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<mods:title id="4D8DC8B72093C1E56C9ABF20D77C31C5">A redefinition of the genus Troglocharinus Reitter, 1908, with description of new species (Coleoptera, Leiodidae, Cholevinae, Leptodirini)</mods:title>
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<mods:namePart id="8E6BA360DA710211F887934E1DCE4AAF">Fresneda, Javier</mods:namePart>
<mods:affiliation id="ACD0BEE916931C81B80BEE52B28683C4">Ca de Massa, 25526 Llesp-El Pont de Suert, Lleida, Spain. &amp; Museu de CieÌncies Naturals (Zoologia), Passeig Picasso s / n, 08003 Barcelona, Spain. comasj 2 @ gmail. com</mods:affiliation>
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<mods:namePart id="7FC49D0513BB279518D563ED3856D748">Rizzo, Valeria</mods:namePart>
<mods:affiliation id="F4D1FB223690D3BA21C47964E80316F6">Myosotis M. M. Coop, Via Ulisse Aldrovandi, 18, 00197, Roma RM, Italy. valeria. rizzo 80 @ gmail. com</mods:affiliation>
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<mods:namePart id="A02A1E453A6CAA7045C0226D09AFA5BF">Comas, Jordi</mods:namePart>
<mods:affiliation id="B26C7D5708BE9C9487EFEAC4E2FD89F9">Museu de CieÌncies Naturals (Zoologia), Passeig Picasso s / n, 08003 Barcelona, Spain. comasj 2 @ gmail. com</mods:affiliation>
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<mods:namePart id="4AF29D35AD07A3408E46FFD51B7800B9">Ribera, Ignacio</mods:namePart>
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<mods:affiliation id="1F49E5A142C987A892342FD388CA7FFF">Institut de Biologia Evolutiva (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37 - 49, 08003 Barcelona, Spain. Corresponding author. ffresned @ gmail. com; https: // orcid. org / 0000 - 0003 - 0701 - 6016</mods:affiliation>
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<taxonomicName id="4C4C4D6FFFB9FFBEFF2A5A462F767341" ID-CoL="598Q9" authority="(Escola, 1971)" baseAuthorityName="Escola" baseAuthorityYear="1971" box="[151,613,1365,1392]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB9FFBEFF2A5A462F767341" bold="true" box="[151,613,1365,1392]" pageId="30" pageNumber="181">
<emphasis id="B938EAFEFFB9FFBEFF2A5A462CA37341" bold="true" box="[151,432,1365,1391]" italics="true" pageId="30" pageNumber="181">Troglocharinus vinyasi</emphasis>
(
<bibRefCitation id="EFDD4B1DFFB9FFBEFE025A462F4E7341" author="Escola, O." box="[447,605,1365,1392]" pageId="30" pageNumber="181" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà, 1971</bibRefCitation>
)
</emphasis>
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<treatmentCitationGroup id="AB5C11C2FFB9FFBEFF2A5A892F1B739C" box="[151,520,1434,1458]" pageId="30" pageNumber="181">
<treatmentCitation id="0AED10FDFFB9FFBEFF2A5A892F1B739C" author="Escola, O." box="[151,520,1434,1458]" page="32" pageId="30" pageNumber="181" year="1971">
<taxonomicName id="4C4C4D6FFFB9FFBEFF2A5A892F1B739C" ID-CoL="4YJ79" authority="Escola, 1971: 32" authorityName="Escola" authorityPageNumber="32" authorityYear="1971" baseAuthorityName="Escola" baseAuthorityYear="1971" box="[151,520,1434,1458]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB9FFBEFF2A5A892C4A739C" box="[151,345,1434,1458]" italics="true" pageId="30" pageNumber="181">Speonomus vinyasi</emphasis>
<bibRefCitation id="EFDD4B1DFFB9FFBEFEE25A892F1B739C" author="Escola, O." box="[351,520,1434,1458]" pageId="30" pageNumber="181" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà, 1971: 32</bibRefCitation>
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<emphasis id="B938EAFEFFB9FFBEFF2A5AAA2C2173FF" box="[151,306,1465,1489]" italics="true" pageId="30" pageNumber="181">Troglocharinus</emphasis>
(
<emphasis id="B938EAFEFFB9FFBEFEFC5AAA2CCF73FF" box="[321,476,1465,1489]" italics="true" pageId="30" pageNumber="181">Troglocharinus</emphasis>
) 
<emphasis id="B938EAFEFFB9FFBEFE565AAA2F2073FE" box="[491,563,1465,1488]" italics="true" pageId="30" pageNumber="181">vinyasi</emphasis>
(Escolà)
</taxonomicName>
: 
<treatmentCitation id="0AED10FDFFB9FFBEFD265AAA2E4F73FF" author="Fresneda, J." box="[667,860,1465,1489]" page="64" pageId="30" pageNumber="181" year="1998">
<bibRefCitation id="EFDD4B1DFFB9FFBEFD265AAA2E4F73FF" author="Fresneda, J." box="[667,860,1465,1489]" pageId="30" pageNumber="181" pagination="53 - 86" refId="ref17908" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda, 1998: 64</bibRefCitation>
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<typeStatus id="54F7884EFFB9FFBEFF2A5AEC2C1E7037" box="[151,269,1535,1561]" pageId="30" pageNumber="181" type="lectotype">Lectotype</typeStatus>
designation of 
<taxonomicName id="4C4C4D6FFFB9FFBEFE7D5AEC2E247037" ID-CoL="4YJ79" authority="Escola, 1971" authorityName="Escola" authorityYear="1971" baseAuthorityName="Escola" baseAuthorityYear="1971" box="[448,823,1534,1561]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB9FFBEFE7D5AEC2F8E7037" bold="true" box="[448,669,1535,1561]" italics="true" pageId="30" pageNumber="181">Speonomus vinyasi</emphasis>
<bibRefCitation id="EFDD4B1DFFB9FFBEFD1F5AED2E247037" author="Escola, O." box="[674,823,1534,1561]" pageId="30" pageNumber="181" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà, 1971</bibRefCitation>
</taxonomicName>
.
</emphasis>
<typeStatus id="54F7884EFFB9FFBEFCFF5AEC2EB97037" box="[834,938,1535,1561]" pageId="30" pageNumber="181" type="holotype">Holotype</typeStatus>
and 
<typeStatus id="54F7884EFFB9FFBEFC625AEC29527037" box="[991,1089,1535,1561]" pageId="30" pageNumber="181" type="syntype">syntypes</typeStatus>
were not designated at the species description (
<bibRefCitation id="EFDD4B1DFFB9FFBEFEE559302CE37013" author="Escola, O." box="[344,496,1571,1597]" pageId="30" pageNumber="181" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escola, 1971</bibRefCitation>
); therefore the specimens used in this study are considered 
<typeStatus id="54F7884EFFB9FFBEFB25593029E97013" box="[1176,1274,1571,1597]" pageId="30" pageNumber="181" type="syntype">syntypes</typeStatus>
(
<bibRefCitation id="EFDD4B1DFFB9FFBEFAB75930288B7013" author="ICZN" box="[1290,1432,1571,1597]" pageId="30" pageNumber="181" refId="ref18205" refString="ICZN (1999) International Commission on Zoological Nomenclature. International Code of Zoological Nomenclature. Fourth edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN, 1999</bibRefCitation>
: 72.1.1). Furthermore the type series of 
<taxonomicName id="4C4C4D6FFFB9FFBEFDE859542E38704F" ID-CoL="4YJ79" authorityName="Escola" authorityYear="1971" baseAuthorityName="Escola" baseAuthorityYear="1971" box="[597,811,1607,1633]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB9FFBEFDE859542E38704F" box="[597,811,1607,1633]" italics="true" pageId="30" pageNumber="181">Speonomus vinyasi</emphasis>
</taxonomicName>
includes specimens of two different species, which requires the designation of a unique 
<typeStatus id="54F7884EFFB9FFBEFDA459782F9370AB" box="[537,640,1643,1669]" pageId="30" pageNumber="181" type="lectotype">lectotype</typeStatus>
to avoid taxonomic misinterpretations.
</paragraph>
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<emphasis id="B938EAFEFFB9FFBEFF7A599C2C5E7087" bold="true" box="[199,333,1679,1705]" pageId="30" pageNumber="181">Type series</emphasis>
. 
<materialsCitation id="3B243CB1FFB9FFBEFEE1599C2F29713B" ID-GBIF-Occurrence="3033943310" collectionCode="MZB, LECTOTY-" collectorName="C. Ribera" county="Paratipo" determinerName="O. Escola" location="Av. del Mort" municipality="Peramola" pageId="30" pageNumber="181" specimenCount="1" typeStatus="lectotype">
<emphasis id="B938EAFEFFB9FFBEFEE1599C2CC17087" bold="true" box="[348,466,1679,1705]" pageId="30" pageNumber="181">
<typeStatus id="54F7884EFFB9FFBEFEE1599C2CC17087" box="[348,466,1679,1705]" pageId="30" pageNumber="181" type="lectotype">Lectotype</typeStatus>
</emphasis>
(by present designation) (
<figureCitation id="13772A69FFB9FFBEFD46599C2E417087" box="[763,850,1679,1705]" captionStart="FIGURE 70" captionStartId="32.[151,250,1180,1204]" captionTargetBox="[152,1435,181,1156]" captionTargetId="figure-24@32.[151,1435,181,1156]" captionTargetPageId="32" captionText="FIGURE 70. Lectotypus of Speonomus vinyasi." figureDoi="http://doi.org/10.5281/zenodo.4505974" httpUri="https://zenodo.org/record/4505974/files/figure.png" pageId="30" pageNumber="181">Fig. 70</figureCitation>
): “ 
<location id="8E936037FFB9FFBEFCCD599C29117087" LSID="urn:lsid:plazi:treatment:03E587FAFFB9FF80FF2A5A462F857137:8E936037FFB9FFBEFCCD599C29117087" box="[880,1026,1679,1705]" county="Paratipo" municipality="Peramola" name="Av. del Mort" pageId="30" pageNumber="181">Av. del Mort</location>
/ 
<collectingMunicipality id="6B97AC96FFB9FFBEFBA7599C29977087" box="[1050,1156,1679,1705]" pageId="30" pageNumber="181">Peramola</collectingMunicipality>
/ 25-X-70 ϐ / 
<collectorName id="26B9533AFFB9FFBEFA8D599C288F7087" box="[1328,1436,1679,1705]" pageId="30" pageNumber="181">C. Ribera</collectorName>
leg.”, “ 
<taxonomicName id="4C4C4D6FFFB9FFBEFF5B59A02C7570E3" box="[230,358,1715,1741]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB9FFBEFF5B59A02C7570E3" box="[230,358,1715,1741]" italics="true" pageId="30" pageNumber="181">Speonomus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4C4D6FFFB9FFBEFEC459A72CDB70E3" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[377,456,1716,1741]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi" status="sp. nov.">
<emphasis id="B938EAFEFFB9FFBEFEC459A72CDB70E3" box="[377,456,1716,1741]" italics="true" pageId="30" pageNumber="181">vinyasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785FFB9FFBEFE7359A02F1170E3" box="[462,514,1715,1741]" pageId="30" pageNumber="181" rank="species">n sp.</taxonomicNameLabel>
/ 
<determinerName id="7E2FDA28FFB9FFBEFDA859A02F9270E3" box="[533,641,1715,1741]" pageId="30" pageNumber="181">O. Escolá</determinerName>
det.”, “ 
<collectingCounty id="62924E60FFB9FFBEFD6E59A02E2570E3" box="[723,822,1715,1741]" pageId="30" pageNumber="181">Paratipo</collectingCounty>
” [invalid label], “ϐ2”, “78-1084 / 
<collectionCode id="ED5DAE29FFB9FFBEFB0859A029E470E3" box="[1205,1271,1715,1741]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
”, “ 
<collectionCode id="ED5DAE29FFB9FFBEFAA959A02DD970DF" pageId="30" pageNumber="181">LECTOTY-PUS</collectionCode>
<taxonomicName id="4C4C4D6FFFB9FFBEFF6C59C42C4270DF" box="[209,337,1751,1777]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB9FFBEFF6C59C42C4270DF" box="[209,337,1751,1777]" italics="true" pageId="30" pageNumber="181">Speonomus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4C4D6FFFB9FFBEFEDB59CB2F5F70DF" authority="Escola, 1971" authorityName="Escola" authorityYear="1971" baseAuthorityName="Escola" baseAuthorityYear="1971" box="[358,588,1751,1777]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB9FFBEFEDB59CB2CA670DF" box="[358,437,1752,1777]" italics="true" pageId="30" pageNumber="181">vinyasi</emphasis>
<bibRefCitation id="EFDD4B1DFFB9FFBEFE0659C42F5F70DF" author="Escola, O." box="[443,588,1751,1777]" pageId="30" pageNumber="181" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà, 1971</bibRefCitation>
</taxonomicName>
/ Fresneda, Rizzo, / Comas &amp; Ribera des. 2020” (
<collectionCode id="ED5DAE29FFB9FFBEFBC559C429A970DF" box="[1144,1210,1751,1777]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
), aedeagus mounted on a transparent card in the same pin.
</materialsCitation>
</paragraph>
<paragraph id="8BF336ECFFB9FFBFFF7A580C2C3177FC" blockId="30.[151,1437,1534,2030]" lastBlockId="31.[151,1437,150,1365]" lastPageId="31" lastPageNumber="182" pageId="30" pageNumber="181">
<emphasis id="B938EAFEFFB9FFBEFF7A580C2C647117" bold="true" box="[199,375,1823,1849]" pageId="30" pageNumber="181">
<typeStatus id="54F7884EFFB9FFBEFF7A580C2C647117" box="[199,375,1823,1849]" pageId="30" pageNumber="181" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D4AFD65FFB9FFBEFE3B580C2CA27117" box="[390,433,1823,1849]" pageId="30" pageNumber="181" type="female">1 ♀</specimenCount>
: “Av. del Mort 
<specimenCount id="9D4AFD65FFB9FFBEFDE2580C2F677117" box="[607,628,1823,1849]" pageId="30" pageNumber="181" type="female">♀</specimenCount>
/ Peramola 1-XI-69 / M.D.Romero-E. Roca leg.”, “ 
<taxonomicName id="4C4C4D6FFFB9FFBEFB05580C282B7117" box="[1208,1336,1823,1849]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB9FFBEFB05580C282B7117" box="[1208,1336,1823,1849]" italics="true" pageId="30" pageNumber="181">Speonomus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4C4D6FFFB9FFBEFAF05833288F7117" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[1357,1436,1824,1849]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi" status="sp. nov.">
<emphasis id="B938EAFEFFB9FFBEFAF05833288F7117" box="[1357,1436,1824,1849]" italics="true" pageId="30" pageNumber="181">vinyasi</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB9FFBEFF2A58502DC47173" bold="true" box="[151,215,1859,1885]" pageId="30" pageNumber="181">
<taxonomicNameLabel id="A20B5785FFB9FFBEFF2A58502DC47173" box="[151,215,1859,1885]" pageId="30" pageNumber="181" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/ O. Escolá det.”, “ 
<typeStatus id="54F7884EFFB9FFBEFE0858502F3A7173" box="[437,553,1859,1885]" pageId="30" pageNumber="181" type="paratype">Paratypus</typeStatus>
” [invalid label], “78-1091 / 
<collectionCode id="ED5DAE29FFB9FFBEFCD658502EB97173" box="[875,938,1859,1885]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
” (
<collectionCode id="ED5DAE29FFB9FFBEFC78585029147173" box="[965,1031,1859,1885]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
). 1 ϐ: “Av. del Mort / Peramola /1- XI-69 / E. Roca-C. Ribera leg.”, “org. cop. ϐ”, “ 
<taxonomicName id="4C4C4D6FFFB9FFBEFD0E58742E2071AF" box="[691,819,1895,1921]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB9FFBEFD0E58742E2071AF" box="[691,819,1895,1921]" italics="true" pageId="30" pageNumber="181">Speonomus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4C4D6FFFB9FFBEFCFA587B2E8571AF" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[839,918,1896,1921]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi" status="sp. nov.">
<emphasis id="B938EAFEFFB9FFBEFCFA587B2E8571AF" box="[839,918,1896,1921]" italics="true" pageId="30" pageNumber="181">vinyasi</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB9FFBEFC2058742EC871AF" bold="true" box="[925,987,1895,1921]" pageId="30" pageNumber="181">
<taxonomicNameLabel id="A20B5785FFB9FFBEFC2058742EC871AF" box="[925,987,1895,1921]" pageId="30" pageNumber="181" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/ O. Escolá det.”, “Tipo” [invalid label], “ϐ1”, “78-1083 / 
<collectionCode id="ED5DAE29FFB9FFBEFEDD58982C8C718B" box="[352,415,1931,1957]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
” (
<collectionCode id="ED5DAE29FFB9FFBEFE0558982CE9718B" box="[440,506,1931,1957]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
). 3 ϐϐ: “Av. del Mort / Peramola / 25-X-70 ϐ / C. Ribera leg.”, “ 
<taxonomicName id="4C4C4D6FFFB9FFBEFB615898284F718B" box="[1244,1372,1931,1957]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB9FFBEFB615898284F718B" box="[1244,1372,1931,1957]" italics="true" pageId="30" pageNumber="181">Speonomus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4C4D6FFFB9FFBEFACC589F2DD771E7" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi" status="sp. nov.">
<emphasis id="B938EAFEFFB9FFBEFACC589F2DD771E7" italics="true" pageId="30" pageNumber="181">vinyasi</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB9FFBEFF7658BC2C1971E7" bold="true" box="[203,266,1967,1993]" pageId="30" pageNumber="181">
<taxonomicNameLabel id="A20B5785FFB9FFBEFF7658BC2C1971E7" box="[203,266,1967,1993]" pageId="30" pageNumber="181" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/ O. Escolá det.”, “Paratipo” [invalid label], “ϐ3 [ϐ4-ϐ5]” “78-1085 / 
<collectionCode id="ED5DAE29FFB9FFBEFB9658BC297B71E7" box="[1067,1128,1967,1993]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
-78-1086 / 
<collectionCode id="ED5DAE29FFB9FFBEFB5658BC283B71E7" box="[1259,1320,1967,1993]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
- 78-1087 / 
<collectionCode id="ED5DAE29FFB9FFBEFF1B58C02DF671C3" box="[166,229,2003,2029]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
” (
<collectionCode id="ED5DAE29FFB9FFBEFF4358C02C5371C3" box="[254,320,2003,2029]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="181" type="Museum">MZB</collectionCode>
). 
<specimenCount id="9D4AFD65FFB9FFBEFEE958C02C8671C3" box="[340,405,2003,2029]" pageId="30" pageNumber="181" type="female">4 ♀♀</specimenCount>
: “Av. del Mort / Peramola / 25-X- 
<specimenCount id="9D4AFD65FFB9FFBEFCAE58C02E5F71C3" box="[787,844,2003,2030]" pageId="30" pageNumber="181" type="female">70 ♀</specimenCount>
/ C. Ribera leg.”, “ 
<taxonomicName id="4C4C4D6FFFB9FFBEFB9958C029B771C3" box="[1060,1188,2003,2029]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB9FFBEFB9958C029B771C3" box="[1060,1188,2003,2029]" italics="true" pageId="30" pageNumber="181">Speonomus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4C4D6FFFB9FFBEFB0758C7281A71C3" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[1210,1289,2004,2029]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="species" species="vinyasi" status="sp. nov.">
<emphasis id="B938EAFEFFB9FFBEFB0758C7281A71C3" box="[1210,1289,2004,2029]" italics="true" pageId="30" pageNumber="181">vinyasi</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB9FFBEFAAD58C0285C71C3" bold="true" box="[1296,1359,2003,2029]" pageId="30" pageNumber="181">
<taxonomicNameLabel id="A20B5785FFB9FFBEFAAD58C0285C71C3" box="[1296,1359,2003,2029]" pageId="30" pageNumber="181" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<specimenCount id="9D4AFD65FFB9FFBEFAEB58C0287871C3" box="[1366,1387,2003,2029]" pageId="30" pageNumber="181" type="female">♀</specimenCount>
/ O. Escolá det.”, “Paratipo” [invalid label], “78-1088 / 
<collectionCode id="ED5DAE29FFB8FFBFFD725F842E1F769F" box="[719,780,151,177]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="182" type="Museum">MZB</collectionCode>
- 78-1089 / 
<collectionCode id="ED5DAE29FFB8FFBFFC2E5F842EC3769F" box="[915,976,151,177]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="182" type="Museum">MZB</collectionCode>
- 78-1090 / 
<collectionCode id="ED5DAE29FFB8FFBFFBEA5F842985769F" box="[1111,1174,151,177]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="182" type="Museum">MZB</collectionCode>
” (
<collectionCode id="ED5DAE29FFB8FFBFFB125F8429E2769F" box="[1199,1265,151,177]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="182" type="Museum">MZB</collectionCode>
). 1 ϐ: “Av. Pau Esteve [= Pou del Dring] / Peramola (Oliana) / 18-I-70 / M. Ubach- ϐ1 / O. Escolá leg.”, “ 
<taxonomicName id="4C4C4D6FFFB8FFBFFB245FA8280A76FB" box="[1177,1305,187,213]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB8FFBFFB245FA8280A76FB" box="[1177,1305,187,213]" italics="true" pageId="31" pageNumber="182">Speonomus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4C4D6FFFB8FFBFFA925FA8286D76FA" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[1327,1406,187,212]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="vinyasi" status="sp. nov.">
<emphasis id="B938EAFEFFB8FFBFFA925FA8286D76FA" box="[1327,1406,187,212]" italics="true" pageId="31" pageNumber="182">vinyasi</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB8FFBFFA3B5FA82DAB76D7" bold="true" pageId="31" pageNumber="182">
<taxonomicNameLabel id="A20B5785FFB8FFBFFA3B5FA82DAB76D7" pageId="31" pageNumber="182" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/ O. Escolá det.”, “Paratipo” [invalid label], “ϐ1”, “78-1092 / 
<collectionCode id="ED5DAE29FFB8FFBFFCC55FCC2EA476D7" box="[888,951,223,249]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="182" type="Museum">MZB</collectionCode>
” (
<collectionCode id="ED5DAE29FFB8FFBFFC6D5FCC290176D7" box="[976,1042,223,249]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="182" type="Museum">MZB</collectionCode>
). 1 ϐ: “Av. Pau Esteve / (= Pou del Dring) / Peramola ϐ / 18-I-70 / O. Escolá leg.”, “ 
<taxonomicName id="4C4C4D6FFFB8FFBFFD755E102E5B7733" box="[712,840,259,285]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB8FFBFFD755E102E5B7733" box="[712,840,259,285]" italics="true" pageId="31" pageNumber="182">Speonomus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4C4D6FFFB8FFBFFCE25E102EBD7732" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[863,942,259,284]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="vinyasi" status="sp. nov.">
<emphasis id="B938EAFEFFB8FFBFFCE25E102EBD7732" box="[863,942,259,284]" italics="true" pageId="31" pageNumber="182">vinyasi</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB8FFBFFC0B5E102EE57733" bold="true" box="[950,1014,259,285]" pageId="31" pageNumber="182">
<taxonomicNameLabel id="A20B5785FFB8FFBFFC0B5E102EE57733" box="[950,1014,259,285]" pageId="31" pageNumber="182" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/ O. Escolá det.”, “Paratipo” [invalid label], “78-1093 / 
<collectionCode id="ED5DAE29FFB8FFBFFEDF5E342CB2776F" box="[354,417,295,321]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="182" type="Museum">MZB</collectionCode>
” (
<collectionCode id="ED5DAE29FFB8FFBFFE045E342CE8776F" box="[441,507,295,321]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="182" type="Museum">MZB</collectionCode>
). 
<specimenCount id="9D4AFD65FFB8FFBFFDB05E342F2B776F" box="[525,568,295,321]" pageId="31" pageNumber="182" type="female">1 ♀</specimenCount>
: “Av. de la Roca / del Corb / Peramola / 29-XI-70 / X. Bellés, / J. Comas leg.”, “Paratipo” [invalid label] (
<collectionCode id="ED5DAE29FFB8FFBFFE795E582CE4774B" box="[452,503,331,357]" pageId="31" pageNumber="182">CJC</collectionCode>
). All the specimens with 
<typeStatus id="54F7884EFFB8FFBFFCA45E582EA3774B" box="[793,944,331,357]" pageId="31" pageNumber="182" type="paralectotype">paralectotype</typeStatus>
label: “ 
<typeStatus id="54F7884EFFB8FFBFFBBA5E582816774B" box="[1031,1285,331,357]" pageId="31" pageNumber="182" type="paralectotype">PARALECTOTYPUS</typeStatus>
/ 
<taxonomicName id="4C4C4D6FFFB8FFBFFAA05E582DF577A6" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB8FFBFFAA05E582DF577A6" italics="true" pageId="31" pageNumber="182">Speonomus vinyasi</emphasis>
</taxonomicName>
/ 
<bibRefCitation id="EFDD4B1DFFB8FFBFFF435E7C2C9C77A7" author="Escola, O." box="[254,399,367,393]" pageId="31" pageNumber="182" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà, 1971</bibRefCitation>
Fresneda, Rizzo, / Comas &amp; Ribera des. 2020”. To these specimens should be added all the 
<typeStatus id="54F7884EFFB8FFBFFF2A5E802C2A7783" box="[151,313,403,429]" pageId="31" pageNumber="182" type="paralectotype">paralectotypes</typeStatus>
of 
<taxonomicName id="4C4C4D6FFFB8FFBFFEE25E802CD87782" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[351,459,403,428]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB8FFBFFEE25E802CD87782" box="[351,459,403,428]" italics="true" pageId="31" pageNumber="182">T. vinyasi</emphasis>
</taxonomicName>
that are designated as 
<typeStatus id="54F7884EFFB8FFBFFD715E802E247783" box="[716,823,403,429]" pageId="31" pageNumber="182" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C4C4D6FFFB8FFBFFCE35E802EF27783" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[862,993,403,429]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="fadriquei" status="sp. nov.">
<emphasis id="B938EAFEFFB8FFBFFCE35E802EF27783" box="[862,993,403,429]" italics="true" pageId="31" pageNumber="182">T. fadriquei</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB8FFBFFC545E8029507783" bold="true" box="[1001,1091,403,429]" pageId="31" pageNumber="182">
<taxonomicNameLabel id="A20B5785FFB8FFBFFC545E8029507783" box="[1001,1091,403,429]" pageId="31" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see list under 
<taxonomicName id="4C4C4D6FFFB8FFBFFB4D5E8028607783" authority="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera, 2021" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[1264,1395,403,429]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="fadriquei" status="sp. nov.">
<emphasis id="B938EAFEFFB8FFBFFB4D5E8028607783" box="[1264,1395,403,429]" italics="true" pageId="31" pageNumber="182">T. fadriquei</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB8FFBFFAC65E802DDB77FF" bold="true" pageId="31" pageNumber="182">
<taxonomicNameLabel id="A20B5785FFB8FFBFFAC65E802DDB77FF" pageId="31" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
above),
</paragraph>
<paragraph id="8BF336ECFFB8FFBFFF7A5EC92DE87413" blockId="31.[151,1437,150,1365]" pageId="31" pageNumber="182">
<emphasis id="B938EAFEFFB8FFBFFF7A5EC92F0977DA" bold="true" box="[199,538,474,500]" pageId="31" pageNumber="182">Additional studied material:</emphasis>
<materialsCitation id="3B243CB1FFB8FFBFFD9C5EC82DE47413" ID-GBIF-Occurrence="3033943318" collectingDate="2017-03-19" collectionCode="IBE" collectorName="Fadrique" country="Spain" county="Avenc de la Roca del Corb" elevation="356379" location="Lleida" municipality="Peramola" pageId="31" pageNumber="182" specimenCode="T 4659411" specimenCount="1">
<collectingCountry id="F35B767CFFB8FFBFFD9C5EC82F7777DB" box="[545,612,475,501]" name="Spain" pageId="31" pageNumber="182">Spain</collectingCountry>
, 
<location id="8E936037FFB8FFBFFDD25EC82FAA77DB" LSID="urn:lsid:plazi:treatment:03E587FAFFB9FF80FF2A5A462F857137:8E936037FFB8FFBFFDD25EC82FAA77DB" box="[623,697,475,501]" country="Spain" county="Avenc de la Roca del Corb" municipality="Peramola" name="Lleida" pageId="31" pageNumber="182">Lleida</location>
, 
<collectingMunicipality id="6B97AC96FFB8FFBFFD795EC82E2277DB" box="[708,817,475,501]" pageId="31" pageNumber="182">Peramola</collectingMunicipality>
, 
<collectingCounty id="62924E60FFB8FFBFFC865EC8297977DB" box="[827,1130,475,501]" pageId="31" pageNumber="182">Avenc de la Roca del Corb</collectingCounty>
, 31 
<specimenCode id="DBEA9E97FFB8FFBFFB325EC8281A77DB" box="[1167,1289,474,501]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="31" pageNumber="182">T 4659411</specimenCode>
<quantity id="4CB49B09FFB8FFBFFAAD5EC92DBC7437" metricMagnitude="5" metricUnit="m" metricValue="3.5637886599999993" pageId="31" pageNumber="182" unit="m" value="356378.866">
<elevation id="0061D1DFFFB8FFBFFAAD5EC92DBC7437" metricMagnitude="5" metricUnit="m" metricValue="3.5637886599999993" pageId="31" pageNumber="182" unit="m" value="356378.866">356378, 866 m</elevation>
</quantity>
, 
<date id="FFF2102CFFB8FFBFFF015EEC2C207437" box="[188,307,511,537]" pageId="31" pageNumber="182" value="2017-03-19">
<collectingDate id="EFB6E9C4FFB8FFBFFF015EEC2C207437" box="[188,307,511,537]" pageId="31" pageNumber="182" value="2017-03-19">19-3-2017</collectingDate>
</date>
, 
<collectorName id="26B9533AFFB8FFBFFE825EEC2CB07437" box="[319,419,511,537]" pageId="31" pageNumber="182">Fadrique</collectorName>
leg., 1 ϐ (
<collectionCode id="ED5DAE29FFB8FFBFFDA05EEC2F5D7437" box="[541,590,511,537]" country="Spain" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="31" pageNumber="182">IBE</collectionCode>
, specimen used for DNA extraction and sequencing, voucher number IBE-AN790)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336ECFFB8FFBFFF7A5D542EC2744F" blockId="31.[151,1437,150,1365]" box="[199,977,582,609]" pageId="31" pageNumber="182">
<emphasis id="B938EAFEFFB8FFBFFF7A5D542CE9744E" bold="true" box="[199,506,582,609]" pageId="31" pageNumber="182">
Type locality (
<typeStatus id="54F7884EFFB8FFBFFED25D552CFD744E" box="[367,494,582,608]" pageId="31" pageNumber="182" type="lectotype">lectotypus</typeStatus>
).
</emphasis>
<materialsCitation id="3B243CB1FFB8FFBFFDBC5D542EDE744F" ID-GBIF-Occurrence="3033943308" box="[513,973,583,609]" location="Avenc del Mort" pageId="31" pageNumber="182" specimenCount="1">
Peramola, 
<location id="8E936037FFB8FFBFFDCA5D542E35744F" LSID="urn:lsid:plazi:treatment:03E587FAFFB9FF80FF2A5A462F857137:8E936037FFB8FFBFFDCA5D542E35744F" box="[631,806,583,609]" name="Avenc del Mort" pageId="31" pageNumber="182">Avenc del Mort</location>
(
<bibRefCitation id="EFDD4B1DFFB8FFBFFC885D542ED7744F" author="Escola, O." box="[821,964,583,609]" pageId="31" pageNumber="182" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà 1971</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3566567FFB8FFBFFF7A5D792C407517" pageId="31" pageNumber="182" type="diagnosis">
<paragraph id="8BF336ECFFB8FFBFFF7A5D792C407517" blockId="31.[151,1437,150,1365]" pageId="31" pageNumber="182">
<emphasis id="B938EAFEFFB8FFBFFF7A5D792CC174AA" bold="true" box="[199,466,618,644]" pageId="31" pageNumber="182">Differential diagnosis.</emphasis>
Bathyscioid species of 
<taxonomicName id="4C4C4D6FFFB8FFBFFD565D782E8474AB" authorityName="Reitter" authorityYear="1908" box="[747,919,619,645]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFB8FFBFFD565D782E8474AB" box="[747,919,619,645]" italics="true" pageId="31" pageNumber="182">Troglocharinus</emphasis>
</taxonomicName>
with well-developed mesoventral keel, with hook-like vertex; first male protarsomere narrower than apex of tibia, as long as 2nd–4th tarsomeres. Median lobe of aedeagus very large, regularly curved in lateral view, apex elongated and sinuate. Endophallus with very large NCs; NFVM deeply notched, larger branch directed towards apex; PF2 triangular. Spiculum of the female urite viii short, wide and with truncated apex. Lobes of the spermatheca ony slightly wider than median region, which is strongly bended.
</paragraph>
</subSubSection>
<subSubSection id="C3566567FFB8FF80FF7A5C512DD773A0" lastPageId="32" lastPageNumber="183" pageId="31" pageNumber="182" type="description">
<paragraph id="8BF336ECFFB8FFBFFF7A5C512C33758B" blockId="31.[151,1437,150,1365]" pageId="31" pageNumber="182">
<emphasis id="B938EAFEFFB8FFBFFF7A5C512CEC7572" bold="true" box="[199,511,834,861]" pageId="31" pageNumber="182">Description of the species.</emphasis>
Habitus as in 
<figureCitation id="13772A69FFB8FFBFFD1F5C502FE67573" box="[674,757,835,861]" captionStart="FIGURE 70" captionStartId="32.[151,250,1180,1204]" captionTargetBox="[152,1435,181,1156]" captionTargetId="figure-24@32.[151,1435,181,1156]" captionTargetPageId="32" captionText="FIGURE 70. Lectotypus of Speonomus vinyasi." figureDoi="http://doi.org/10.5281/zenodo.4505974" httpUri="https://zenodo.org/record/4505974/files/figure.png" pageId="31" pageNumber="182">Fig. 70</figureCitation>
. Body length 3.2 mm. Body shape oval, wide. Body surface finely punctated on pronotum, with transverse striolas on elytra. With fine, yellowish, recumbent inconspicuous pubescence.
</paragraph>
<paragraph id="8BF336ECFFB8FFBFFF7A5CBC293275C3" blockId="31.[151,1437,150,1365]" pageId="31" pageNumber="182">
Antennomere length as in 
<tableCitation id="C6CE0357FFB8FFBFFE545CBC2F2E75E7" box="[489,573,943,969]" captionStart="TABLE 3" captionStartId="31.[151,244,1410,1436]" captionTargetPageId="31" captionText="TABLE 3. Length of the antennomeres. All values in mm." httpUri="http://table.plazi.org/id/DF336664FFB8FFBFFF2A5A912E3573B3" pageId="31" pageNumber="182" tableUuid="DF336664FFB8FFBFFF2A5A912E3573B3">Table 3</tableCitation>
. Pronotum transverse, lateral margins regularly arched, slightly narrower than elytra, with maximum width at base (maximum width = 1.6 x length = 0.8 mm.).
</paragraph>
<paragraph id="8BF336ECFFB8FFBFFF7A5CE4297A723F" blockId="31.[151,1437,150,1365]" box="[199,1129,1015,1041]" pageId="31" pageNumber="182">Elytra elongated, strongly tapering towards apex, maximum width on anterior half.</paragraph>
<paragraph id="8BF336ECFFB8FFBFFF7A5B082E52721B" blockId="31.[151,1437,150,1365]" box="[199,833,1051,1077]" pageId="31" pageNumber="182">Mesoventral keel well developed, with hook-like vertex.</paragraph>
<paragraph id="8BF336ECFFB8FFBFFF7A5B2C2F2F7253" blockId="31.[151,1437,150,1365]" pageId="31" pageNumber="182">
Legs long, especially posterior legs. Mesotibia weakly arched, first protrarsomere (
<figureCitation id="13772A69FFB8FFBFFBDB5B2C29A67277" box="[1126,1205,1087,1113]" captionStart="FIGURES 7–11" captionStartId="18.[151,264,1918,1942]" captionTargetBox="[152,1436,1198,1894]" captionTargetId="figure-338@18.[152,1436,1198,1894]" captionTargetPageId="18" captionText="FIGURES 7–11. Male protarsus of Troglocharinus sendrai sp. nov. (7), T. fadriquei sp. nov. (8), T. mengeli (9)—Coves Encantades de Toloriu—, T. mercedesi (10)—Bòfia de Sant Jaume—and T. vinyasi (11)—Avenc de la Roca del Corb—." figureDoi="http://doi.org/10.5281/zenodo.4505952" httpUri="https://zenodo.org/record/4505952/files/figure.png" pageId="31" pageNumber="182">Fig. 11</figureCitation>
) narrower than apex of tibiae, as long as 2nd–4th together.
</paragraph>
<paragraph id="8BF336ECFFB8FFBFFF7A5B942EB672EB" blockId="31.[151,1437,150,1365]" pageId="31" pageNumber="182">
Median lobe of aedeagus with strongly sinuous margins in dorsal view (
<figureCitation id="13772A69FFB8FFBFFC5B5B94292A728F" box="[998,1081,1159,1185]" captionStart="FIGURES 20–24" captionStartId="22.[151,264,1951,1975]" captionTargetBox="[252,1333,182,1914]" captionTargetId="figure-24@22.[251,1336,181,1927]" captionTargetPageId="22" captionText="FIGURES 20–24. Aedeagus in dorsal view of Troglocharinus sendrai sp. nov. (20), T. fadriquei sp. nov. (21), T. mengeli (22)—Cabana d’en Garraba—, T. mercedesi (23)—Bòfia de Sant Jaume—and T. vinyasi (24)—Avenc de la Roca del Corb." figureDoi="http://doi.org/10.5281/zenodo.4505958" httpUri="https://zenodo.org/record/4505958/files/figure.png" pageId="31" pageNumber="182">Fig. 24</figureCitation>
), with apex apparently notched; regularly curved in lateral view (
<figureCitation id="13772A69FFB8FFBFFDBB5BB82F4A72EB" box="[518,601,1195,1221]" captionStart="FIGURES 15–19" captionStartId="20.[151,264,1699,1723]" captionTargetBox="[151,1424,382,1676]" captionTargetId="figure-113@20.[151,1436,381,1676]" captionTargetPageId="20" captionText="FIGURES 15–19. Aedeagus in lateral view of Troglocharinus sendrai sp. nov. (15), T. fadriquei sp. nov. (16), T. mengeli (17)—Coves Encantades de Toloriu—, T. mercedesi (18)—Bofia de Sant Jaume—and T. vinyasi (19)—Avenc de la Roca del Corb." figureDoi="http://doi.org/10.5281/zenodo.4505956" httpUri="https://zenodo.org/record/4505956/files/figure.png" pageId="31" pageNumber="182">Fig. 19</figureCitation>
); apex elongated and sinuate.
</paragraph>
<paragraph id="8BF336ECFFB8FFBFFF7A5BDC2FAF737B" blockId="31.[151,1437,150,1365]" pageId="31" pageNumber="182">Lateral stylus of the tegmen slightly sinuate, somewhat shorter than median lobe and with three long, setae on apex, two inserted laterally, one curved and other sinuate; third seta shorter, thin and straight, inserted on the opposed side. Membranous lamina long, elongated, with a long and dense tuff of setae (“penicillum”) inserted in the apical region, including the membranous lamina.</paragraph>
<caption id="DF336664FFB8FFBFFF2A5A912E3573B3" ID-Table-UUID="DF336664FFB8FFBFFF2A5A912E3573B3" box="[151,806,1410,1437]" httpUri="http://table.plazi.org/id/DF336664FFB8FFBFFF2A5A912E3573B3" pageId="31" pageNumber="182" targetBox="[166,1415,1452,1920]" targetIsTable="true" targetPageId="31">
<paragraph id="8BF336ECFFB8FFBFFF2A5A912E3573B3" blockId="31.[151,806,1410,1437]" box="[151,806,1410,1437]" pageId="31" pageNumber="182">
<emphasis id="B938EAFEFFB8FFBFFF2A5A912C0373B2" bold="true" box="[151,272,1410,1436]" pageId="31" pageNumber="182">TABLE 3.</emphasis>
Length of the antennomeres. All values in mm.
</paragraph>
</caption>
<paragraph id="8BF336ECFFB8FFBFFF1B5ABE289471AE" pageId="31" pageNumber="182">
<table id="F94CC44CFFB8005FFF1B5ABF289471AE" box="[166,1415,1452,1920]" gridcols="12" gridrows="12" pageId="31" pageNumber="182">
<tr id="357C34AEFFB8005FFF1B5ABF289473EB" box="[166,1415,1452,1477]" gridrow="0" pageId="31" pageNumber="182" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-8="1" rowspan-9="1">
<th id="76AD5DD2FFB8005FFF1B5ABF2C9873EB" box="[166,395,1452,1477]" gridcol="0" gridrow="0" pageId="31" pageNumber="182">Species</th>
<th id="76AD5DD2FFB8005FFCF45ABF293773EB" box="[841,1060,1452,1477]" colspan="3" colspanRight="2" gridcol="5" gridrow="0" pageId="31" pageNumber="182">
<emphasis id="B938EAFEFFB8FFBFFCD65ABF291B73EA" bold="true" box="[875,1032,1452,1476]" pageId="31" pageNumber="182">Antennomeres</emphasis>
</th>
</tr>
<tr id="357C34AEFFB8005FFF1B5AC6289473C3" box="[166,1415,1493,1517]" gridrow="1" pageId="31" pageNumber="182" rowspan-0="1">
<td id="76AD5DD2FFB8005FFE565AC62F0473C3" box="[491,535,1493,1517]" gridcol="1" gridrow="1" pageId="31" pageNumber="182">I</td>
<td id="76AD5DD2FFB8005FFDFE5AC62F7C73C3" box="[579,623,1493,1517]" gridcol="2" gridrow="1" pageId="31" pageNumber="182">II</td>
<td id="76AD5DD2FFB8005FFD275AC62FD573C3" box="[666,710,1493,1517]" gridcol="3" gridrow="1" pageId="31" pageNumber="182">III</td>
<td id="76AD5DD2FFB8005FFD4F5AC62E0D73C3" box="[754,798,1493,1517]" gridcol="4" gridrow="1" pageId="31" pageNumber="182">IV</td>
<td id="76AD5DD2FFB8005FFCF45AC62E6673C3" box="[841,885,1493,1517]" gridcol="5" gridrow="1" pageId="31" pageNumber="182">V</td>
<td id="76AD5DD2FFB8005FFC1C5AC62EDE73C3" box="[929,973,1493,1517]" gridcol="6" gridrow="1" pageId="31" pageNumber="182">VI</td>
<td id="76AD5DD2FFB8005FFC455AC6293773C3" box="[1016,1060,1493,1517]" gridcol="7" gridrow="1" pageId="31" pageNumber="182">VII</td>
<td id="76AD5DD2FFB8005FFBEF5AC6296D73C3" box="[1106,1150,1493,1517]" gridcol="8" gridrow="1" pageId="31" pageNumber="182">VIII</td>
<td id="76AD5DD2FFB8005FFB115AC629CB73C3" box="[1196,1240,1493,1517]" gridcol="9" gridrow="1" pageId="31" pageNumber="182">IX</td>
<td id="76AD5DD2FFB8005FFABE5AC6283C73C3" box="[1283,1327,1493,1517]" gridcol="10" gridrow="1" pageId="31" pageNumber="182">X</td>
<td id="76AD5DD2FFB8005FFAE65AC6289473C3" box="[1371,1415,1493,1517]" gridcol="11" gridrow="1" pageId="31" pageNumber="182">
<emphasis id="B938EAFEFFB8FFBFFADE5AC6286C73C3" bold="true" box="[1379,1407,1493,1517]" pageId="31" pageNumber="182">XI</emphasis>
</td>
</tr>
<tr id="357C34AEFFB8005FFF1B5AEE28947036" box="[166,1415,1533,1560]" gridrow="2" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B5AEE2C987036" box="[166,395,1533,1560]" gridcol="0" gridrow="2" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B5AED2C19703B" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[166,266,1533,1557]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="sendrai">
<emphasis id="B938EAFEFFB8FFBFFF1B5AED2C19703B" box="[166,266,1533,1557]" italics="true" pageId="31" pageNumber="182">T. sendrai</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB8FFBFFEAC5AEE2C72703B" bold="true" box="[273,353,1533,1557]" pageId="31" pageNumber="182">
sp. no 
<emphasis id="B938EAFEFFB8FFBFFEED5AED2C72703B" bold="true" box="[336,353,1534,1557]" italics="true" pageId="31" pageNumber="182">v.</emphasis>
</emphasis>
ϐ
</th>
<td id="76AD5DD2FFB8005FFE565AEE2F047036" box="[491,535,1533,1560]" gridcol="1" gridrow="2" pageId="31" pageNumber="182">0.28</td>
<td id="76AD5DD2FFB8005FFDFE5AEE2F7C7036" box="[579,623,1533,1560]" gridcol="2" gridrow="2" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFD275AEE2FD57036" box="[666,710,1533,1560]" gridcol="3" gridrow="2" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFD4F5AEE2E0D7036" box="[754,798,1533,1560]" gridcol="4" gridrow="2" pageId="31" pageNumber="182">0.26</td>
<td id="76AD5DD2FFB8005FFCF45AEE2E667036" box="[841,885,1533,1560]" gridcol="5" gridrow="2" pageId="31" pageNumber="182">0.31</td>
<td id="76AD5DD2FFB8005FFC1C5AEE2EDE7036" box="[929,973,1533,1560]" gridcol="6" gridrow="2" pageId="31" pageNumber="182">0.29</td>
<td id="76AD5DD2FFB8005FFC455AEE29377036" box="[1016,1060,1533,1560]" gridcol="7" gridrow="2" pageId="31" pageNumber="182">0.29</td>
<td id="76AD5DD2FFB8005FFBEF5AEE296D7036" box="[1106,1150,1533,1560]" gridcol="8" gridrow="2" pageId="31" pageNumber="182">0.19</td>
<td id="76AD5DD2FFB8005FFB115AEE29CB7036" box="[1196,1240,1533,1560]" gridcol="9" gridrow="2" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFABE5AEE283C7036" box="[1283,1327,1533,1560]" gridcol="10" gridrow="2" pageId="31" pageNumber="182">0.18</td>
<td id="76AD5DD2FFB8005FFAE65AEE28947036" box="[1371,1415,1533,1560]" gridcol="11" gridrow="2" pageId="31" pageNumber="182">0.25</td>
</tr>
<tr id="357C34AEFFB8005FFF1B593628947010" box="[166,1415,1573,1598]" gridrow="3" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B59362C987010" box="[166,395,1573,1598]" gridcol="0" gridrow="3" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B59352C197013" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[166,266,1573,1597]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="sendrai">
<emphasis id="B938EAFEFFB8FFBFFF1B59352C197013" box="[166,266,1573,1597]" italics="true" pageId="31" pageNumber="182">T. sendrai</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB8FFBFFEAC59352C727013" bold="true" box="[273,353,1574,1598]" pageId="31" pageNumber="182">
sp. no 
<emphasis id="B938EAFEFFB8FFBFFEED59352C727013" bold="true" box="[336,353,1574,1597]" italics="true" pageId="31" pageNumber="182">v.</emphasis>
</emphasis>
♀
</th>
<td id="76AD5DD2FFB8005FFE5659362F047010" box="[491,535,1573,1598]" gridcol="1" gridrow="3" pageId="31" pageNumber="182">0.28</td>
<td id="76AD5DD2FFB8005FFDFE59362F7C7010" box="[579,623,1573,1598]" gridcol="2" gridrow="3" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD2759362FD57010" box="[666,710,1573,1598]" gridcol="3" gridrow="3" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFD4F59362E0D7010" box="[754,798,1573,1598]" gridcol="4" gridrow="3" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFCF459362E667010" box="[841,885,1573,1598]" gridcol="5" gridrow="3" pageId="31" pageNumber="182">0.25</td>
<td id="76AD5DD2FFB8005FFC1C59362EDE7010" box="[929,973,1573,1598]" gridcol="6" gridrow="3" pageId="31" pageNumber="182">0.23</td>
<td id="76AD5DD2FFB8005FFC45593629377010" box="[1016,1060,1573,1598]" gridcol="7" gridrow="3" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFBEF5936296D7010" box="[1106,1150,1573,1598]" gridcol="8" gridrow="3" pageId="31" pageNumber="182">0.15</td>
<td id="76AD5DD2FFB8005FFB11593629CB7010" box="[1196,1240,1573,1598]" gridcol="9" gridrow="3" pageId="31" pageNumber="182">0.17</td>
<td id="76AD5DD2FFB8005FFABE5936283C7010" box="[1283,1327,1573,1598]" gridcol="10" gridrow="3" pageId="31" pageNumber="182">0.15</td>
<td id="76AD5DD2FFB8005FFAE6593628947010" box="[1371,1415,1573,1598]" gridcol="11" gridrow="3" pageId="31" pageNumber="182">0.23</td>
</tr>
<tr id="357C34AEFFB8005FFF1B595D28947046" box="[166,1415,1614,1640]" gridrow="4" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B595D2C987046" box="[166,395,1614,1640]" gridcol="0" gridrow="4" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B595D2C087048" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[166,283,1614,1638]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="fadriquei">
<emphasis id="B938EAFEFFB8FFBFFF1B595D2C087048" box="[166,283,1614,1638]" italics="true" pageId="31" pageNumber="182">T. fadriquei</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB8FFBFFE9F595D2C61704B" bold="true" box="[290,370,1614,1638]" pageId="31" pageNumber="182">
sp. no 
<emphasis id="B938EAFEFFB8FFBFFEDC595D2C61704B" bold="true" box="[353,370,1614,1637]" italics="true" pageId="31" pageNumber="182">v.</emphasis>
</emphasis>
ϐ
</th>
<td id="76AD5DD2FFB8005FFE56595D2F047046" box="[491,535,1614,1640]" gridcol="1" gridrow="4" pageId="31" pageNumber="182">0.27</td>
<td id="76AD5DD2FFB8005FFDFE595D2F7C7046" box="[579,623,1614,1640]" gridcol="2" gridrow="4" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD27595D2FD57046" box="[666,710,1614,1640]" gridcol="3" gridrow="4" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFD4F595D2E0D7046" box="[754,798,1614,1640]" gridcol="4" gridrow="4" pageId="31" pageNumber="182">0.27</td>
<td id="76AD5DD2FFB8005FFCF4595D2E667046" box="[841,885,1614,1640]" gridcol="5" gridrow="4" pageId="31" pageNumber="182">0.32</td>
<td id="76AD5DD2FFB8005FFC1C595D2EDE7046" box="[929,973,1614,1640]" gridcol="6" gridrow="4" pageId="31" pageNumber="182">0.29</td>
<td id="76AD5DD2FFB8005FFC45595D29377046" box="[1016,1060,1614,1640]" gridcol="7" gridrow="4" pageId="31" pageNumber="182">0.33</td>
<td id="76AD5DD2FFB8005FFBEF595D296D7046" box="[1106,1150,1614,1640]" gridcol="8" gridrow="4" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFB11595D29CB7046" box="[1196,1240,1614,1640]" gridcol="9" gridrow="4" pageId="31" pageNumber="182">0.23</td>
<td id="76AD5DD2FFB8005FFABE595D283C7046" box="[1283,1327,1614,1640]" gridcol="10" gridrow="4" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFAE6595D28947046" box="[1371,1415,1614,1640]" gridcol="11" gridrow="4" pageId="31" pageNumber="182">0.25</td>
</tr>
<tr id="357C34AEFFB8005FFF1B5965289470A0" box="[166,1415,1654,1678]" gridrow="5" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B59652C9870A0" box="[166,395,1654,1678]" gridcol="0" gridrow="5" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B59642C0870A0" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[166,283,1654,1678]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="fadriquei">
<emphasis id="B938EAFEFFB8FFBFFF1B59642C0870A0" box="[166,283,1654,1678]" italics="true" pageId="31" pageNumber="182">T. fadriquei</emphasis>
</taxonomicName>
<emphasis id="B938EAFEFFB8FFBFFE9F59652C6170A3" bold="true" box="[290,370,1654,1678]" pageId="31" pageNumber="182">
sp. no 
<emphasis id="B938EAFEFFB8FFBFFEDC59652C6170A3" bold="true" box="[353,370,1654,1677]" italics="true" pageId="31" pageNumber="182">v.</emphasis>
</emphasis>
♀
</th>
<td id="76AD5DD2FFB8005FFE5659652F0470A0" box="[491,535,1654,1678]" gridcol="1" gridrow="5" pageId="31" pageNumber="182">0.28</td>
<td id="76AD5DD2FFB8005FFDFE59652F7C70A0" box="[579,623,1654,1678]" gridcol="2" gridrow="5" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFD2759652FD570A0" box="[666,710,1654,1678]" gridcol="3" gridrow="5" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD4F59652E0D70A0" box="[754,798,1654,1678]" gridcol="4" gridrow="5" pageId="31" pageNumber="182">0.26</td>
<td id="76AD5DD2FFB8005FFCF459652E6670A0" box="[841,885,1654,1678]" gridcol="5" gridrow="5" pageId="31" pageNumber="182">0.30</td>
<td id="76AD5DD2FFB8005FFC1C59652EDE70A0" box="[929,973,1654,1678]" gridcol="6" gridrow="5" pageId="31" pageNumber="182">0.25</td>
<td id="76AD5DD2FFB8005FFC455965293770A0" box="[1016,1060,1654,1678]" gridcol="7" gridrow="5" pageId="31" pageNumber="182">0.27</td>
<td id="76AD5DD2FFB8005FFBEF5965296D70A0" box="[1106,1150,1654,1678]" gridcol="8" gridrow="5" pageId="31" pageNumber="182">0.15</td>
<td id="76AD5DD2FFB8005FFB11596529CB70A0" box="[1196,1240,1654,1678]" gridcol="9" gridrow="5" pageId="31" pageNumber="182">0.19</td>
<td id="76AD5DD2FFB8005FFABE5965283C70A0" box="[1283,1327,1654,1678]" gridcol="10" gridrow="5" pageId="31" pageNumber="182">0.18</td>
<td id="76AD5DD2FFB8005FFAE65965289470A0" box="[1371,1415,1654,1678]" gridcol="11" gridrow="5" pageId="31" pageNumber="182">0.24</td>
</tr>
<tr id="357C34AEFFB8005FFF1B598D28947097" box="[166,1415,1694,1721]" gridrow="6" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B598D2C987097" box="[166,395,1694,1721]" gridcol="0" gridrow="6" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B598C2C1D7098" box="[166,270,1694,1718]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="mengeli">
<emphasis id="B938EAFEFFB8FFBFFF1B598C2C1D7098" box="[166,270,1694,1718]" italics="true" pageId="31" pageNumber="182">T. mengeli</emphasis>
</taxonomicName>
ϐ
</th>
<td id="76AD5DD2FFB8005FFE56598D2F047097" box="[491,535,1694,1721]" gridcol="1" gridrow="6" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFDFE598D2F7C7097" box="[579,623,1694,1721]" gridcol="2" gridrow="6" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD27598D2FD57097" box="[666,710,1694,1721]" gridcol="3" gridrow="6" pageId="31" pageNumber="182">0.17</td>
<td id="76AD5DD2FFB8005FFD4F598D2E0D7097" box="[754,798,1694,1721]" gridcol="4" gridrow="6" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFCF4598D2E667097" box="[841,885,1694,1721]" gridcol="5" gridrow="6" pageId="31" pageNumber="182">0.21</td>
<td id="76AD5DD2FFB8005FFC1C598D2EDE7097" box="[929,973,1694,1721]" gridcol="6" gridrow="6" pageId="31" pageNumber="182">0.19</td>
<td id="76AD5DD2FFB8005FFC45598D29377097" box="[1016,1060,1694,1721]" gridcol="7" gridrow="6" pageId="31" pageNumber="182">0.19</td>
<td id="76AD5DD2FFB8005FFBEF598D296D7097" box="[1106,1150,1694,1721]" gridcol="8" gridrow="6" pageId="31" pageNumber="182">0.11</td>
<td id="76AD5DD2FFB8005FFB11598D29CB7097" box="[1196,1240,1694,1721]" gridcol="9" gridrow="6" pageId="31" pageNumber="182">0.13</td>
<td id="76AD5DD2FFB8005FFABE598D283C7097" box="[1283,1327,1694,1721]" gridcol="10" gridrow="6" pageId="31" pageNumber="182">0.13</td>
<td id="76AD5DD2FFB8005FFAE6598D28947097" box="[1371,1415,1694,1721]" gridcol="11" gridrow="6" pageId="31" pageNumber="182">0.22</td>
</tr>
<tr id="357C34AEFFB8005FFF1B59D5289470F1" box="[166,1415,1734,1759]" gridrow="7" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B59D52C9870F1" box="[166,395,1734,1759]" gridcol="0" gridrow="7" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B59D42C1D70F0" box="[166,270,1734,1758]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="mengeli">
<emphasis id="B938EAFEFFB8FFBFFF1B59D42C1D70F0" box="[166,270,1734,1758]" italics="true" pageId="31" pageNumber="182">T. mengeli</emphasis>
</taxonomicName>
♀
</th>
<td id="76AD5DD2FFB8005FFE5659D52F0470F1" box="[491,535,1734,1759]" gridcol="1" gridrow="7" pageId="31" pageNumber="182">0.23</td>
<td id="76AD5DD2FFB8005FFDFE59D52F7C70F1" box="[579,623,1734,1759]" gridcol="2" gridrow="7" pageId="31" pageNumber="182">0.21</td>
<td id="76AD5DD2FFB8005FFD2759D52FD570F1" box="[666,710,1734,1759]" gridcol="3" gridrow="7" pageId="31" pageNumber="182">0.17</td>
<td id="76AD5DD2FFB8005FFD4F59D52E0D70F1" box="[754,798,1734,1759]" gridcol="4" gridrow="7" pageId="31" pageNumber="182">0.18</td>
<td id="76AD5DD2FFB8005FFCF459D52E6670F1" box="[841,885,1734,1759]" gridcol="5" gridrow="7" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFC1C59D52EDE70F1" box="[929,973,1734,1759]" gridcol="6" gridrow="7" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFC4559D5293770F1" box="[1016,1060,1734,1759]" gridcol="7" gridrow="7" pageId="31" pageNumber="182">0.19</td>
<td id="76AD5DD2FFB8005FFBEF59D5296D70F1" box="[1106,1150,1734,1759]" gridcol="8" gridrow="7" pageId="31" pageNumber="182">0.12</td>
<td id="76AD5DD2FFB8005FFB1159D529CB70F1" box="[1196,1240,1734,1759]" gridcol="9" gridrow="7" pageId="31" pageNumber="182">0.14</td>
<td id="76AD5DD2FFB8005FFABE59D5283C70F1" box="[1283,1327,1734,1759]" gridcol="10" gridrow="7" pageId="31" pageNumber="182">0.12</td>
<td id="76AD5DD2FFB8005FFAE659D5289470F1" box="[1371,1415,1734,1759]" gridcol="11" gridrow="7" pageId="31" pageNumber="182">0.20</td>
</tr>
<tr id="357C34AEFFB8005FFF1B59FC28947127" box="[166,1415,1775,1801]" gridrow="8" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B59FC2C987127" box="[166,395,1775,1801]" gridcol="0" gridrow="8" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B59E32C377129" box="[166,292,1775,1799]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="mercedesi">
<emphasis id="B938EAFEFFB8FFBFFF1B59E32C377129" box="[166,292,1775,1799]" italics="true" pageId="31" pageNumber="182">T. mercedesi</emphasis>
</taxonomicName>
ϐ
</th>
<td id="76AD5DD2FFB8005FFE5659FC2F047127" box="[491,535,1775,1801]" gridcol="1" gridrow="8" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFDFE59FC2F7C7127" box="[579,623,1775,1801]" gridcol="2" gridrow="8" pageId="31" pageNumber="182">0.23</td>
<td id="76AD5DD2FFB8005FFD2759FC2FD57127" box="[666,710,1775,1801]" gridcol="3" gridrow="8" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD4F59FC2E0D7127" box="[754,798,1775,1801]" gridcol="4" gridrow="8" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFCF459FC2E667127" box="[841,885,1775,1801]" gridcol="5" gridrow="8" pageId="31" pageNumber="182">0.28</td>
<td id="76AD5DD2FFB8005FFC1C59FC2EDE7127" box="[929,973,1775,1801]" gridcol="6" gridrow="8" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFC4559FC29377127" box="[1016,1060,1775,1801]" gridcol="7" gridrow="8" pageId="31" pageNumber="182">0.24</td>
<td id="76AD5DD2FFB8005FFBEF59FC296D7127" box="[1106,1150,1775,1801]" gridcol="8" gridrow="8" pageId="31" pageNumber="182">0.13</td>
<td id="76AD5DD2FFB8005FFB1159FC29CB7127" box="[1196,1240,1775,1801]" gridcol="9" gridrow="8" pageId="31" pageNumber="182">0.19</td>
<td id="76AD5DD2FFB8005FFABE59FC283C7127" box="[1283,1327,1775,1801]" gridcol="10" gridrow="8" pageId="31" pageNumber="182">0.17</td>
<td id="76AD5DD2FFB8005FFAE659FC28947127" box="[1371,1415,1775,1801]" gridcol="11" gridrow="8" pageId="31" pageNumber="182">0.23</td>
</tr>
<tr id="357C34AEFFB8005FFF1B580428947101" box="[166,1415,1815,1839]" gridrow="9" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B58042C987101" box="[166,395,1815,1839]" gridcol="0" gridrow="9" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B580B2C377101" box="[166,292,1815,1839]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="mercedesi">
<emphasis id="B938EAFEFFB8FFBFFF1B580B2C377101" box="[166,292,1815,1839]" italics="true" pageId="31" pageNumber="182">T. mercedesi</emphasis>
</taxonomicName>
♀
</th>
<td id="76AD5DD2FFB8005FFE5658042F047101" box="[491,535,1815,1839]" gridcol="1" gridrow="9" pageId="31" pageNumber="182">0.21</td>
<td id="76AD5DD2FFB8005FFDFE58042F7C7101" box="[579,623,1815,1839]" gridcol="2" gridrow="9" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFD2758042FD57101" box="[666,710,1815,1839]" gridcol="3" gridrow="9" pageId="31" pageNumber="182">0.19</td>
<td id="76AD5DD2FFB8005FFD4F58042E0D7101" box="[754,798,1815,1839]" gridcol="4" gridrow="9" pageId="31" pageNumber="182">0.19</td>
<td id="76AD5DD2FFB8005FFCF458042E667101" box="[841,885,1815,1839]" gridcol="5" gridrow="9" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFC1C58042EDE7101" box="[929,973,1815,1839]" gridcol="6" gridrow="9" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFC45580429377101" box="[1016,1060,1815,1839]" gridcol="7" gridrow="9" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFBEF5804296D7101" box="[1106,1150,1815,1839]" gridcol="8" gridrow="9" pageId="31" pageNumber="182">0.12</td>
<td id="76AD5DD2FFB8005FFB11580429CB7101" box="[1196,1240,1815,1839]" gridcol="9" gridrow="9" pageId="31" pageNumber="182">0.15</td>
<td id="76AD5DD2FFB8005FFABE5804283C7101" box="[1283,1327,1815,1839]" gridcol="10" gridrow="9" pageId="31" pageNumber="182">0.14</td>
<td id="76AD5DD2FFB8005FFAE6580428947101" box="[1371,1415,1815,1839]" gridcol="11" gridrow="9" pageId="31" pageNumber="182">0.22</td>
</tr>
<tr id="357C34AEFFB8005FFF1B582C28947174" box="[166,1415,1855,1882]" gridrow="10" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B582C2C987174" box="[166,395,1855,1882]" gridcol="0" gridrow="10" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B58532C157179" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[166,262,1856,1879]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB8FFBFFF1B58532C157179" box="[166,262,1856,1879]" italics="true" pageId="31" pageNumber="182">T. vinyasi</emphasis>
</taxonomicName>
ϐ
</th>
<td id="76AD5DD2FFB8005FFE56582C2F047174" box="[491,535,1855,1882]" gridcol="1" gridrow="10" pageId="31" pageNumber="182">0.27</td>
<td id="76AD5DD2FFB8005FFDFE582C2F7C7174" box="[579,623,1855,1882]" gridcol="2" gridrow="10" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD27582C2FD57174" box="[666,710,1855,1882]" gridcol="3" gridrow="10" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD4F582C2E0D7174" box="[754,798,1855,1882]" gridcol="4" gridrow="10" pageId="31" pageNumber="182">0.25</td>
<td id="76AD5DD2FFB8005FFCF4582C2E667174" box="[841,885,1855,1882]" gridcol="5" gridrow="10" pageId="31" pageNumber="182">0.30</td>
<td id="76AD5DD2FFB8005FFC1C582C2EDE7174" box="[929,973,1855,1882]" gridcol="6" gridrow="10" pageId="31" pageNumber="182">0.27</td>
<td id="76AD5DD2FFB8005FFC45582C29377174" box="[1016,1060,1855,1882]" gridcol="7" gridrow="10" pageId="31" pageNumber="182">0.30</td>
<td id="76AD5DD2FFB8005FFBEF582C296D7174" box="[1106,1150,1855,1882]" gridcol="8" gridrow="10" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFB11582C29CB7174" box="[1196,1240,1855,1882]" gridcol="9" gridrow="10" pageId="31" pageNumber="182">0.21</td>
<td id="76AD5DD2FFB8005FFABE582C283C7174" box="[1283,1327,1855,1882]" gridcol="10" gridrow="10" pageId="31" pageNumber="182">0.20</td>
<td id="76AD5DD2FFB8005FFAE6582C28947174" box="[1371,1415,1855,1882]" gridcol="11" gridrow="10" pageId="31" pageNumber="182">0.24</td>
</tr>
<tr id="357C34AEFFB8005FFF1B5874289471AE" box="[166,1415,1895,1920]" gridrow="11" pageId="31" pageNumber="182">
<th id="76AD5DD2FFB8005FFF1B58742C9871AE" box="[166,395,1895,1920]" gridcol="0" gridrow="11" pageId="31" pageNumber="182">
<taxonomicName id="4C4C4D6FFFB8FFBFFF1B587B2C157151" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[166,262,1896,1919]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFB8FFBFFF1B587B2C157151" box="[166,262,1896,1919]" italics="true" pageId="31" pageNumber="182">T. vinyasi</emphasis>
</taxonomicName>
♀
</th>
<td id="76AD5DD2FFB8005FFE5658742F0471AE" box="[491,535,1895,1920]" gridcol="1" gridrow="11" pageId="31" pageNumber="182">0.26</td>
<td id="76AD5DD2FFB8005FFDFE58742F7C71AE" box="[579,623,1895,1920]" gridcol="2" gridrow="11" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD2758742FD571AE" box="[666,710,1895,1920]" gridcol="3" gridrow="11" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFD4F58742E0D71AE" box="[754,798,1895,1920]" gridcol="4" gridrow="11" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFCF458742E6671AE" box="[841,885,1895,1920]" gridcol="5" gridrow="11" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFC1C58742EDE71AE" box="[929,973,1895,1920]" gridcol="6" gridrow="11" pageId="31" pageNumber="182">0.28</td>
<td id="76AD5DD2FFB8005FFC455874293771AE" box="[1016,1060,1895,1920]" gridcol="7" gridrow="11" pageId="31" pageNumber="182">0.28</td>
<td id="76AD5DD2FFB8005FFBEF5874296D71AE" box="[1106,1150,1895,1920]" gridcol="8" gridrow="11" pageId="31" pageNumber="182">0.16</td>
<td id="76AD5DD2FFB8005FFB11587429CB71AE" box="[1196,1240,1895,1920]" gridcol="9" gridrow="11" pageId="31" pageNumber="182">0.22</td>
<td id="76AD5DD2FFB8005FFABE5874283C71AE" box="[1283,1327,1895,1920]" gridcol="10" gridrow="11" pageId="31" pageNumber="182">0.18</td>
<td id="76AD5DD2FFB8005FFAE65874289471AE" box="[1371,1415,1895,1920]" gridcol="11" gridrow="11" pageId="31" pageNumber="182">0.20</td>
</tr>
</table>
</paragraph>
<caption id="DF336664FF87FF80FF2A5B8F2F93729A" ID-DOI="http://doi.org/10.5281/zenodo.4505974" ID-Zenodo-Dep="4505974" box="[151,640,1180,1205]" httpUri="https://zenodo.org/record/4505974/files/figure.png" pageId="32" pageNumber="183" startId="32.[151,250,1180,1204]" targetBox="[152,1435,181,1156]" targetPageId="32">
<paragraph id="8BF336ECFF87FF80FF2A5B8F2F93729A" blockId="32.[151,640,1180,1205]" box="[151,640,1180,1205]" pageId="32" pageNumber="183">
<emphasis id="B938EAFEFF87FF80FF2A5B8F2C33729B" bold="true" box="[151,288,1180,1205]" pageId="32" pageNumber="183">FIGURE 70.</emphasis>
Lectotypus of 
<taxonomicName id="4C4C4D6FFF87FF80FE055B8E2F69729A" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" box="[440,634,1181,1204]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFF87FF80FE055B8E2F69729A" box="[440,634,1181,1204]" italics="true" pageId="32" pageNumber="183">Speonomus vinyasi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF336ECFF87FF80FF7A5BF02CDF730F" blockId="32.[151,1437,1251,1601]" pageId="32" pageNumber="183">
Intersal sac of the aedeagus (
<figureCitation id="13772A69FF87FF80FDAC5BF02F7072D3" box="[529,611,1251,1277]" captionStart="FIGURES 20–24" captionStartId="22.[151,264,1951,1975]" captionTargetBox="[252,1333,182,1914]" captionTargetId="figure-24@22.[251,1336,181,1927]" captionTargetPageId="22" captionText="FIGURES 20–24. Aedeagus in dorsal view of Troglocharinus sendrai sp. nov. (20), T. fadriquei sp. nov. (21), T. mengeli (22)—Cabana d’en Garraba—, T. mercedesi (23)—Bòfia de Sant Jaume—and T. vinyasi (24)—Avenc de la Roca del Corb." figureDoi="http://doi.org/10.5281/zenodo.4505958" httpUri="https://zenodo.org/record/4505958/files/figure.png" pageId="32" pageNumber="183">Fig. 24</figureCitation>
) with very large NCs; NFVM deeply notched, larger branch directed towards apex; PF2 triangular.
</paragraph>
<paragraph id="8BF336ECFF87FF80FF7A5A382DD773A0" blockId="32.[151,1437,1251,1601]" pageId="32" pageNumber="183">
<emphasis id="B938EAFEFF87FF80FF7A5A382C36736B" bold="true" box="[199,293,1323,1349]" pageId="32" pageNumber="183">Female.</emphasis>
Protarsi not dilated, filiform; antennae shorter. Spiculum of the urite viii short, wide and with truncated apex (
<figureCitation id="13772A69FF87FF80FF635A5C2C277344" box="[222,308,1359,1386]" captionStart="FIGURES 58–63" captionStartId="29.[151,264,1784,1808]" captionTargetBox="[151,1436,489,1757]" captionTargetId="figure-128@29.[151,1436,489,1760]" captionTargetPageId="29" captionText="FIGURES 58–63. viii female urite of Troglocharinus sendrai sp. nov. (58), T. fadriquei sp. nov. (59), T. mengeli (60)—Coves Encantades de Toloriu—, T. mercedesi (61)—Bòfia de Sant Jaume—, T. vinyasi (62)—Avenc de l’Home Mort—and Speonomites velox (63)—Querant del riu Merlé, Vilanova de Meià—. Spermathecal complex of T. mercedesi: CE, spermaduct; E, spermatheca; LA, apical lobe; LBS, basal lobe; RM middle region; VIII, eight urite." figureDoi="http://doi.org/10.5281/zenodo.4505970" httpUri="https://zenodo.org/record/4505970/files/figure.png" pageId="32" pageNumber="183">Fig. 62</figureCitation>
). Lobes of the spermatheca ony slightly wider than median region, which is strongly bended (
<figureCitation id="13772A69FF87FF80FAD25A5C2DA473A0" captionStart="FIGURES 52–57" captionStartId="28.[151,264,1907,1931]" captionTargetBox="[151,1436,527,1884]" captionTargetId="figure-144@28.[151,1436,525,1884]" captionTargetPageId="28" captionText="FIGURES 52–57. Spermatheca of Troglocharinus sendrai sp. nov. (52), T. fadriquei sp. nov. (53), T. mengeli (54)—Coves Encantades de Toloriu—, T. mercedesi (55)—Bòfia de Sant Jaume—, T. mercedesi (56)—Avenc del Xato—and T. vinyasi (57)—Avenc de l’Home Mort." figureDoi="http://doi.org/10.5281/zenodo.4505968" httpUri="https://zenodo.org/record/4505968/files/figure.png" pageId="32" pageNumber="183">Fig. 57</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3566567FF87FF80FF7A5A842F857137" pageId="32" pageNumber="183" type="distribution">
<paragraph id="8BF336ECFF87FF80FF7A5A842CD97033" blockId="32.[151,1437,1251,1601]" pageId="32" pageNumber="183">
<emphasis id="B938EAFEFF87FF80FF7A5A842C4A739F" bold="true" box="[199,345,1431,1457]" pageId="32" pageNumber="183">Distribution</emphasis>
(
<figureCitation id="13772A69FF87FF80FED45A842CAE739F" box="[361,445,1431,1457]" captionStart="FIGURE 71" captionStartId="33.[151,250,1993,2017]" captionTargetBox="[212,1380,187,1961]" captionTargetId="figure-17@33.[195,1393,174,1969]" captionTargetPageId="33" captionText="FIGURE 71. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.4505976" httpUri="https://zenodo.org/record/4505976/files/figure.png" pageId="32" pageNumber="183">Fig. 71</figureCitation>
). Known from several caves in Serra de l’Arcosa, right margin of the river Segre, in the surrounding area of Peramola, Lleida (
<bibRefCitation id="EFDD4B1DFF87FF80FDF35AA82FCE73FB" author="Escola, O." box="[590,733,1467,1493]" pageId="32" pageNumber="183" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà 1971</bibRefCitation>
, 
<bibRefCitation id="EFDD4B1DFF87FF80FD575AA82E3673FB" author="Escola, O." box="[746,805,1467,1493]" pageId="32" pageNumber="183" pagination="91 - 93" refId="ref17452" refString="Escola, O. (1973) Addenda a &quot; un nuevo Speonomus de la provincia de Lerida &quot;. Speleon, 20, 91 - 93." type="journal article" year="1973">1973</bibRefCitation>
, 
<bibRefCitation id="EFDD4B1DFF87FF80FC8F5AA82E7E73FB" author="Escola, O." box="[818,877,1467,1493]" pageId="32" pageNumber="183" pagination="141 - 147" refId="ref17480" refString="Escola, O. (1980) Sur la coexistence des Bathysciinae dans les grottes d'Espagne. Memories de Biospeologie, 7, 141 - 147." type="journal article" year="1980">1980</bibRefCitation>
, 
<bibRefCitation id="EFDD4B1DFF87FF80FCC45AA82EA773FB" author="Escola, O." box="[889,948,1467,1493]" pageId="32" pageNumber="183" pagination="129 - 133" refId="ref17507" refString="Escola, O. (1986) Updating of Bathysciinae of the Iberian Peninsula (Col. Catopidae). Comunicacions del 9 Congres Internacional d'Espeleologia, Barcelona, 2, 129 - 133." type="journal article" year="1986">1986</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FC7C5AA8294C73FB" author="Espanol, F." box="[961,1119,1467,1493]" pageId="32" pageNumber="183" refId="ref17690" refString="Espanol, F. (1976) Invertebrats cavernicoles dels Paisos Catalans. In: Natura us o abus? Institucio Catalana d'Historia Natural. Vol. 9. Ed. Barcino, Barcelona, pp. 230 - 236" type="book" year="1976">Español 1976</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FBD15AA8281273FB" author="Belles, X." box="[1132,1281,1467,1493]" pageId="32" pageNumber="183" pagination="114 - 121" refId="ref16828" refString="Belles, X. (1978 a) Notes biospeleologiques. In: Borras, J., Minarro, J. M. &amp; Talavera, F. Cataleg espeleologic de Catalunya, 1, L'Alt Urgell i la Cerdanya. Ed. Poliglota, Barcelona, pp. 23 - 33, 114 - 121." type="journal article" year="1978">Bellés 1978a</bibRefCitation>
, 
<bibRefCitation id="EFDD4B1DFF87FF80FAB35AA8285A73FB" author="Belles, X." box="[1294,1353,1467,1493]" pageId="32" pageNumber="183" pagination="1 - 207" refId="ref17009" refString="Belles, X. (1987) Fauna cavernicola i intersticial de la Peninsula Iberica i les illes Balears. C. S. I. C. Monografies Cientifiques, 4, 1 - 207." type="journal article" year="1987">1987</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FAEB5AA82C5073D7" author="Belles, X. &amp; Escola, O." pageId="32" pageNumber="183" pagination="569 - 572" refId="ref17158" refString="Belles, X. &amp; Escola, O. (1979) Addicions al cataleg de Bathysciinae (Col. Catopidae) iberics d'enca la classificacio de Laneyrie (1967). Espeleoleg, 28, 569 - 572." type="journal article" year="1979">Bellés &amp; Escolà 1979</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FEED5ACC2F5673D7" author="Belles, X. &amp; Deliot, Ph." box="[336,581,1503,1529]" pageId="32" pageNumber="183" pagination="237 - 243" refId="ref17124" refString="Belles, X. &amp; Deliot, Ph. (1983) Nouveaux Bathysciinae (Coleopteres Catopidae) des Pyrenees francaises et espagnoles. Memoires de Biospeologie, 10, 237 - 243." type="journal article" year="1983">Bellés &amp; Déliot 1983</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FDEF5ACC2FE973D7" author="Fresneda, J." box="[594,762,1503,1529]" pageId="32" pageNumber="183" pagination="53 - 86" refId="ref17908" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda 1998</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FCBA5ACC2EB273D7" author="Perreau, M." box="[775,929,1503,1529]" pageId="32" pageNumber="183" pagination="1 - 460" refId="ref18493" refString="Perreau, M. (2000) Catalogue des Coleopteres Leiodidae Cholevinae et Platypsyllinae. Memoires de la Societe entomologique de France, 4, 1 - 460." type="journal article" year="2000">Perreau 2000</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FC105ACC29BE73D7" author="Lobl, I. &amp; Smetana A." box="[941,1197,1503,1529]" pageId="32" pageNumber="183" refId="ref18428" refString="Lobl, I. &amp; Smetana A. (2004) Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Histeroidea-Staphylinoidea. Apollo Books, Stenstrup. 942 pp." type="book" year="2004">Löbl &amp; Smetana 2004</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FB045ACC288473D7" author="Salgado, J. M. &amp; Blas, M. &amp; Fresneda, J." box="[1209,1431,1503,1529]" pageId="32" pageNumber="183" refId="ref18846" refString="Salgado, J. M., Blas, M. &amp; Fresneda, J. (2008) Coleoptera, Cholevidae. In: Fauna Iberica. Vol. 31. Museo Nacional de Ciencias Naturales, CSIC, Madrid, 799 pp." type="book" year="2008">
Salgado 
<emphasis id="B938EAFEFF87FF80FAA65AF3285E73D7" box="[1307,1357,1503,1529]" italics="true" pageId="32" pageNumber="183">et al</emphasis>
. 2008
</bibRefCitation>
; 
<bibRefCitation id="EFDD4B1DFF87FF80FF2A59102CAD7033" author="Fresneda, J. &amp; Salgado, J. M." box="[151,446,1539,1565]" pageId="32" pageNumber="183" refId="ref18026" refString="Fresneda, J. &amp; Salgado, J. M. (2017) Catalogo de los Coleopteros Leiodidae Cholevinae Kirby, 1837 de la Peninsula Iberica e Islas Baleares. Monografies del Museu de Ciencies Naturals, 7 [2016]. Consorci del Museu de Ciencies Naturals de Barcelona, 308 pp." type="book" year="2017">Fresneda &amp; Salgado 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BF336ECFF87FF80FF7A59342C08706F" blockId="32.[151,1437,1251,1601]" box="[199,283,1575,1601]" pageId="32" pageNumber="183">
<emphasis id="B938EAFEFF87FF80FF7A59342C00706F" bold="true" box="[199,275,1575,1601]" pageId="32" pageNumber="183">Lleida</emphasis>
:
</paragraph>
<paragraph id="8BF336ECFF87FF80FF2A597C285970A7" blockId="32.[151,1436,1647,1817]" box="[151,1354,1647,1673]" pageId="32" pageNumber="183">
<emphasis id="B938EAFEFF87FF80FF2A597C2DBE70A7" bold="true" box="[151,173,1647,1673]" pageId="32" pageNumber="183">1.</emphasis>
<materialsCitation id="3B243CB1FF87FF80FF7B597C285570A7" ID-GBIF-Occurrence="3033943322" box="[198,1350,1647,1673]" collectingDate="1971-03-07" collectionCode="MZB" collectorName="Belles &amp; Comas" elevation="940" pageId="32" pageNumber="183" specimenCount="1">
Peramola, Avenc Viladrich, 
<quantity id="4CB49B09FF87FF80FDBD597C2F5A70A7" box="[512,585,1647,1673]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="32" pageNumber="183" unit="m" value="940.0">
<elevation id="0061D1DFFF87FF80FDBD597C2F5A70A7" box="[512,585,1647,1673]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="32" pageNumber="183" unit="m" value="940.0">940 m</elevation>
</quantity>
, 
<date id="FFF2102CFF87FF80FDE9597C2FAE70A7" box="[596,701,1647,1673]" pageId="32" pageNumber="183" value="1971-03-07">
<collectingDate id="EFB6E9C4FF87FF80FDE9597C2FAE70A7" box="[596,701,1647,1673]" pageId="32" pageNumber="183" value="1971-03-07">7-3-1971</collectingDate>
</date>
, 
<collectorName id="26B9533AFF87FF80FD74597C2E1C70A7" box="[713,783,1647,1673]" pageId="32" pageNumber="183">Bellés</collectorName>
&amp; 
<collectorName id="26B9533AFF87FF80FC8E597C2E9270A7" box="[819,897,1647,1673]" pageId="32" pageNumber="183">Comas</collectorName>
leg. (
<collectionCode id="ED5DAE29FF87FF80FC7D597C291370A7" box="[960,1024,1647,1673]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="32" pageNumber="183" type="Museum">MZB</collectionCode>
) (
<bibRefCitation id="EFDD4B1DFF87FF80FBAB597C282C70A7" author="Fresneda, J. &amp; Salgado, J. M." box="[1046,1343,1647,1673]" pageId="32" pageNumber="183" refId="ref18026" refString="Fresneda, J. &amp; Salgado, J. M. (2017) Catalogo de los Coleopteros Leiodidae Cholevinae Kirby, 1837 de la Peninsula Iberica e Islas Baleares. Monografies del Museu de Ciencies Naturals, 7 [2016]. Consorci del Museu de Ciencies Naturals de Barcelona, 308 pp." type="book" year="2017">Fresneda &amp; Salgado 2017</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336ECFF87FF80FF2A59802C1C70FF" blockId="32.[151,1436,1647,1817]" pageId="32" pageNumber="183">
<emphasis id="B938EAFEFF87FF80FF2A59802DBE7083" bold="true" box="[151,173,1683,1709]" pageId="32" pageNumber="183">2.</emphasis>
Peramola, Pou del Dring (
<bibRefCitation id="EFDD4B1DFF87FF80FE4059802F867083" author="Escola, O." box="[509,661,1683,1709]" pageId="32" pageNumber="183" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà, 1971</bibRefCitation>
) (Avenc de Pau Esteve, 
<bibRefCitation id="EFDD4B1DFF87FF80FC7D598029417083" author="Escola, O." box="[960,1106,1683,1709]" pageId="32" pageNumber="183" pagination="91 - 93" refId="ref17452" refString="Escola, O. (1973) Addenda a &quot; un nuevo Speonomus de la provincia de Lerida &quot;. Speleon, 20, 91 - 93." type="journal article" year="1973">Escolà 1973</bibRefCitation>
); 
<date id="FFF2102CFF87FF80FBD6598029F17083" box="[1131,1250,1683,1709]" pageId="32" pageNumber="183" value="1970-01-18">18-1-1970</date>
, 2 exx. (
<bibRefCitation id="EFDD4B1DFF87FF80FAED59802C1170FF" author="Escola, O." pageId="32" pageNumber="183" pagination="141 - 147" refId="ref17480" refString="Escola, O. (1980) Sur la coexistence des Bathysciinae dans les grottes d'Espagne. Memories de Biospeologie, 7, 141 - 147." type="journal article" year="1980">Escolà 1980</bibRefCitation>
).
</paragraph>
<paragraph id="8BF336ECFF87FF80FF2A59C82E0070DB" blockId="32.[151,1436,1647,1817]" box="[151,787,1755,1781]" pageId="32" pageNumber="183">
<emphasis id="B938EAFEFF87FF80FF2A59C82DBF70DB" bold="true" box="[151,172,1755,1781]" pageId="32" pageNumber="183">3.</emphasis>
Peramola, Avenc de la Roca del Corb (
<bibRefCitation id="EFDD4B1DFF87FF80FDCA59C82E1570DB" author="Escola, O." box="[631,774,1755,1781]" pageId="32" pageNumber="183" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà 1971</bibRefCitation>
).
</paragraph>
<paragraph id="8BF336ECFF87FF80FF2A59EC2F857137" blockId="32.[151,1436,1647,1817]" box="[151,662,1791,1817]" pageId="32" pageNumber="183">
<emphasis id="B938EAFEFF87FF80FF2A59EC2DBE7137" bold="true" box="[151,173,1791,1817]" pageId="32" pageNumber="183">4.</emphasis>
Peramola, Avenc del Mort (
<bibRefCitation id="EFDD4B1DFF87FF80FE4759EC2F9A7137" author="Escola, O." box="[506,649,1791,1817]" pageId="32" pageNumber="183" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>