumber="160">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDFFFCDFF64FD039665C28D" blockId="19.[151,1437,583,1760]" pageId="19" pageNumber="160">
|
||
2) Late Blancan (ca. 2.5 Ma) Haile 15A locality, Alachua County. A nuchal (UF-VP 259613) possesses no Pleural 1 sulci dorsally, the same condition as in
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFD19FD6E9107C292" box="[698,863,728,752]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="auffenbergi">
|
||
<emphasis id="B90AEAA2FFDFFFCDFD19FD6E9107C292" box="[698,863,728,752]" italics="true" pageId="19" pageNumber="160">M. auffenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFC35FD6E966EC292" box="[918,1078,728,752]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="temminckii">
|
||
<emphasis id="B90AEAA2FFDFFFCDFC35FD6E966EC292" box="[918,1078,728,752]" italics="true" pageId="19" pageNumber="160">M. temminckii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDFFFCDFF64FD4B90A9C33F" blockId="19.[151,1437,583,1760]" pageId="19" pageNumber="160">
|
||
3) Latest Blancan (ca. 2 Ma) De Soto Shell Pit locality (pits 1 and 3A), De Soto County. Records from this locality are farther south than the current range of the genus. A pygal (UF-VP
|
||
<date id="FFC01070FFDFFFCDFBB3FCA8963EC35A" box="[1040,1126,799,824]" pageId="19" pageNumber="160" value="15-09-24">240915</date>
|
||
) from De Soto 3A is wider than long with two serrations and unsutured medially.
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDFFFCDFF64FCDF936BC4A6" blockId="19.[151,1437,583,1760]" pageId="19" pageNumber="160">
|
||
4) Early Irvingtonian (ca. 1.6-1.0 Ma) (
|
||
<bibRefCitation id="EFEF4B41FFDFFFCDFD20FCDF91C3C3E2" author="Morgan" box="[643,923,871,896]" pageId="19" pageNumber="160" refString="Morgan, G. S. & Hulbert Jr., R. C. (1995) Overview of the geology and vertebrate biochronology of the Leisey Shell Pit local fauna, Hillsborough County, Florida. Bulletin of the Florida Museum of Natural History, 37, 1 - 92." type="journal article" year="1995">Morgan & Hulbert 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B41FFDFFFCDFC0BFCD09666C3E2" author="Meylan" box="[936,1086,871,896]" pageId="19" pageNumber="160" refString="Meylan, P. A. (1995) Pleistocene amphibians and reptiles from the Leisey Shell Pit, Hillsborough, County, Florida. Bulletin of the Florida Museum of Natural History, 37, 273 - 297." type="journal article" year="1995">Meylan 1995</bibRefCitation>
|
||
) Leisey Shell Pits (sites 1, 1B, and 3B), Hillsborough County. Both
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFD91FC3B909AC3C6" box="[562,706,908,932]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFD91FC3B909AC3C6" box="[562,706,908,932]" italics="true" pageId="19" pageNumber="160">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFD54FC3B9107C3C6" box="[759,863,908,932]" class="Reptilia" family="Chelydridae" genus="Chelydra" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFD54FC3B9107C3C6" box="[759,863,908,932]" italics="true" pageId="19" pageNumber="160">Chelydra</emphasis>
|
||
</taxonomicName>
|
||
occur at this locality, and we feel that there is some confusion with regard to Meylan’s (1995)
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFDD6FC07915DC3AA" box="[629,773,944,968]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFDD6FC07915DC3AA" box="[629,773,944,968]" italics="true" pageId="19" pageNumber="160">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
vouchers. Some of the vouchers represent other taxa (e.g., UF-VP 84005 from pit 1A is half of an emydid bridge peripheral), including a giant
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFBDDFC6396BEC38E" box="[1150,1254,980,1004]" class="Reptilia" family="Chelydridae" genus="Chelydra" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFBDDFC6396BEC38E" box="[1150,1254,980,1004]" italics="true" pageId="19" pageNumber="160">Chelydra</emphasis>
|
||
</taxonomicName>
|
||
species (e.g., a partial peripheral UF-VP 81198 from pit 1A and a partial shell UF-VP 125099 from pit 2).
|
||
<bibRefCitation id="EFEF4B41FFDFFFCDFB63FC4097C4C472" author="Meylan" box="[1216,1436,1015,1040]" pageId="19" pageNumber="160" refString="Meylan, P. A. (1995) Pleistocene amphibians and reptiles from the Leisey Shell Pit, Hillsborough, County, Florida. Bulletin of the Florida Museum of Natural History, 37, 273 - 297." type="journal article" year="1995">Meylan (1995:285)</bibRefCitation>
|
||
regarded some of his chelydrid identifications as tentative, being aware of an unnamed contemporaneous giant
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFF34FBF792A7C43A" box="[151,255,1088,1112]" class="Reptilia" family="Chelydridae" genus="Chelydra" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFF34FBF792A7C43A" box="[151,255,1088,1112]" italics="true" pageId="19" pageNumber="160">Chelydra</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B90AEAA2FFDFFFCDFEB0FBF6936FC43A" box="[275,311,1089,1112]" italics="true" pageId="19" pageNumber="160">see</emphasis>
|
||
below for discussion of
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFDFDFBF7909EC43A" box="[606,710,1088,1112]" class="Reptilia" family="Chelydridae" genus="Chelydra" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFDFDFBF7909EC43A" box="[606,710,1088,1112]" italics="true" pageId="19" pageNumber="160">Chelydra</emphasis>
|
||
</taxonomicName>
|
||
species). Also, the
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFC12FBF79618C43A" box="[945,1088,1088,1112]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFC12FBF79618C43A" box="[945,1088,1088,1112]" italics="true" pageId="19" pageNumber="160">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
left dentary (UF-VP 116093) reported from pit 3A is actually from pit 3B. The
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFD46FBD3912DC41E" box="[741,885,1124,1148]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEAA2FFDFFFCDFD46FBD3912DC41E" box="[741,885,1124,1148]" italics="true" pageId="19" pageNumber="160">Macrochelys</emphasis>
|
||
</taxonomicName>
|
||
fossils occur south of the extant range and are significant in that the dentaries are very robust with expansive triturating surfaces like those in modern
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFADEFB3E9374C4A6" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="suwanniensis">
|
||
<emphasis id="B90AEAA2FFDFFFCDFADEFB3E9374C4A6" italics="true" pageId="19" pageNumber="160">M. suwanniensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDFFFCDFF64FB789347C51A" blockId="19.[151,1437,583,1760]" pageId="19" pageNumber="160">
|
||
5) Latest Irvingtonian to earliest Rancholabrean (ca. 0.3 Ma) Oldsmar locality, Pinellas County (
|
||
<bibRefCitation id="EFEF4B41FFDFFFCDFB55FB7897D7C48A" author="Meylan" box="[1270,1423,1231,1256]" pageId="19" pageNumber="160" refString="Meylan, P. A. (1995) Pleistocene amphibians and reptiles from the Leisey Shell Pit, Hillsborough, County, Florida. Bulletin of the Florida Museum of Natural History, 37, 273 - 297." type="journal article" year="1995">Meylan 1995</bibRefCitation>
|
||
). UF-VP 135629 represents a partial posterior carapace with the pygal region well preserved. As with the Leisey specimens, this specimen most closely resembles
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFD67FAAE91D0C552" box="[708,904,1304,1328]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="apalachicolae">
|
||
<emphasis id="B90AEAA2FFDFFFCDFD67FAAE91D0C552" box="[708,904,1304,1328]" italics="true" pageId="19" pageNumber="160">M. apalachicolae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D33FFDFFFCDFC1EFAAE9620C54D" box="[957,1144,1304,1328]" class="Reptilia" family="Chelydridae" genus="Macrochelys" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="160" phylum="Chordata" rank="species" species="suwanniensis">
|
||
<emphasis id="B90AEAA2FFDFFFCDFC1EFAAE9620C54D" box="[957,1144,1304,1328]" italics="true" pageId="19" pageNumber="160">M. suwanniensis</emphasis>
|
||
</taxonomicName>
|
||
. The pygal is much wider than long, is sutured medially, and possesses two serrations, the condition most typically seen in the central assemblage.
|
||
</paragraph>
|
||
<paragraph id="8BC136B0FFDFFFCDFF64FA3392AFC682" blockId="19.[151,1437,583,1760]" pageId="19" pageNumber="160">
|
||
6) Numerous late Pleistocene Rancholabrean NALMA records exist from Florida, including: Jug Springs, Ichetucknee River,
|
||
<collectingCountry id="F3697620FFDFFFCDFED2FA109387C5A2" box="[369,479,1447,1472]" name="United States of America" pageId="19" pageNumber="160">Columbia</collectingCountry>
|
||
County (
|
||
<bibRefCitation id="EFEF4B41FFDFFFCDFDEBFA109155C5A2" author="Auffenberg" box="[584,781,1447,1472]" pageId="19" pageNumber="160" refString="Auffenberg, W. (1957) The status of the turtle Macroclemys floridana Hay. Herpetologica, 13, 123 - 126." type="journal article" year="1957">Auffenberg 1957</bibRefCitation>
|
||
); Suwannee River sites; Hornsby Springs, Santa Fe River, Alachua County; and Aucilla River 1A, Taylor County; as well as extralimital occurences from Wekiva Spring, Levy County; Rock Springs, Orange County; Oklawaha 1, Oklawaha River, Marion County; and Buzzard Island, Putnam County (
|
||
<bibRefCitation id="EFEF4B41FFDFFFCDFEF6F9A393B7C64F" author="Meylan" box="[341,495,1556,1581]" pageId="19" pageNumber="160" refString="Meylan, P. A. (1995) Pleistocene amphibians and reptiles from the Leisey Shell Pit, Hillsborough, County, Florida. Bulletin of the Florida Museum of Natural History, 37, 273 - 297." type="journal article" year="1995">Meylan 1995</bibRefCitation>
|
||
). Most of the Rancholabrean fossils are difficult to assess at the species level due to their incompleteness. Specimens from the Ichetucknee River (Suwannee River drainage), including some previously discussed by
|
||
<bibRefCitation id="EFEF4B41FFDFFFCDFE0AF9EB9025C617" author="Auffenberg" box="[425,637,1628,1653]" pageId="19" pageNumber="160" refString="Auffenberg, W. (1957) The status of the turtle Macroclemys floridana Hay. Herpetologica, 13, 123 - 126." type="journal article" year="1957">Auffenberg (1957)</bibRefCitation>
|
||
, consist of nearly complete shells and skulls (e.g., UF-VP 259848, UF- VP 259849, and UF-VP 259842). However, the squamosal and pygal regions are not preserved in these fossils. In UF-VP 259848 and UF-VP 259849, peripheral 11 possesses only one serration, indicating the pygal was also serrated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |