628 lines
80 KiB
XML
628 lines
80 KiB
XML
<document id="8E920C23DD3260C4631C298BF3A81399" ID-DOI="10.5281/zenodo.5172580" ID-ISSN="1942-1354" ID-Zenodo-Dep="5172580" ID-ZooBank="F8317A32-DFD7-42EA-8BC6-DCB5A3659F6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628270718788" checkinUser="felipe" docAuthor="Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa" docDate="2017" docId="A24087A7396A0427FF7FFB71FA32FF4C" docLanguage="en" docName="InsectaMundi.0547.1-36.pdf" docOrigin="Insecta Mundi 2017 (547)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Tomarus roigjunenti Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa 2017, New Species" docType="treatment" docVersion="3" lastPageNumber="10" masterDocId="5E79FFDF3963042CFFBFFF93FF92FF96" masterDocTitle="The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay" masterLastPageNumber="36" masterPageNumber="1" pageNumber="8" updateTime="1699038600670" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
|
||
<mods:mods id="09C3976DD15A00231920008873A14570" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="EC4E39950BB8908F9DA53B44C6D21777">
|
||
<mods:title id="46B40F3C9F72A04CFE9F8898B6EBE2F0">The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="2D13E5C9EC8CB5107C9F0EAFDF7CA52E" type="personal">
|
||
<mods:role id="4AD1DCC3F9C7501A6247B6159C0AFA23">
|
||
<mods:roleTerm id="02281D5FA285F1D15A5BB385BBC06A13">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4BD167C3EF1FBD4B334545EF1CA51DE0">Neita-Moreno, Jhon César</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0A59E3B59F2D54373254F0B9D59E49AF" type="personal">
|
||
<mods:role id="ABF0D8CD86BA440D73FED44783A921CE">
|
||
<mods:roleTerm id="C4B9B057E061E59459EC293CA41AFA03">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C2E4E9701FBFE006B2A44D07A2CC43D2">Alexander, Recursos Biológicos</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FD1D762320CD99741A1140AB68757C4E" type="personal">
|
||
<mods:role id="23174E82C9FEBE2B0D3BF4DDC2B5BE79">
|
||
<mods:roleTerm id="8747E4FF5838B1374EE940DCF4739B5F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D0098D8D950728548616EBA11AAB0D62">Agustín, San</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="5DC8D73E28A36DE1725DC153D6BDD863" type="personal">
|
||
<mods:role id="1AEA4E6B1298C39AADC84914DCDE6CBC">
|
||
<mods:roleTerm id="79901BA615412941B6AB51979A4D6F3F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1A6A79F689565A71E781350F24455CB1">de, Villa</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F73B0D1CA03D2F4715DD676F05FC5B64" type="personal">
|
||
<mods:role id="F2BDB6EA94E6AB8ECCEE9C0B510A31EE">
|
||
<mods:roleTerm id="5F0CA8EDAC59F0B07B3D920189148630">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="62CC15E1304EF68C767E6115A091FC18">Leyva</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="249D14C5BF5E67ED6BBAF2D3420387D1" type="personal">
|
||
<mods:role id="4BA5FD7A67B63F57AD222E230E0C4B58">
|
||
<mods:roleTerm id="B340C543025FC94ED2433DB34C1AA3A6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2715A79E2E70815B4A8343CCC8AD9DDD">Boyacá</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="9C1FB6341CD4D771D511C0844165E92D" type="personal">
|
||
<mods:role id="A27DC0EF16624B754CA9E964CC5F7E68">
|
||
<mods:roleTerm id="BB01656E478774FED8EA33EA1D91A26D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D2D8FB58E5B57DAC62125E816A1F9CE0">Ratcliffe, Brett C.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="66CD815734534C95175755DFEEEE78E9" type="personal">
|
||
<mods:role id="B6F4A4BD47E821A9F3476EDCCCE169E0">
|
||
<mods:roleTerm id="02C07C07A7BBA8136CCD8F2B0280FA19">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FCE092D8172896F7EEDB099BC80CFE52">Collections, Systematics Research</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="85527414E152D00F43C824BB7823645C" type="personal">
|
||
<mods:role id="3F92F533FB078C07DCE67D017C8C5809">
|
||
<mods:roleTerm id="C9D623FF2FF58CA0C1E1B5E46BC6E57D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="31B8DE4759807CF3FEED377BA1723463">of, University</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="C1C6C9474EE0A0639BCE088720B8B3B0" type="personal">
|
||
<mods:role id="6AC894C3D754B10B81422F167D3011E9">
|
||
<mods:roleTerm id="DACB3420689156C9355F6412DC0BAA1A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CB37FDF1E4B9F02402DB505DDC6321F8">Museum, Nebraska State</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="2E7570C9DC827EABF26AFC715B0951D8" type="personal">
|
||
<mods:role id="ACF5B243135B68A7F893958C9330336F">
|
||
<mods:roleTerm id="50A3BB9024B26A18DDAD2DDA12BE8090">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="55098D0532B27F6DB0EBA73C563CDF59">Hall, Nebraska</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D1ABB06E780BCC720AA28A84A597F57F" type="personal">
|
||
<mods:role id="5B73C82D6FB18255FD352760F83480A8">
|
||
<mods:roleTerm id="C926E75306938FCA5749FB373C119E12">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FE36F557F4C176A5C53A07B358E5BFE6">of, University</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FE7E6E09EC78FA7F72E773CDAD826AFC" type="personal">
|
||
<mods:role id="949F64DFC7157C2D0A528C8292EEBBF3">
|
||
<mods:roleTerm id="8428D0CA38E7BE264EB4FC90A5F0DED7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="97B4CC24DE517E291EEDDCDF350518C6">Nebraska</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="DA96A26DA5C5C8AD5D82117A3600BF43" type="personal">
|
||
<mods:role id="F4E53BE5B3852AF31791413593CB7D2E">
|
||
<mods:roleTerm id="EFDAB53BDD2EF0CD4758986279FA918C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9FE7B7D2C47C6B6ABA537142813D489C">Lincoln</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="90FC756E69D0D6B7F5FDEF0044DBCBD7" type="personal">
|
||
<mods:role id="98C80033C85DD32B71B89271F6B189EC">
|
||
<mods:roleTerm id="220B164CAF15920C1F78D1E1FC80E3C8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DB335EB692A4C2830816C7A27E688037">Ne</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B1DC82E786280610B38A1D3913E5A3BD" type="personal">
|
||
<mods:role id="EBEE8768E16B332B305D734669B92EA6">
|
||
<mods:roleTerm id="BE7A438A5CC8F52CD113A897D0786F7B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F983DECD940C68C821618CF020CBCC47">Usa</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3F55C8B9F2F02CAE2872BE86477B616D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="C528B617070EFE2BC88DAB3C18E618EC" type="host">
|
||
<mods:titleInfo id="CA1ACCE9145561CAEFD5F108C07F695F">
|
||
<mods:title id="C8D14E8A9273085B590DA3253E57E00A">Insecta Mundi</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="E744BDCBEAB698E5CE7E4BF7E9F5D5C3">
|
||
<mods:date id="4F7DB4FF674D485B3C74DDB7DEE6329A">2017</mods:date>
|
||
<mods:detail id="EB605E46C5FF302263FC26538CCCF071" type="pubDate">
|
||
<mods:number id="74B3C68E49B0A4BCA9C7C52761E12102">2017-04-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="EA0712117749E630505C1365AC2897F5" type="volume">
|
||
<mods:number id="4035EF55E3F263F0680BEA565AC2FE03">2017</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="978A83EC9B9C7664E244B4D53E2DFE5A" type="issue">
|
||
<mods:number id="52E5746FB047E05D813CC3AAC35A73A6">547</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="2E4F9789739AA309D00F6E99EADCF67F" unit="page">
|
||
<mods:start id="D9E05A02B5A6AF3AEAC0265A6AA1F32E">1</mods:start>
|
||
<mods:end id="ACECD80F24EDC707A0D13DCC7E0278E8">36</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="BCCF367A950A732F6846DB12E50B4913">journal article</mods:classification>
|
||
<mods:identifier id="01E6D443829247B9AA57336CF945491B" type="DOI">10.5281/zenodo.5172580</mods:identifier>
|
||
<mods:identifier id="3C6583E4A406D3B32E21EF038B7D5DFC" type="ISSN">1942-1354</mods:identifier>
|
||
<mods:identifier id="FA371956B8F7801AC91853F67336E47C" type="Zenodo-Dep">5172580</mods:identifier>
|
||
<mods:identifier id="7616407111AB523A0811A25555CD6DBF" type="ZooBank">F8317A32-DFD7-42EA-8BC6-DCB5A3659F6C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="A24087A7396A0427FF7FFB71FA32FF4C" LSID="urn:lsid:plazi:treatment:A24087A7396A0427FF7FFB71FA32FF4C" httpUri="http://treatment.plazi.org/id/A24087A7396A0427FF7FFB71FA32FF4C" lastPageId="11" lastPageNumber="10" pageId="9" pageNumber="8">
|
||
<subSubSection id="62F3653A396A0425FF7FFB71FC3AFB6F" box="[192,936,1249,1274]" pageId="9" pageNumber="8" type="nomenclature">
|
||
<paragraph id="2A5636B1396A0425FF7FFB71FC3AFB6F" blockId="9.[192,974,1249,1306]" box="[192,936,1249,1274]" pageId="9" pageNumber="8">
|
||
<heading id="711E81DD396A0425FF7FFB71FC3AFB6F" bold="true" box="[192,936,1249,1274]" fontSize="10" level="2" pageId="9" pageNumber="8" reason="3">
|
||
<emphasis id="189DEAA3396A0425FF7FFB71FC3AFB6F" bold="true" box="[192,936,1249,1274]" pageId="9" pageNumber="8">
|
||
<taxonomicName id="EDE94D32396A0425FF7FFB71FD7FFB6F" authority="Neita and Ratcliffe" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[192,749,1249,1274]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="roigjunenti" status="sp. nov.">
|
||
<emphasis id="189DEAA3396A0425FF7FFB71FE4CFB6F" bold="true" box="[192,478,1249,1273]" italics="true" pageId="9" pageNumber="8">Tomarus roigjunenti</emphasis>
|
||
Neita and Ratcliffe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="03AE57D8396A0425FD44FB71FC3AFB6F" box="[763,936,1249,1274]" pageId="9" pageNumber="8" rank="species">New Species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A396A0425FF7FFA91FC5CFA8C" box="[192,974,1281,1306]" pageId="9" pageNumber="8" type="description">
|
||
<paragraph id="2A5636B1396A0425FF7FFA91FC5CFA8C" blockId="9.[192,974,1249,1306]" box="[192,974,1281,1306]" pageId="9" pageNumber="8">
|
||
(
|
||
<figureCitation id="B2D22A34396A0425FF76FA91FEA5FA8C" box="[201,311,1282,1306]" captionStart="Figure 1" captionStartId="19.[192,276,1846,1868]" captionTargetBox="[207,1426,292,1810]" captionTargetId="figure-64@19.[192,1441,192,1828]" captionTargetPageId="19" captionText="Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype." figureDoi="http://doi.org/10.5281/zenodo.5172584" httpUri="https://zenodo.org/record/5172584/files/figure.png" pageId="9" pageNumber="8">Fig. 1e–f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FEFCFA91FEF3FA8C" box="[323,353,1282,1306]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="9" pageNumber="8">4e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FED1FA92FE1EFA8F" box="[366,396,1281,1305]" captionStart="Figure 5" captionStartId="23.[192,276,1853,1875]" captionTargetBox="[215,1440,192,1823]" captionTargetId="figure-63@23.[192,1443,192,1843]" captionTargetPageId="23" captionText="Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172599" httpUri="https://zenodo.org/record/5172599/files/figure.png" pageId="9" pageNumber="8">5e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FE26FA92FE58FA8F" box="[409,458,1281,1305]" captionStart="Figure 6" captionStartId="24.[192,276,1739,1761]" captionTargetBox="[211,1436,218,1720]" captionTargetId="figure-63@24.[192,1440,192,1730]" captionTargetPageId="24" captionText="Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172601" httpUri="https://zenodo.org/record/5172601/files/figure.png" pageId="9" pageNumber="8">6i–j</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FE69FA91FE66FA8F" box="[470,500,1282,1305]" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="9" pageNumber="8">7e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FDBEFA91FD8DFA8C" box="[513,543,1282,1306]" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="9" pageNumber="8">8e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FD93FA91FDF9FA8C" box="[556,619,1282,1306]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="9" pageNumber="8">10i–j</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FDC8FA91FD36FA8C" box="[631,676,1282,1306]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="9" pageNumber="8">11e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FD0EFA91FD4CFA8C" box="[689,734,1282,1306]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="9" pageNumber="8">12e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FD54FA91FC8AFA8C" box="[747,792,1282,1306]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="9" pageNumber="8">13e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FC9AFA91FCC0FA8C" box="[805,850,1282,1306]" captionStart="Figure 14" captionStartId="32.[192,276,1832,1854]" captionTargetBox="[358,1267,208,1798]" captionTargetId="figure-70@32.[349,1298,192,1821]" captionTargetPageId="32" captionText="Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite." figureDoi="http://doi.org/10.5281/zenodo.5172626" httpUri="https://zenodo.org/record/5172626/files/figure.png" pageId="9" pageNumber="8">14e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FCE1FA92FC1EFA8F" box="[862,908,1281,1305]" captionStart="Figure 15" captionStartId="33.[192,276,1817,1839]" captionTargetBox="[238,1409,229,1784]" captionTargetId="figure-88@33.[192,1440,192,1807]" captionTargetPageId="33" captionText="Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness." figureDoi="http://doi.org/10.5281/zenodo.5172630" httpUri="https://zenodo.org/record/5172630/files/figure.png" pageId="9" pageNumber="8">15e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A34396A0425FC27FA91FC56FA8C" box="[920,964,1282,1306]" captionStart="Figure 17" captionStartId="35.[192,276,1818,1840]" captionTargetBox="[235,1415,214,1804]" captionTargetId="figure-44@35.[192,1440,192,1807]" captionTargetPageId="35" captionText="Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172634" httpUri="https://zenodo.org/record/5172634/files/figure.png" pageId="9" pageNumber="8">17a</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A396A0426FF7FFAD1FAD7FBEC" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="materials_examined">
|
||
<paragraph id="2A5636B1396A0426FF7FFAD1FCEBFF6C" blockId="9.[192,1441,1345,1914]" lastBlockId="10.[192,1440,193,250]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
|
||
<materialsCitation id="9A813CEC396A0425FF7FFAD1FCBEFAEC" collectionCode="IADIZA" collectorName="Leg. M. Gentili" pageId="9" pageNumber="8" specimenCount="2" typeStatus="holotype">
|
||
<emphasis id="189DEAA3396A0425FF7FFAD1FEACFACC" bold="true" box="[192,318,1346,1370]" pageId="9" pageNumber="8">
|
||
<typeStatus id="F5528813396A0425FF7FFAD1FEACFACC" box="[192,318,1346,1370]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
male labeled: “VATRA LAUFQUEN / 14580 M.S.N.M/Neuquén-Arg. / 13-XI-69 /
|
||
<collectorName id="871C5367396A0425FAFEFAD1FE8AFAEC" pageId="9" pageNumber="8">Leg. M. Gentili</collectorName>
|
||
”, and with our red
|
||
<typeStatus id="F5528813396A0425FDBDFAF1FDF9FAEC" box="[514,619,1378,1402]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
label [
|
||
<collectionCode id="4CF8AE74396A0425FD04FAF1FCB7FAEC" box="[699,805,1378,1402]" pageId="9" pageNumber="8">IADIZA</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="9A813CEC396A0425FC89FAF2FDDAFA2C" collectingDate="2007-01-15" collectionCode="IADIZA" collectorName="R. A. Neuquen & G. San Blas Col." location="Lag. Blanca" pageId="9" pageNumber="8" specimenCount="2" typeStatus="allotype">
|
||
<emphasis id="189DEAA3396A0425FC89FAF2FC38FAEF" bold="true" box="[822,938,1377,1401]" pageId="9" pageNumber="8">
|
||
<typeStatus id="F5528813396A0425FC89FAF2FC38FAEF" box="[822,938,1377,1401]" pageId="9" pageNumber="8" type="allotype">Allotype</typeStatus>
|
||
</emphasis>
|
||
female labeled: “
|
||
<collectorName id="871C5367396A0425FBC3FAF1FABCFAEC" box="[1148,1326,1378,1402]" pageId="9" pageNumber="8">R. A. Neuquén</collectorName>
|
||
/
|
||
<quantity id="ED119B54396A0425FAFBFAF2FA16FAEC" box="[1348,1412,1377,1402]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="8" unit="km" value="5.0">5 km</quantity>
|
||
. S de
|
||
<location id="2F36606A396A0425FF5AFA11FEEAFA0C" LSID="urn:lsid:plazi:treatment:A24087A7396A0427FF7FFB71FA32FF4C:2F36606A396A0425FF5AFA11FEEAFA0C" box="[229,376,1410,1434]" name="Lag. Blanca" pageId="9" pageNumber="8">Lag. Blanca</location>
|
||
/
|
||
<date id="5E571071396A0425FE2FFA12FD87FA0F" box="[400,533,1409,1433]" pageId="9" pageNumber="8" value="2007-01-15">
|
||
<collectingDate id="4E13E999396A0425FE2FFA12FD87FA0F" box="[400,533,1409,1433]" pageId="9" pageNumber="8" value="2007-01-15">15/01/2007</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="871C5367396A0425FD93FA11FD6AFA0C" box="[556,760,1410,1434]" pageId="9" pageNumber="8">G. San Blas Col.</collectorName>
|
||
/ 39°06´56.1´´S/70°00´10.1´´W. / 1975 msnm”, and with our red
|
||
<typeStatus id="F5528813396A0425FE9DFA31FE16FA2C" box="[290,388,1442,1466]" pageId="9" pageNumber="8" type="allotype">allotype</typeStatus>
|
||
label [
|
||
<collectionCode id="4CF8AE74396A0425FE68FA31FDD3FA2C" box="[471,577,1442,1466]" pageId="9" pageNumber="8">IADIZA</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="189DEAA3396A0425FDECFA31FD72FA2C" bold="true" box="[595,736,1442,1466]" pageId="9" pageNumber="8">
|
||
<typeStatus id="F5528813396A0425FDECFA31FD72FA2C" box="[595,736,1442,1466]" pageId="9" pageNumber="8" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
labeled: “Buenos Aires: Prov. de / Buenos Aires / J. Boso Col.” (
|
||
<specimenCount id="3CEFFD38396A0425FEB2FA51FEFDFA4C" box="[269,367,1474,1498]" pageId="9" pageNumber="8" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="3CEFFD38396A0425FEC2FA51FE75FA4C" box="[381,487,1474,1498]" pageId="9" pageNumber="8" type="female">1 female</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FE40FA51FDC5FA4C" box="[511,599,1474,1498]" pageId="9" pageNumber="8">MLPA</collectionCode>
|
||
]. “Buenos Aires” (
|
||
<specimenCount id="3CEFFD38396A0425FC89FA52FC0AFA4C" box="[822,920,1473,1498]" pageId="9" pageNumber="8" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="3CEFFD38396A0425FC19FA52FB8CFA4C" box="[934,1054,1473,1498]" pageId="9" pageNumber="8" type="female">5 females</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FB8AFA51FB1FFA4C" box="[1077,1165,1474,1498]" pageId="9" pageNumber="8">MLPA</collectionCode>
|
||
]. “Chubut: Chubut” (
|
||
<specimenCount id="3CEFFD38396A0425FA2BFA51FF6FFA6C" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FEABFA71FEE0FA6C" box="[276,370,1506,1530]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="9" pageNumber="8" type="Museum">MACN</collectionCode>
|
||
]. “Mendoza: RA. Mendoza / Uspallata /
|
||
<date id="5E571071396A0425FCE6FA71FC74FA6F" box="[857,998,1505,1529]" pageId="9" pageNumber="8" value="1975-01">Enero 1975</date>
|
||
/ S. Roig” (
|
||
<specimenCount id="3CEFFD38396A0425FBCDFA71FB56FA6C" box="[1138,1220,1506,1530]" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FB65FA71FAD6FA6C" box="[1242,1348,1506,1530]" pageId="9" pageNumber="8">IADIZA</collectionCode>
|
||
]. “Mendoza / Tunuyan /
|
||
<date id="5E571071396A0425FE1CF991FE60F98C" box="[419,498,1538,1562]" pageId="9" pageNumber="8" value="1940-01">I-1940</date>
|
||
” (
|
||
<specimenCount id="3CEFFD38396A0425FDACF991FDFAF98C" box="[531,616,1538,1562]" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FD3EF991FD4BF98C" box="[641,729,1538,1562]" pageId="9" pageNumber="8">MLPA</collectionCode>
|
||
]. “Neuquén: Neuquén / Covunco /
|
||
<date id="5E571071396A0425FB2FF992FAB1F98F" box="[1168,1315,1537,1561]" pageId="9" pageNumber="8" value="1963-12-19">19-XII-1963</date>
|
||
” (
|
||
<specimenCount id="3CEFFD38396A0425FAFCF991FA0AF98C" box="[1347,1432,1538,1562]" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FF78F9B1FEA3F9AC" box="[199,305,1570,1594]" pageId="9" pageNumber="8">IADIZA</collectionCode>
|
||
]. “R. A. Neuquén / Barrancas / 5-1970 Col. A. Roig / CE 00283” (
|
||
<specimenCount id="3CEFFD38396A0425FB88F9B1FB0FF9AC" box="[1079,1181,1570,1594]" pageId="9" pageNumber="8" type="female">1 female</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FB0CF9B1FA8FF9AC" box="[1203,1309,1570,1594]" pageId="9" pageNumber="8">IADIZA</collectionCode>
|
||
]. “Covunco / Neuquén-Arg. / 29-X-62 / Leg. M. Gentili” (
|
||
<specimenCount id="3CEFFD38396A0425FD5CF9D1FCA6F9CC" box="[739,820,1602,1626]" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FCF4F9D1FC27F9CC" box="[843,949,1602,1626]" pageId="9" pageNumber="8">IADIZA</collectionCode>
|
||
]. “Neuquén / L. Victoria / Febr. 1958” (
|
||
<specimenCount id="3CEFFD38396A0425FA2BF9D1FE81F9EC" pageId="9" pageNumber="8" type="female">1 female</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FE94F9F1FE07F9EC" box="[299,405,1634,1658]" pageId="9" pageNumber="8">IADIZA</collectionCode>
|
||
]. “Neuquén / Covunco /
|
||
<date id="5E571071396A0425FD00F9F2FCC0F9EF" box="[703,850,1633,1657]" pageId="9" pageNumber="8" value="1963-12-19">19-XII-1963</date>
|
||
” (
|
||
<specimenCount id="3CEFFD38396A0425FCCDF9F1FC56F9EC" box="[882,964,1634,1658]" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FC63F9F1FBD4F9EC" box="[988,1094,1634,1658]" pageId="9" pageNumber="8">IADIZA</collectionCode>
|
||
]. “
|
||
<date id="5E571071396A0425FBDBF9F2FB7AF9EF" box="[1124,1256,1633,1657]" pageId="9" pageNumber="8" value="1955-12-08">8-XII-1955</date>
|
||
/ Collón Cura/ Pcio. Neuquén / Leg. S. Sochajovskoy” (
|
||
<specimenCount id="3CEFFD38396A0425FD16F911FCB3F90C" box="[681,801,1666,1690]" pageId="9" pageNumber="8" type="female">2 females</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FC88F911FC1DF90C" box="[823,911,1666,1690]" pageId="9" pageNumber="8">MLPA</collectionCode>
|
||
]. “Río Negro: Arg. Río Negro /Gral. Rocas / Cnia. Catriel
|
||
<quantity id="ED119B54396A0425FED6F931FE24F92F" box="[361,438,1698,1722]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="9" pageNumber="8" unit="m" value="311.0">311 m</quantity>
|
||
/ S. Roig / G.
|
||
<collectingCountry id="52FE7621396A0425FDDDF931FD3DF92C" box="[610,687,1698,1722]" name="Indonesia" pageId="9" pageNumber="8">Flores</collectingCountry>
|
||
/04/I/05 // 37°52´83´´ S/67°50´16´´W” (
|
||
<specimenCount id="3CEFFD38396A0425FB39F931FB7AF92C" box="[1158,1256,1698,1722]" pageId="9" pageNumber="8" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="3CEFFD38396A0425FB4AF931FACDF92C" box="[1269,1375,1698,1722]" pageId="9" pageNumber="8" type="female">1 female</specimenCount>
|
||
) [IA-
|
||
<collectionCode id="4CF8AE74396A0425FF7FF951FE9BF94C" box="[192,265,1730,1754]" pageId="9" pageNumber="8">DIZA</collectionCode>
|
||
]. “REP.
|
||
<collectingCountry id="52FE7621396A0425FED1F951FD8AF94C" box="[366,536,1730,1754]" name="Argentina" pageId="9" pageNumber="8">ARGENTINA</collectingCountry>
|
||
/ Río Negro / H. Ritcher” (
|
||
<specimenCount id="3CEFFD38396A0425FCDEF951FC51F94C" box="[865,963,1730,1754]" pageId="9" pageNumber="8" type="male">7 males</specimenCount>
|
||
,
|
||
<specimenCount id="3CEFFD38396A0425FC6EF951FBA9F94C" box="[977,1083,1730,1754]" pageId="9" pageNumber="8" type="female">1 female</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FBEDF951FB38F94C" box="[1106,1194,1730,1754]" pageId="9" pageNumber="8">MLPA</collectionCode>
|
||
]. “
|
||
<date id="5E571071396A0425FB77F952FAD6F94F" box="[1224,1348,1729,1753]" pageId="9" pageNumber="8" value="1929-03-05">III-5-1929</date>
|
||
/ S.O. / Río Negro” (
|
||
<specimenCount id="3CEFFD38396A0425FEE6F971FE53F96C" box="[345,449,1762,1786]" pageId="9" pageNumber="8" type="female">1 female</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FE67F971FDA4F96C" box="[472,566,1762,1786]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="9" pageNumber="8" type="Museum">MACN</collectionCode>
|
||
]. “Río Negro / Allen” (
|
||
<specimenCount id="3CEFFD38396A0425FCFBF971FC39F96C" box="[836,939,1762,1786]" pageId="9" pageNumber="8" type="female">1 female</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FC7DF971FBB2F96C" box="[962,1056,1762,1786]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="9" pageNumber="8" type="Museum">MACN</collectionCode>
|
||
]. No Data (
|
||
<specimenCount id="3CEFFD38396A0425FB13F971FAB0F96C" box="[1196,1314,1762,1786]" pageId="9" pageNumber="8" type="female">3 females</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FA87F971FA04F96C" box="[1336,1430,1762,1786]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="9" pageNumber="8" type="Museum">MACN</collectionCode>
|
||
]. “Col. Breyer” (
|
||
<specimenCount id="3CEFFD38396A0425FEC7F891FE59F88C" box="[376,459,1794,1818]" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FE5CF891FDD3F88C" box="[483,577,1794,1818]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="9" pageNumber="8" type="Museum">MACN</collectionCode>
|
||
]. “REP.
|
||
<collectingCountry id="52FE7621396A0425FD16F891FCC1F88C" box="[681,851,1794,1818]" name="Argentina" pageId="9" pageNumber="8">ARGENTINA</collectingCountry>
|
||
/ A. BREYER / Col. Breyer” (
|
||
<specimenCount id="3CEFFD38396A0425FB72F891FAB2F88C" box="[1229,1312,1794,1818]" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FA87F891FA04F88C" box="[1336,1430,1794,1818]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="9" pageNumber="8" type="Museum">MACN</collectionCode>
|
||
]. “
|
||
<collectingRegion id="E82DF853396A0425FF74F8B1FEA9F8AC" box="[203,315,1826,1850]" country="Argentina" name="Neuquen" pageId="9" pageNumber="8">Neuquen</collectingRegion>
|
||
/ Capital / i.iii.961 //
|
||
<collectingCountry id="52FE7621396A0425FD80F8B1FD2EF8AC" box="[575,700,1826,1850]" name="Argentina" pageId="9" pageNumber="8">Argentina</collectingCountry>
|
||
/ 1968 colln. / J. Daguerre” (
|
||
<specimenCount id="3CEFFD38396A0425FBA0F8B1FBE0F8AC" box="[1055,1138,1826,1850]" pageId="9" pageNumber="8" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="3CEFFD38396A0425FBC0F8B1FB7BF8AC" box="[1151,1257,1826,1850]" pageId="9" pageNumber="8" type="female">1 female</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FABFF8B1FACFF8AC" box="[1280,1373,1826,1850]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="8" type="Museum">USNM</collectionCode>
|
||
]. “21- II-60 / Cipolletti / Prov Río Negro / lg. S. Schajovskoy / Howden coll. ex. / A. Martínez coll.” (
|
||
<specimenCount id="3CEFFD38396A0425FA87F8D1FA09F8CC" box="[1336,1435,1858,1882]" pageId="9" pageNumber="8" type="male">4 males</specimenCount>
|
||
,
|
||
<specimenCount id="3CEFFD38396A0425FF7FF8F1FEA5F8EC" box="[192,311,1890,1914]" pageId="9" pageNumber="8" type="female">2 females</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE74396A0425FEF2F8F1FE3AF8EC" box="[333,424,1890,1914]" pageId="9" pageNumber="8">CMNC</collectionCode>
|
||
,
|
||
<collectionCode id="4CF8AE74396A0425FE0CF8F1FD94F8EC" box="[435,518,1890,1914]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="9" pageNumber="8">BCRC</collectionCode>
|
||
]. “
|
||
<date id="5E571071396A0425FD9CF8F2FD39F8EF" box="[547,683,1889,1913]" pageId="9" pageNumber="8" value="1955-11-24">24-XI-1955</date>
|
||
/ Villa Regina / Pcia. Rio-Negro / Lg. E. Fleiss // Howden coll. ex. / A. Martínez coll. // Lgyrus [sic] (T.) / gyas / niger / (Blanch.) / A. MARTÍNEZ-DET. 1990” (
|
||
<specimenCount id="3CEFFD3839690426FAF8FF51FA0AFF4C" box="[1351,1432,194,218]" pageId="10" pageNumber="9" type="male">1 male</specimenCount>
|
||
) [
|
||
<collectionCode id="4CF8AE7439690426FF78FF71FEB6FF6C" box="[199,292,226,250]" pageId="10" pageNumber="9">CMNC</collectionCode>
|
||
].
|
||
<typeStatus id="F552881339690426FE89FF71FE21FF6C" box="[310,435,226,250]" pageId="10" pageNumber="9" type="paratype">Paratypes</typeStatus>
|
||
each with our yellow
|
||
<typeStatus id="F552881339690426FD7FFF71FCBCFF6C" box="[704,814,226,250]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
label.
|
||
</paragraph>
|
||
<paragraph id="2A5636B139690426FF7FFEB1FD19FC4F" blockId="10.[192,1441,289,1146]" pageId="10" pageNumber="9">
|
||
Description of
|
||
<typeStatus id="F552881339690426FEC9FEB1FE7BFEAC" box="[374,489,290,314]" pageId="10" pageNumber="9" type="holotype">Holotype</typeStatus>
|
||
. Length
|
||
<quantity id="ED119B5439690426FDEAFEB2FD56FEAF" box="[597,708,289,313]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="10" pageNumber="9" unit="mm" value="26.8">26.8 mm</quantity>
|
||
; width 13.0 mm. Color dark reddish brown to black. Head: Frons and clypeus coarsely rugose. Frontoclypeal region with 2 distinctly transverse tubercles; tubercles broadly separated, usually about 3 tubercle diameters apart (
|
||
<figureCitation id="B2D22A3439690426FC08FEF1FB9CFEEC" box="[951,1038,354,378]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="10" pageNumber="9">Fig. 4e</figureCitation>
|
||
). Clypeus strongly rugopunctate to transversely rugose, tapering to slightly emarginate, broad apex with 2 reflexed teeth. Mandibles with 2 teeth, apical tooth subacute, subapical notch narrow (
|
||
<figureCitation id="B2D22A3439690426FC25FE31FB93FE2F" box="[922,1025,417,442]" captionStart="Figure 6" captionStartId="24.[192,276,1739,1761]" captionTargetBox="[211,1436,218,1720]" captionTargetId="figure-63@24.[192,1440,192,1730]" captionTargetPageId="24" captionText="Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172601" httpUri="https://zenodo.org/record/5172601/files/figure.png" pageId="10" pageNumber="9">Fig. 6i–j</figureCitation>
|
||
); labium with lateral edge densely setose, paraglossa developed, apex slightly concave at center (
|
||
<figureCitation id="B2D22A3439690426FC79FE51FB8FFE4C" box="[966,1053,450,474]" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="10" pageNumber="9">Fig. 7e</figureCitation>
|
||
); maxilla with galea developed and with 6 teeth (
|
||
<figureCitation id="B2D22A3439690426FE21FE71FE67FE6F" box="[414,501,481,506]" captionStart="Figure 5" captionStartId="23.[192,276,1853,1875]" captionTargetBox="[215,1440,192,1823]" captionTargetId="figure-63@23.[192,1443,192,1843]" captionTargetPageId="23" captionText="Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172599" httpUri="https://zenodo.org/record/5172599/files/figure.png" pageId="10" pageNumber="9">Fig. 5e</figureCitation>
|
||
). Pronotum: Surface on disc moderately punctate, sides and anterior angles densely punctate to rugopunctate; punctures moderate in size on disc, becoming larger laterally; fovea rugose within. Anterior margin with a strong, median tubercle; fovea small, narrower than distance between eyes (
|
||
<figureCitation id="B2D22A3439690426FEC7FDD1FE42FDCC" box="[376,464,578,602]" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="10" pageNumber="9">Fig. 8e</figureCitation>
|
||
). Elytra: Surface densely punctate; punctures mostly moderately in size with sparser, minute punctures between larger punctures; larger punctures usually ocellate, some in double rows and with 3 pairs of double rows. Pygidium: Surface densely punctate, punctures becoming denser in basal and lateral angles; punctures small to moderate in size, weakly ocellate; in lateral view with convex surface. Legs: Protibia tridentate and with weak, basal convexity suggestive of 4
|
||
<superScript id="DD9C9BF939690426FB44FD53FA99FD59" attach="left" box="[1275,1291,704,719]" fontSize="6" pageId="10" pageNumber="9">th</superScript>
|
||
tooth, teeth subequally spaced from each other. Protarsus simple, not enlarged. Mesotibia on transverse carinae with spinules densely (subcontiguous) to broadly (separated by 1 spinule diameter) spaced (
|
||
<figureCitation id="B2D22A3439690426FAA2FC91FA01FC8C" box="[1309,1427,770,794]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="10" pageNumber="9">Fig. 10i–j</figureCitation>
|
||
). Metatibia with 1 basal, transverse carina interrupted at middle (carina with spine-like setae) and 1 medial, transverse carina at middle; apex with about 21 spinules (
|
||
<figureCitation id="B2D22A3439690426FC0FFCD1FB9FFCCC" box="[944,1037,834,858]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="10" pageNumber="9">Fig. 10j</figureCitation>
|
||
). Venter: Prosternal process long, thick, apex longitudinally oval. Mesosternal surface glabrous, slightly concave at middle. Metasternal surface with sparse, ocellate punctures. Male Genitalia: Parameres (
|
||
<figureCitation id="B2D22A3439690426FBB6FC11FBFFFC0C" box="[1033,1133,898,922]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="10" pageNumber="9">Fig. 11e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FBC5FC11FB35FC0C" box="[1146,1191,898,922]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="10" pageNumber="9">12e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FB0CFC11FB4CFC0C" box="[1203,1246,898,922]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="10" pageNumber="9">13e</figureCitation>
|
||
) subtriangular, shaft strongly narrowed to apices, mid-shaft with large, curved tooth extending laterally. Internal sac with 3 copulatory lamellae (
|
||
<figureCitation id="B2D22A3439690426FDA7FC51FDEFFC4F" box="[536,637,961,986]" captionStart="Figure 15" captionStartId="33.[192,276,1817,1839]" captionTargetBox="[238,1409,229,1784]" captionTargetId="figure-88@33.[192,1440,192,1807]" captionTargetPageId="33" captionText="Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness." figureDoi="http://doi.org/10.5281/zenodo.5172630" httpUri="https://zenodo.org/record/5172630/files/figure.png" pageId="10" pageNumber="9">Fig. 15e</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A5636B139690426FF4FFC72FDC6FBAC" blockId="10.[192,1441,289,1146]" pageId="10" pageNumber="9">
|
||
<emphasis id="189DEAA339690426FF4FFC72FE40FC6C" bold="true" box="[240,466,993,1018]" pageId="10" pageNumber="9">
|
||
<typeStatus id="F552881339690426FF4FFC72FEF6FC6F" box="[240,356,993,1017]" pageId="10" pageNumber="9" type="allotype">Allotype</typeStatus>
|
||
Female
|
||
</emphasis>
|
||
. Length
|
||
<quantity id="ED119B5439690426FDFAFC72FD26FC6F" box="[581,692,993,1017]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="10" pageNumber="9" unit="mm" value="27.5">27.5 mm</quantity>
|
||
; width
|
||
<quantity id="ED119B5439690426FCADFC71FC10FC6F" box="[786,898,994,1018]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="10" pageNumber="9" unit="mm" value="13.9">13.9 mm</quantity>
|
||
. Similar to male, but pronotal surface with punctures slightly less dense, pygidial surface weakly convex. Gonocoxite larger than gonocoxal sternite, with a semicircular furrow (14e).
|
||
</paragraph>
|
||
<paragraph id="2A5636B139690426FF4FFBD1FAD7FBEC" blockId="10.[192,1441,289,1146]" pageId="10" pageNumber="9">
|
||
<emphasis id="189DEAA339690426FF4FFBD1FEEFFBCC" bold="true" box="[240,381,1090,1114]" pageId="10" pageNumber="9">
|
||
<typeStatus id="F552881339690426FF4FFBD1FEEFFBCC" box="[240,381,1090,1114]" pageId="10" pageNumber="9" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
. Length
|
||
<quantity id="ED119B5439690426FE54FBD2FD08FBCF" box="[491,666,1089,1113]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" metricValueMax="2.75" metricValueMin="2.63" pageId="10" pageNumber="9" unit="mm" value="26.9" valueMax="27.5" valueMin="26.3">26.3–27.5 mm</quantity>
|
||
; width
|
||
<quantity id="ED119B5439690426FD4BFBD1FC37FBCF" box="[756,933,1090,1114]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" metricValueMax="1.39" metricValueMin="1.29" pageId="10" pageNumber="9" unit="mm" value="13.4" valueMax="13.9" valueMin="12.9">12.9–13.9 mm</quantity>
|
||
. The
|
||
<specimenCount id="3CEFFD3839690426FC59FBD1FBD5FBCC" box="[998,1095,1090,1114]" pageId="10" pageNumber="9" type="male">32 male</specimenCount>
|
||
and
|
||
<specimenCount id="3CEFFD3839690426FB3BFBD1FB68FBCC" box="[1156,1274,1090,1114]" pageId="10" pageNumber="9" type="female">20 female</specimenCount>
|
||
<typeStatus id="F552881339690426FABDFBD1FAEEFBCC" box="[1282,1404,1090,1114]" pageId="10" pageNumber="9" type="paratype">paratypes</typeStatus>
|
||
do not significantly differ from their respective primary type other than slight differences in size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39690426FF7FFB31FD15FB4C" pageId="10" pageNumber="9" type="etymology">
|
||
<paragraph id="2A5636B139690426FF7FFB31FD15FB4C" blockId="10.[192,1440,1186,1242]" pageId="10" pageNumber="9">
|
||
<emphasis id="189DEAA339690426FF7FFB31FEC6FB2C" bold="true" box="[192,340,1186,1210]" pageId="10" pageNumber="9">Etymology</emphasis>
|
||
. This species is named in honor of Dr. Sergio Roig-Juñent in recognition of his invaluable entomological research in
|
||
<collectingCountry id="52FE762139690426FDBDFB51FD11FB4C" box="[514,643,1218,1242]" name="Argentina" pageId="10" pageNumber="9">Argentina</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39690426FF7FFA92FEF8FA4C" pageId="10" pageNumber="9" type="distribution">
|
||
<paragraph id="2A5636B139690426FF7FFA92FED7FAAC" blockId="10.[192,1440,1281,1338]" pageId="10" pageNumber="9">
|
||
<emphasis id="189DEAA339690426FF7FFA92FEFEFA8F" bold="true" box="[192,364,1281,1305]" pageId="10" pageNumber="9">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="EDE94D3239690426FEC3FA91FDEEFA8F" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[380,636,1282,1305]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="189DEAA339690426FEC3FA91FDEEFA8F" box="[380,636,1282,1305]" italics="true" pageId="10" pageNumber="9">Tomarus roigjunenti</emphasis>
|
||
</taxonomicName>
|
||
occurs in the provinces of
|
||
<collectingRegion id="E82DF85339690426FC74FA91FBAFFA8C" box="[971,1085,1282,1306]" country="Argentina" name="Mendoza" pageId="10" pageNumber="9">Mendoza</collectingRegion>
|
||
,
|
||
<collectingRegion id="E82DF85339690426FBF5FA91FB2FFA8C" box="[1098,1213,1282,1306]" country="Argentina" name="Neuquen" pageId="10" pageNumber="9">Neuquén</collectingRegion>
|
||
, and Río Negro in
|
||
<collectingCountry id="52FE762139690426FF7FFAB1FED3FAAC" box="[192,321,1314,1338]" name="Argentina" pageId="10" pageNumber="9">Argentina</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2A5636B139690426FF7FFAF2FB57FAEC" blockId="10.[192,1221,1377,1402]" box="[192,1221,1377,1402]" pageId="10" pageNumber="9">
|
||
<emphasis id="189DEAA339690426FF7FFAF2FE38FAEC" bold="true" box="[192,426,1377,1402]" pageId="10" pageNumber="9">Locality Records</emphasis>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FE05FAF1FDB2FAEC" box="[442,544,1378,1402]" captionStart="Figure 17" captionStartId="35.[192,276,1818,1840]" captionTargetBox="[235,1415,214,1804]" captionTargetId="figure-44@35.[192,1440,192,1807]" captionTargetPageId="35" captionText="Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172634" httpUri="https://zenodo.org/record/5172634/files/figure.png" pageId="10" pageNumber="9">Fig. 17a</figureCitation>
|
||
).
|
||
<specimenCount id="3CEFFD3839690426FD89FAF2FD48FAEC" box="[566,730,1377,1402]" pageId="10" pageNumber="9" type="generic">52 specimens</specimenCount>
|
||
examined (see
|
||
<typeStatus id="F552881339690426FC27FAF1FC5FFAEC" box="[920,973,1378,1402]" pageId="10" pageNumber="9">type</typeStatus>
|
||
information above).
|
||
</paragraph>
|
||
<paragraph id="2A5636B139690426FF7FFA31FEF8FA4C" blockId="10.[192,1440,1441,1498]" pageId="10" pageNumber="9">
|
||
<emphasis id="189DEAA339690426FF7FFA31FE69FA2F" bold="true" box="[192,507,1441,1466]" pageId="10" pageNumber="9">Temporal Distribution</emphasis>
|
||
. January (7), February (5), March (4), May (1), October (1), November (2), December (3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39690427FF7FF992FA32FF4C" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9" type="diagnosis">
|
||
<paragraph id="2A5636B139690426FF7FF992FDC4F8EF" blockId="10.[192,1440,1537,1914]" pageId="10" pageNumber="9">
|
||
<emphasis id="189DEAA339690426FF7FF992FEDAF98F" bold="true" box="[192,328,1537,1561]" pageId="10" pageNumber="9">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="EDE94D3239690426FEE9F991FDC6F98F" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[342,596,1538,1561]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="189DEAA339690426FEE9F991FDC6F98F" box="[342,596,1538,1561]" italics="true" pageId="10" pageNumber="9">Tomarus roigjunenti</emphasis>
|
||
</taxonomicName>
|
||
is externally similar to
|
||
<taxonomicName id="EDE94D3239690426FCC4F991FC41F98F" authorityName="Erichson" authorityYear="1848" box="[891,979,1538,1561]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="189DEAA339690426FCC4F991FC41F98F" box="[891,979,1538,1561]" italics="true" pageId="10" pageNumber="9">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EDE94D3239690426FBAFF991FB02F98F" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[1040,1168,1538,1561]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339690426FBAFF991FBB0F98F" box="[1040,1058,1538,1561]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FB8EF991FB02F98F" box="[1073,1168,1538,1561]" italics="true" pageId="10" pageNumber="9">rosettae</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="EDE94D3239690426FB7EF991FAF8F98F" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[1217,1386,1538,1561]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="189DEAA339690426FB7EF991FAF8F98F" box="[1217,1386,1538,1561]" italics="true" pageId="10" pageNumber="9">T. roigjunenti</emphasis>
|
||
</taxonomicName>
|
||
, the apex of the labium is slightly concave (
|
||
<figureCitation id="B2D22A3439690426FD14F9B1FC91F9AC" box="[683,771,1570,1594]" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="10" pageNumber="9">Fig. 7e</figureCitation>
|
||
), while in
|
||
<taxonomicName id="EDE94D3239690426FC35F9B1FC76F9AF" authorityName="Erichson" authorityYear="1848" box="[906,996,1570,1593]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="189DEAA339690426FC35F9B1FC0EF9AF" box="[906,924,1570,1593]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FC12F9B1FC76F9AF" box="[941,996,1570,1593]" italics="true" pageId="10" pageNumber="9">gyas</emphasis>
|
||
</taxonomicName>
|
||
the labium has a small notch (
|
||
<figureCitation id="B2D22A3439690426FACFF9B1FF49F9CC" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="10" pageNumber="9">Fig. 7c</figureCitation>
|
||
) and in
|
||
<taxonomicName id="EDE94D3239690426FEFAF9D1FE54F9CF" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[325,454,1602,1625]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339690426FEFAF9D1FE54F9CF" box="[325,454,1602,1625]" italics="true" pageId="10" pageNumber="9">T. rosettae</emphasis>
|
||
</taxonomicName>
|
||
there is a small tooth (
|
||
<figureCitation id="B2D22A3439690426FD53F9D1FCD2F9CC" box="[748,832,1602,1626]" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="10" pageNumber="9">Fig. 7f</figureCitation>
|
||
). The clypeal teeth are distinctly separated in
|
||
<taxonomicName id="EDE94D3239690426FA39F9D1FE8DF9EF" baseAuthorityName="Endrodi" baseAuthorityYear="1968" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339690426FA39F9D1FE8DF9EF" italics="true" pageId="10" pageNumber="9">T. rosettae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EDE94D3239690426FEE1F9F1FE24F9EF" authorityName="Erichson" authorityYear="1848" box="[350,438,1634,1657]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="189DEAA339690426FEE1F9F1FE24F9EF" box="[350,438,1634,1657]" italics="true" pageId="10" pageNumber="9">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FE78F9F1FDA6F9EC" box="[455,564,1634,1658]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="10" pageNumber="9">Fig. 4c, f</figureCitation>
|
||
) but nearly contiguous in
|
||
<taxonomicName id="EDE94D3239690426FCC5F9F1FBB6F9EF" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[890,1060,1634,1657]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="189DEAA339690426FCC5F9F1FC1EF9EF" box="[890,908,1634,1657]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FC23F9F1FBB6F9EF" box="[924,1060,1634,1657]" italics="true" pageId="10" pageNumber="9">roigjunenti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FB8AF9F1FB1EF9EC" box="[1077,1164,1634,1658]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="10" pageNumber="9">Fig. 4e</figureCitation>
|
||
). The tubercles on the head are relatively close together in
|
||
<taxonomicName id="EDE94D3239690426FD39F911FC9FF90F" authorityName="Erichson" authorityYear="1847" box="[646,781,1666,1689]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="maimon">
|
||
<emphasis id="189DEAA339690426FD39F911FD0AF90F" box="[646,664,1666,1689]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FD17F911FC9FF90F" box="[680,781,1666,1689]" italics="true" pageId="10" pageNumber="9">maimon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EDE94D3239690426FCA2F911FC0CF90F" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[797,926,1666,1689]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339690426FCA2F911FC0CF90F" box="[797,926,1666,1689]" italics="true" pageId="10" pageNumber="9">T. rosettae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EDE94D3239690426FC5BF911FB1CF90F" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[996,1166,1666,1689]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="189DEAA339690426FC5BF911FC64F90F" box="[996,1014,1666,1689]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FBB9F911FB1CF90F" box="[1030,1166,1666,1689]" italics="true" pageId="10" pageNumber="9">roigjunenti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FB20F911FA9FF90C" box="[1183,1293,1666,1690]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="10" pageNumber="9">Fig. 4d–f</figureCitation>
|
||
) but widely separated from each other in
|
||
<taxonomicName id="EDE94D3239690426FD8CF931FD1EF92F" authorityName="Erichson" authorityYear="1848" box="[563,652,1698,1721]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="189DEAA339690426FD8CF931FDD7F92F" box="[563,581,1698,1721]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FDEAF931FD1EF92F" box="[597,652,1698,1721]" italics="true" pageId="10" pageNumber="9">gyas</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FD21F931FD67F92C" box="[670,757,1698,1722]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="10" pageNumber="9">Fig. 4c</figureCitation>
|
||
). The pronotal fovea is narrow or nearly obsolete in
|
||
<taxonomicName id="EDE94D3239690426FA39F931FEB7F94F" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="maimon">
|
||
<emphasis id="189DEAA339690426FA39F931FA0AF92F" box="[1414,1432,1698,1721]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FF7FF951FEB7F94F" box="[192,293,1730,1753]" italics="true" pageId="10" pageNumber="9">maimon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EDE94D3239690426FE8CF951FE21F94F" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[307,435,1730,1753]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339690426FE8CF951FE21F94F" box="[307,435,1730,1753]" italics="true" pageId="10" pageNumber="9">T. rosettae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EDE94D3239690426FE49F951FD0CF94F" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[502,670,1730,1753]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="189DEAA339690426FE49F951FD9AF94F" box="[502,520,1730,1753]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FDA9F951FD0CF94F" box="[534,670,1730,1753]" italics="true" pageId="10" pageNumber="9">roigjunenti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FD11F951FC89F94C" box="[686,795,1730,1754]" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="10" pageNumber="9">Fig. 8d–f</figureCitation>
|
||
) but nearly round and usally as wide as the distance between the eyes (rarely less) in
|
||
<taxonomicName id="EDE94D3239690426FDEAF971FD3FF96F" authorityName="Erichson" authorityYear="1848" box="[597,685,1762,1785]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="189DEAA339690426FDEAF971FD3FF96F" box="[597,685,1762,1785]" italics="true" pageId="10" pageNumber="9">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FD02F971FC80F96C" box="[701,786,1762,1786]" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="10" pageNumber="9">Fig. 8c</figureCitation>
|
||
). The protibia has three teeth in
|
||
<taxonomicName id="EDE94D3239690426FB14F971FAA3F96F" authorityName="Erichson" authorityYear="1847" box="[1195,1329,1762,1785]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="maimon">
|
||
<emphasis id="189DEAA339690426FB14F971FAA3F96F" box="[1195,1329,1762,1785]" italics="true" pageId="10" pageNumber="9">T. maimon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FAFDF971FA0AF96C" box="[1346,1432,1762,1786]" captionStart="Figure 9" captionStartId="27.[192,276,742,764]" captionTargetBox="[211,1420,216,723]" captionTargetId="figure-35@27.[192,1439,192,740]" captionTargetPageId="27" captionText="Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon." figureDoi="http://doi.org/10.5281/zenodo.5172612" httpUri="https://zenodo.org/record/5172612/files/figure.png" pageId="10" pageNumber="9">Fig. 9b</figureCitation>
|
||
) and 4 teeth in
|
||
<taxonomicName id="EDE94D3239690426FECEF891FE5BF88F" authorityName="Erichson" authorityYear="1848" box="[369,457,1794,1817]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="189DEAA339690426FECEF891FE11F88F" box="[369,387,1794,1817]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FE2DF891FE5BF88F" box="[402,457,1794,1817]" italics="true" pageId="10" pageNumber="9">gyas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EDE94D3239690426FE68F891FDC4F88F" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[471,598,1794,1817]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339690426FE68F891FDC4F88F" box="[471,598,1794,1817]" italics="true" pageId="10" pageNumber="9">T. rosettae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EDE94D3239690426FD26F891FCD0F88F" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[665,834,1794,1817]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="189DEAA339690426FD26F891FD39F88F" box="[665,683,1794,1817]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FD05F891FCD0F88F" box="[698,834,1794,1817]" italics="true" pageId="10" pageNumber="9">roigjunenti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2D22A3439690426FCEDF891FC3AF88C" box="[850,936,1794,1818]" captionStart="Figure 9" captionStartId="27.[192,276,742,764]" captionTargetBox="[211,1420,216,723]" captionTargetId="figure-35@27.[192,1439,192,740]" captionTargetPageId="27" captionText="Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon." figureDoi="http://doi.org/10.5281/zenodo.5172612" httpUri="https://zenodo.org/record/5172612/files/figure.png" pageId="10" pageNumber="9">Fig. 9a</figureCitation>
|
||
). The parameres are different among the species (compare
|
||
<figureCitation id="B2D22A3439690426FE25F8B1FE6DF8AC" box="[410,511,1826,1850]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="10" pageNumber="9">Fig. 11c</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FDB1F8B1FDA8F8AC" box="[526,570,1826,1850]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="10" pageNumber="9">12c</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FDF7F8B1FDE6F8AC" box="[584,628,1826,1850]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="10" pageNumber="9">13c</figureCitation>
|
||
;
|
||
<figureCitation id="B2D22A3439690426FD3DF8B1FD23F8AC" box="[642,689,1826,1850]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="10" pageNumber="9">11d</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FD00F8B1FD7DF8AC" box="[703,751,1826,1850]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="10" pageNumber="9">12d</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FD42F8B1FCBEF8AC" box="[765,812,1826,1850]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="10" pageNumber="9">13d</figureCitation>
|
||
;
|
||
<figureCitation id="B2D22A3439690426FC84F8B1FCF5F8AC" box="[827,871,1826,1850]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="10" pageNumber="9">11e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FCC9F8B1FC31F8AC" box="[886,931,1826,1850]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="10" pageNumber="9">12e</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FC0DF8B1FC4DF8AC" box="[946,991,1826,1850]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="10" pageNumber="9">13e</figureCitation>
|
||
, and
|
||
<figureCitation id="B2D22A3439690426FB9AF8B1FBDCF8AC" box="[1061,1102,1826,1850]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="10" pageNumber="9">11f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FBE3F8B1FB17F8AC" box="[1116,1157,1826,1850]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="10" pageNumber="9">12f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439690426FB2CF8B1FB2FF8AC" box="[1171,1213,1826,1850]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="10" pageNumber="9">13f</figureCitation>
|
||
). The internal sac of
|
||
<taxonomicName id="EDE94D3239690426FF61F8D1FEF6F8CF" authorityName="Erichson" authorityYear="1847" box="[222,356,1858,1881]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="maimon">
|
||
<emphasis id="189DEAA339690426FF61F8D1FF62F8CF" box="[222,240,1858,1881]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FF40F8D1FEF6F8CF" box="[255,356,1858,1881]" italics="true" pageId="10" pageNumber="9">maimon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EDE94D3239690426FECBF8D1FD8FF8CF" authorityName="Neita-Moreno & Alexander & Agustín & de & Leyva & Boyacá & Ratcliffe & Collections & of & Museum & Hall & of & Nebraska & Lincoln & Ne & Usa" authorityYear="2017" box="[372,541,1858,1881]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="189DEAA339690426FECBF8D1FE14F8CF" box="[372,390,1858,1881]" italics="true" pageId="10" pageNumber="9">T</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339690426FE2AF8D1FD8FF8CF" box="[405,541,1858,1881]" italics="true" pageId="10" pageNumber="9">roigjunenti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EDE94D3239690426FDE4F8D1FD49F8CF" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[603,731,1858,1881]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339690426FDE4F8D1FD49F8CF" box="[603,731,1858,1881]" italics="true" pageId="10" pageNumber="9">T. rosettae</emphasis>
|
||
</taxonomicName>
|
||
has three copulatory lamellae (
|
||
<figureCitation id="B2D22A3439690426FBDDF8D1FB73F8CF" box="[1122,1249,1857,1882]" captionStart="Figure 15" captionStartId="33.[192,276,1817,1839]" captionTargetBox="[238,1409,229,1784]" captionTargetId="figure-88@33.[192,1440,192,1807]" captionTargetPageId="33" captionText="Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness." figureDoi="http://doi.org/10.5281/zenodo.5172630" httpUri="https://zenodo.org/record/5172630/files/figure.png" pageId="10" pageNumber="9">Fig. 15d–f</figureCitation>
|
||
), but in
|
||
<taxonomicName id="EDE94D3239690426FAF7F8D1FA32F8CF" authorityName="Erichson" authorityYear="1848" box="[1352,1440,1858,1881]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="189DEAA339690426FAF7F8D1FA32F8CF" box="[1352,1440,1858,1881]" italics="true" pageId="10" pageNumber="9">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
there are five lamellae (
|
||
<figureCitation id="B2D22A3439690426FE5AF8F1FDDAF8EF" box="[485,584,1889,1914]" captionStart="Figure 15" captionStartId="33.[192,276,1817,1839]" captionTargetBox="[238,1409,229,1784]" captionTargetId="figure-88@33.[192,1440,192,1807]" captionTargetPageId="33" captionText="Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness." figureDoi="http://doi.org/10.5281/zenodo.5172630" httpUri="https://zenodo.org/record/5172630/files/figure.png" pageId="10" pageNumber="9">Fig. 15c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A5636B139680427FF7FFF52FA32FF4C" blockId="11.[192,1440,193,218]" box="[192,1440,193,218]" pageId="11" pageNumber="10">
|
||
<emphasis id="189DEAA339680427FF7FFF52FEF4FF4F" bold="true" box="[192,358,193,217]" pageId="11" pageNumber="10">Life History</emphasis>
|
||
. Adults are attracted to lights at night. Some specimens have been taken in sand dunes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |