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<document id="40D5A1124D2F2E1F7350E81995C5CF56" ID-DOI="10.11646/phytotaxa.233.3.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13633146" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725320595591" checkinUser="felipe" docAuthor="Majewska, Roksana, Kociolek, J. P., Thomas, Evan W., Stefano, Mario De, Santoro, Mario, Bolaños, Federico &amp; Vijver, Bart Van De" docDate="2015" docId="03E587D85D38FFC34A8FFF0BFF22C215" docLanguage="en" docName="phytotaxa.233.3.2.pdf" docOrigin="Phytotaxa 233 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.233.3.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Chelonicola costaricensis Majewska, De Stefano &amp; Van de Vijver 2015, sp. nov." docType="treatment" docVersion="2" lastPageNumber="243" masterDocId="FFDCFFA05D3EFFC44A07FF95FFD7C050" masterDocTitle="Chelonicola and Poulinea, two new gomphonemoid diatom genera (Bacillariophyta) living on marine turtles from Costa Rica" masterLastPageNumber="250" masterPageNumber="236" pageNumber="242" updateTime="1725320997363" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D77FDF5801C978D91E9B1FB7588D6F79">Chelonicola and Poulinea, two new gomphonemoid diatom genera (Bacillariophyta) living on marine turtles from Costa Rica</mods:title>
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<mods:namePart id="BBF4CF1A86017E563282E592790B9F0F">Majewska, Roksana</mods:namePart>
<mods:affiliation id="DDC292A26706D10551D6F2B661139FA6">Department of Environmental, Biological and Pharmaceutical Sciences and Technologies, II University of Naples, via Vivaldi 43, 81100 Caserta, Italy &amp; BioNEM Laboratory, Department of Experimental and Clinical Medicine, University “ Magna Graecia ” of Catanzaro, Loc. Germaneto, 88100 Catanzaro, Italy</mods:affiliation>
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<mods:namePart id="C7855D13966E3298A9CB6E37C78CEA5F">Kociolek, J. P.</mods:namePart>
<mods:affiliation id="6F3DF6AE0C35652C211DE0BA497A16D5">Museum of Natural History and &amp; Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, CO 80309, USA</mods:affiliation>
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<mods:namePart id="C23E864030606207A48FF4B515112954">Thomas, Evan W.</mods:namePart>
<mods:affiliation id="02CF422C4A765E5ADE08B27CF415A381">Museum of Natural History and &amp; Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, CO 80309, USA</mods:affiliation>
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<mods:namePart id="0CD6C5DB4D26C42B8BB317C12C1C2088">Stefano, Mario De</mods:namePart>
<mods:affiliation id="F1381F175A1A5933833098A6B396BF62">Department of Environmental, Biological and Pharmaceutical Sciences and Technologies, II University of Naples, via Vivaldi 43, 81100 Caserta, Italy</mods:affiliation>
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<mods:namePart id="3BF751F6033F0F209426588899469D14">Santoro, Mario</mods:namePart>
<mods:affiliation id="06A4906F8544FFBDDD47D718BCCAE137">Department of Public Health and Infectious Diseases, Section of Parasitology, Sapienza University of Rome, Rome, Italy &amp; Istituto Zooprofilattico Sperimentale del Mezzogiorno, Portici, Italy</mods:affiliation>
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<mods:namePart id="EFE8564C8AA77FA1AC852A745441EC9B">Bolaños, Federico</mods:namePart>
<mods:affiliation id="4E05A3C2FD6497FDA5941198D64DDC3B">Escuela de Biología, Universidad de Costa Rica, San José, Costa Rica</mods:affiliation>
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<mods:namePart id="78CB2A0FE80C3D5053699E06E35558F9">Vijver, Bart Van De</mods:namePart>
<mods:affiliation id="E25131A60E52BDC9C1915C11C17647BC">Botanic Garden Meise, Department of Bryophyta &amp; Thallophyta, Nieuwelaan 38, B- 1860 Meise, Belgium &amp; University of Antwerp, Department of Biology, ECOBE, Universiteitsplein 1, B- 2610 Wilrijk, Belgium</mods:affiliation>
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<mods:title id="2883B4CB969FAA9FEF2EC9DCE19F0AEA">Phytotaxa</mods:title>
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<treatment id="03E587D85D38FFC34A8FFF0BFF22C215" ID-DOI="http://doi.org/10.5281/zenodo.13633156" ID-Zenodo-Dep="13633156" LSID="urn:lsid:plazi:treatment:03E587D85D38FFC34A8FFF0BFF22C215" httpUri="http://treatment.plazi.org/id/03E587D85D38FFC34A8FFF0BFF22C215" lastPageId="7" lastPageNumber="243" pageId="6" pageNumber="242">
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<heading id="D0BB81A25D38FFC24A8FFF0BFBA0C0E9" bold="true" box="[136,1143,158,185]" fontSize="11" level="1" pageId="6" pageNumber="242" reason="1">
<emphasis id="B938EADC5D38FFC24A8FFF0BFBA0C0E9" bold="true" box="[136,1143,158,185]" pageId="6" pageNumber="242">
<taxonomicName id="4C4C4D4D5D38FFC24A8FFF0BFC5EC0E9" authority="Majewska, De Stefano &amp; Van de Vijver" authorityName="Majewska, De Stefano &amp; Van de Vijver" authorityYear="2015" box="[136,905,158,185]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Chelonicola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="6" pageNumber="242" phylum="Ochrophyta" rank="species" species="costaricensis" status="sp. nov.">
<emphasis id="B938EADC5D38FFC24A8FFF0BFE78C0E8" bold="true" box="[136,431,158,185]" italics="true" pageId="6" pageNumber="242">Chelonicola costaricensis</emphasis>
Majewska, De Stefano &amp; Van de Vijver
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,
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<emphasis id="B938EADC5D38FFC24992FF0AFC79C0E9" bold="true" box="[917,942,159,185]" italics="true" pageId="6" pageNumber="242">sp</emphasis>
.
<emphasis id="B938EADC5D38FFC249BBFF0AFC31C0E9" bold="true" box="[956,998,159,185]" italics="true" pageId="6" pageNumber="242">nov</emphasis>
.
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(
<figureCitation id="13772A4B5D38FFC249FCFF35FBB8C0E9" box="[1019,1135,160,185]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStartId-0="5.[136,243,1851,1872]" captionStartId-1="6.[136,243,1627,1648]" captionTargetBox-0="[164,1423,635,1826]" captionTargetBox-1="[151,1436,491,1602]" captionTargetId-0="figure-195@5.[164,1424,635,1826]" captionTargetId-1="figure-150@6.[151,1436,491,1602]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." captionText-1="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13633148" figureDoi-1="http://doi.org/10.5281/zenodo.13633152" httpUri-0="https://zenodo.org/record/13633148/files/figure.png" httpUri-1="https://zenodo.org/record/13633152/files/figure.png" pageId="6" pageNumber="242">Figs 1–12</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF336CE5D38FFC24A8FFF71FC68C1D9" blockId="6.[136,1452,228,465]" pageId="6" pageNumber="242">
Frustules wedge-shaped in girdle view showing conspicuous septa at both poles. Valves small, heteropolar, typically clavate with a broadly rounded, non-protracted headpole and an acutely terminating footpole. Septa visible in LM and SEM on both poles. Valve dimensions (n=50): length 6.0–17.5 μm, width 1.7–3.1 μm. Axial area very narrow, not discernible in LM. Central area very small. Raphe filiform, straight with simple, indistinct proximal raphe endings. Distal raphe endings not discernible in LM. Striae almost parallel throughout the entire valve, very faintly visible in LM,
<quantity id="4CB49B2B5D38FFC24918FEE1FCA3C1D9" box="[799,884,372,393]" metricMagnitude="0" metricUnit="m" metricValue="1.0541" metricValueMax="1.1938" metricValueMin="0.9144" pageId="6" pageNumber="242" unit="in" value="41.5" valueMax="47.0" valueMin="36.0">36–47 in</quantity>
10 μm.
</paragraph>
</subSubSection>
<subSubSection id="C35665455D38FFC34A8FFE0DFC24C1AD" lastPageId="7" lastPageNumber="243" pageId="6" pageNumber="242" type="materials_examined">
<paragraph id="8BF336CE5D38FFC24A8FFE0DFF2BC181" blockId="6.[136,1452,228,465]" pageId="6" pageNumber="242">
<emphasis id="B938EADC5D38FFC24A8FFE0DFF6DC1FD" bold="true" box="[136,186,408,429]" pageId="6" pageNumber="242">Type</emphasis>
:—
<collectingCountry id="F35B765E5D38FFC24AD1FE0DFEBEC1FD" box="[214,361,408,429]" name="Costa Rica" pageId="6" pageNumber="242">COSTA RICA</collectingCountry>
. Olive ridley sea turtle,
<geoCoordinate id="EE7850095D38FFC2485EFE0DFD3CC1FD" box="[601,747,408,429]" degrees="9" direction="north" minutes="59" orientation="latitude" pageId="6" pageNumber="242" precision="1" seconds="23.7" value="9.989917">9º 59’ 23.7” N</geoCoordinate>
,
<geoCoordinate id="EE7850095D38FFC248F0FE0DFC4FC1FD" box="[759,920,408,429]" degrees="85" direction="west" minutes="41" orientation="longitude" pageId="6" pageNumber="242" precision="1" seconds="52.6" value="-85.697945">85º 41’ 52.6” W</geoCoordinate>
,
<emphasis id="B938EADC5D38FFC249A3FE0DFC6FC1FD" box="[932,952,408,429]" italics="true" pageId="6" pageNumber="242">M</emphasis>
.
<emphasis id="B938EADC5D38FFC249C1FE0DFBE5C1FD" box="[966,1074,408,429]" italics="true" pageId="6" pageNumber="242">De Stefano</emphasis>
,
<emphasis id="B938EADC5D38FFC24E47FE0DFB30C1FD" box="[1088,1255,408,429]" italics="true" pageId="6" pageNumber="242">
<date id="FFF2100E5D38FFC24E47FE0DFB30C1FD" box="[1088,1255,408,429]" pageId="6" pageNumber="242" value="2013-10-27">27 October 2013</date>
</emphasis>
(
<typeStatus id="54F7886C5D38FFC24EF1FE0DFA9BC1FD" box="[1270,1356,408,429]" pageId="6" pageNumber="242" type="holotype">holotype</typeStatus>
<collectionCode id="ED5DAE0B5D38FFC24F54FE0DFAAFC1FD" box="[1363,1400,408,429]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="6" pageNumber="242">BR</collectionCode>
! stub 4420).
</paragraph>
<caption id="DF3366465D38FFC24A8FF9CEFD02C750" ID-DOI="http://doi.org/10.5281/zenodo.13633152" ID-Zenodo-Dep="13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="6" pageNumber="242" startId="6.[136,243,1627,1648]" targetBox="[151,1436,491,1602]" targetPageId="6" targetType="figure">
<paragraph id="8BF336CE5D38FFC24A8FF9CEFD02C750" blockId="6.[136,1452,1627,1792]" pageId="6" pageNumber="242">
<emphasis id="B938EADC5D38FFC24A8FF9CEFEFEC620" bold="true" box="[136,297,1627,1648]" pageId="6" pageNumber="242">FIGURES 7–12</emphasis>
.
<taxonomicName id="4C4C4D4D5D38FFC24B30F9CEFDE7C620" authorityName="Majewska, De Stefano &amp; Van de Vijver" authorityYear="2015" box="[311,560,1627,1648]" class="Bacillariophyceae" family="Rhoicospheniaceae" genus="Chelonicola" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="6" pageNumber="242" phylum="Ochrophyta" rank="species" species="costaricensis">
<emphasis id="B938EADC5D38FFC24B30F9CEFDE7C620" box="[311,560,1627,1648]" italics="true" pageId="6" pageNumber="242">Chelonicola costaricensis</emphasis>
</taxonomicName>
. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm.
</paragraph>
</caption>
<paragraph id="8BF336CE5D38FFC34A8FF8A5FC24C1AD" blockId="6.[136,1452,1840,2081]" lastBlockId="7.[136,1452,160,581]" lastPageId="7" lastPageNumber="243" pageId="6" pageNumber="242">
<emphasis id="B938EADC5D38FFC24A8FF8A5FE22C719" bold="true" box="[136,501,1840,1865]" pageId="6" pageNumber="242">Scanning Electron Microscopy</emphasis>
:—(
<figureCitation id="13772A4B5D38FFC24827F8A5FD45C719" box="[544,658,1840,1865]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStartId-0="5.[136,243,1851,1872]" captionStartId-1="6.[136,243,1627,1648]" captionTargetBox-0="[164,1423,635,1826]" captionTargetBox-1="[151,1436,491,1602]" captionTargetId-0="figure-195@5.[164,1424,635,1826]" captionTargetId-1="figure-150@6.[151,1436,491,1602]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." captionText-1="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13633148" figureDoi-1="http://doi.org/10.5281/zenodo.13633152" httpUri-0="https://zenodo.org/record/13633148/files/figure.png" httpUri-1="https://zenodo.org/record/13633152/files/figure.png" pageId="6" pageNumber="242">Figs 1–12</figureCitation>
) Frustules isovalvar, clavate in girdle view, attached by the footpole on short mucilaginous stalks (
<figureCitation id="13772A4B5D38FFC24BBFF8C1FE29C73D" box="[440,510,1876,1901]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="6" pageNumber="242">Fig. 3</figureCitation>
). Valve face flat in both valves with a clear angle to the very shallow mantle (
<figureCitation id="13772A4B5D38FFC24F7DF8C1FF40C7C1" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="6" pageNumber="242">Figs 3</figureCitation>
,
<figureCitation id="13772A4B5D38FFC24AA0F8EDFF60C7C1" box="[167,183,1912,1937]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="6" pageNumber="242">7</figureCitation>
). The mantle is equally high in its distal and proximal part but larger in the central part (
<figureCitation id="13772A4B5D38FFC24EC5F8EDFADEC7C1" box="[1218,1289,1912,1937]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="6" pageNumber="242">Fig. 7</figureCitation>
). Pseudosepta absent (
<figureCitation id="13772A4B5D38FFC24AE7F809FE53C7E5" box="[224,388,1948,1973]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="6" pageNumber="242">Figs 9, 10, 12</figureCitation>
). Axial area very narrow, linear (
<figureCitation id="13772A4B5D38FFC248F8F809FCBFC7E5" box="[767,872,1948,1973]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="6" pageNumber="242">Figs 4–6</figureCitation>
). Central area very small, bordered on one or both sides by one slightly shortened central stria (
<figureCitation id="13772A4B5D38FFC24879F855FD31C789" box="[638,742,1984,2009]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStartId-0="5.[136,243,1851,1872]" captionStartId-1="6.[136,243,1627,1648]" captionTargetBox-0="[164,1423,635,1826]" captionTargetBox-1="[151,1436,491,1602]" captionTargetId-0="figure-195@5.[164,1424,635,1826]" captionTargetId-1="figure-150@6.[151,1436,491,1602]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." captionText-1="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13633148" figureDoi-1="http://doi.org/10.5281/zenodo.13633152" httpUri-0="https://zenodo.org/record/13633148/files/figure.png" httpUri-1="https://zenodo.org/record/13633152/files/figure.png" pageId="6" pageNumber="242">Figs 4–7</figureCitation>
). Fascia never present (
<figureCitation id="13772A4B5D38FFC249E8F855FBE3C789" box="[1007,1076,1984,2009]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="6" pageNumber="242">Fig. 5</figureCitation>
). External raphe branches almost straight to very weakly curving (
<figureCitation id="13772A4B5D38FFC24BF8F871FDB0C7AD" box="[511,615,2020,2045]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="6" pageNumber="242">Figs 4–6</figureCitation>
). External proximal raphe endings slightly expanded, unilaterally weakly deflected (
<figureCitation id="13772A4B5D38FFC24B07F79DFE91C871" box="[256,326,2056,2081]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="6" pageNumber="242">Fig. 5</figureCitation>
). External distal raphe fissures elongated, weakly deflected, continuing shortly onto the mantle on both poles (
<figureCitation id="13772A4B5D39FFC34B0FFF35FE98C0E9" box="[264,335,160,185]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="7" pageNumber="243">Figs 6</figureCitation>
,
<figureCitation id="13772A4B5D39FFC34B5DFF35FEBDC0E9" box="[346,362,160,185]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">7</figureCitation>
). Striae uniseriate, equally spaced throughout the entire valve, composed of a series of 3–5 slightly transapically elongated areolae (
<figureCitation id="13772A4B5D39FFC34BE8FF51FD83C08D" box="[495,596,196,221]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStartId-0="5.[136,243,1851,1872]" captionStartId-1="6.[136,243,1627,1648]" captionTargetBox-0="[164,1423,635,1826]" captionTargetBox-1="[151,1436,491,1602]" captionTargetId-0="figure-195@5.[164,1424,635,1826]" captionTargetId-1="figure-150@6.[151,1436,491,1602]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." captionText-1="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13633148" figureDoi-1="http://doi.org/10.5281/zenodo.13633152" httpUri-0="https://zenodo.org/record/13633148/files/figure.png" httpUri-1="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">Figs 4–7</figureCitation>
). Areolae bordering the axial area being the largest (
<figureCitation id="13772A4B5D39FFC34E92FF51FB0FC08D" box="[1173,1240,196,221]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="7" pageNumber="243">Fig. 6</figureCitation>
). Striae continuing without interruption onto the shallow mantle (
<figureCitation id="13772A4B5D39FFC34884FF7DFD3FC151" box="[643,744,232,257]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStartId-0="5.[136,243,1851,1872]" captionStartId-1="6.[136,243,1627,1648]" captionTargetBox-0="[164,1423,635,1826]" captionTargetBox-1="[151,1436,491,1602]" captionTargetId-0="figure-195@5.[164,1424,635,1826]" captionTargetId-1="figure-150@6.[151,1436,491,1602]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." captionText-1="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13633148" figureDoi-1="http://doi.org/10.5281/zenodo.13633152" httpUri-0="https://zenodo.org/record/13633148/files/figure.png" httpUri-1="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">Figs 4–7</figureCitation>
). Apical pore field absent on both poles (
<figureCitation id="13772A4B5D39FFC34EAEFF7DFADCC151" box="[1193,1291,232,257]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="7" pageNumber="243">Figs 4, 6</figureCitation>
,
<figureCitation id="13772A4B5D39FFC34F11FF7DFAF1C151" box="[1302,1318,232,257]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">8</figureCitation>
). Internally, raphe straight to weakly curved, positioned asymmetrically in a raised raphe sternum (
<figureCitation id="13772A4B5D39FFC34E4AFE99FB15C175" box="[1101,1218,268,293]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">Figs 9–12</figureCitation>
). Primary side of the sternum thickened, opening the raphe in a lateral position (
<figureCitation id="13772A4B5D39FFC34928FEA5FC71C119" box="[815,934,304,329]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">Figs 9–12</figureCitation>
). Proximal raphe endings covered by a silica flap and distal raphe endings straight, terminating on weakly developed helictoglossae (
<figureCitation id="13772A4B5D39FFC34E5CFEC1FB07C13D" box="[1115,1232,340,365]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">Figs 9–12</figureCitation>
). Areolae internally slightly sunken between thickened interstriae, covered by hymenes (
<figureCitation id="13772A4B5D39FFC349A4FEEDFBFCC1C1" box="[931,1067,376,401]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">Figs 11, 12</figureCitation>
). Cingulum composed of a large number (up to 12) of open copulae, each with one row of apically elongated, slit-like poroids in the advalvar position (
<figureCitation id="13772A4B5D39FFC34A97FE55FF24C189" box="[144,243,448,473]" captionStart="FIGURES 1–6" captionStartId="5.[136,243,1851,1872]" captionTargetBox="[164,1423,635,1826]" captionTargetId="figure-195@5.[164,1424,635,1826]" captionTargetPageId="5" captionText="FIGURES 1–6. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 1. Three frustules of Chelonicola attached to the substrate together with one frustule (upper one) of Poulinea. 2. Several valves of Chelonicola attached to the substrate showing their valve face and the girdle composition. 3. One frustule attached to the substrate by a mucilaginous stalk. The girdle structure composed of at least 10 bands is shown. 4. Valve face view showing the head pole with the distal raphe fissure continuing onto the mantle. 5. External detail of the central area showing the lack of fascia and the stria structure. 6. External detail of a head pole showing the distal raphe fissure. Scale bar represents 10 μm except for Figs 5, 6 where scale bar = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633148" httpUri="https://zenodo.org/record/13633148/files/figure.png" pageId="7" pageNumber="243">Figs 3, 4</figureCitation>
&amp;
<figureCitation id="13772A4B5D39FFC34B11FE55FEF1C189" box="[278,294,448,473]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">7</figureCitation>
). First band, the valvocopula, with a small, but distinct septum at the head pole (
<figureCitation id="13772A4B5D39FFC34EA9FE55FB25C189" box="[1198,1266,448,473]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">Fig. 9</figureCitation>
). Second copula with a small septum at the footpole (
<figureCitation id="13772A4B5D39FFC34826FE71FDA3C1AD" box="[545,628,484,509]" captionStart="FIGURES 7–12" captionStartId="6.[136,243,1627,1648]" captionTargetBox="[151,1436,491,1602]" captionTargetId="figure-150@6.[151,1436,491,1602]" captionTargetPageId="6" captionText="FIGURES 7–12. Chelonicola costaricensis. SEM pictures taken from the holotype population from the olive ridley sea turtle carapace collected on Ostional Beach, Costa Rica. 7. External valve face view showing the mantle and the areolae. 8. External detail of a footpole showing the absence of an apical pore field and the distal raphe fissure continuing onto the mantle. 9 &amp; 10. Internal views of an entire valve showing the raphe sternum, the septate girdle bands and the covered areolae. 11. Internal detail of the central area. 12. Internal detail of the footpole with helictoglossa. Scale bar represents 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13633152" httpUri="https://zenodo.org/record/13633152/files/figure.png" pageId="7" pageNumber="243">Fig. 10</figureCitation>
). Other copulae lacking a septum.
</paragraph>
</subSubSection>
<subSubSection id="C35665455D39FFC34ABAFD9DFF22C215" pageId="7" pageNumber="243" type="etymology">
<paragraph id="8BF336CE5D39FFC34ABAFD9DFF22C215" blockId="7.[136,1452,160,581]" pageId="7" pageNumber="243">
<emphasis id="B938EADC5D39FFC34ABAFD9DFEE9C271" bold="true" box="[189,318,520,545]" pageId="7" pageNumber="243">Etymology</emphasis>
:—The specific epithet refers to the geographical locality,
<collectingCountry id="F35B765E5D39FFC349E3FD9DFBB3C271" box="[996,1124,520,545]" name="Costa Rica" pageId="7" pageNumber="243">Costa Rica</collectingCountry>
, where the species was first observed.
</paragraph>
</subSubSection>
</treatment>
</document>
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