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<mods:title id="3D757E0D53F9F8880DB142502520B9B5">The cicadas (Hemiptera: Cicadoidea: Cicadidae) of Madagascar including a new tribe, five new genera, twelve new species, four new species synonymies, five revised species status, ten new combinations, new tribal assignments for four genera, one new subtribe synonymy, a checklist and key to the species</mods:title>
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<paragraph id="8BF33695FF8CFFDE46BAF9ABFE56F9D5" blockId="18.[151,412,1616,1642]" box="[151,412,1616,1642]" pageId="18" pageNumber="19">
<heading id="D0BB81F9FF8CFFDE46BAF9ABFE56F9D5" bold="true" box="[151,412,1616,1642]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="3">
<emphasis id="B938EA87FF8CFFDE46BAF9ABFE56F9D5" bold="true" box="[151,412,1616,1642]" pageId="18" pageNumber="19">
<taxonomicName id="4C4C4D16FF8CFFDE46BAF9ABFEF0F9D5" ID-CoL="8KNKM" authority="Sanborn, 2021" authorityName="Sanborn" authorityYear="2021" box="[151,314,1616,1642]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Antankariini">Antankariini</taxonomicName>
<taxonomicNameLabel id="A20B57FCFF8CFFDE476FF9ABFE56F9D5" box="[322,412,1616,1642]" pageId="18" pageNumber="19" rank="tribe">n. tribe</taxonomicNameLabel>
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<paragraph id="8BF33695FF8CFFDE46BAF96EFDA6F912" blockId="18.[151,895,1684,1740]" box="[151,620,1684,1709]" pageId="18" pageNumber="19">
<typeStatus id="54F78837FF8CFFDE46BAF96EFF03F912" box="[151,201,1685,1709]" pageId="18" pageNumber="19">Type</typeStatus>
genus.—
<taxonomicName id="4C4C4D16FF8CFFDE4706F96FFDA2F912" authority="Distant 1904 b: 429" authorityName="Distant" authorityPageNumber="429" authorityYear="1904" box="[299,616,1684,1709]" class="Insecta" family="Cicadidae" genus="Antankaria" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE4706F96FFE57F913" box="[299,413,1684,1708]" italics="true" pageId="18" pageNumber="19">Antankaria</emphasis>
<bibRefCitation id="EFDD4B64FF8CFFDE478EF96FFDA2F912" author="Distant, W. L." box="[419,616,1684,1709]" pageId="18" pageNumber="19" pagination="425 - 430" refId="ref68150" refString="Distant, W. L. (1904 b) Rhynchotal notes XXVIII. Annals and Magazine of Natural History, Series 7, 14, 425 - 430. https: // doi. org / 10.1080 / 03745480409443032" type="journal article" year="1904">Distant 1904b: 429</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF33695FF8CFFDE46BAF948FCB5F974" blockId="18.[151,895,1684,1740]" box="[151,895,1715,1740]" pageId="18" pageNumber="19">
Included genera.—
<taxonomicName id="4C4C4D16FF8CFFDE4774F948FDA9F974" authority="Distant 1904 b" authorityName="Distant" authorityYear="1904" box="[345,611,1715,1739]" class="Insecta" family="Cicadidae" genus="Antankaria" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE4774F948FE01F974" box="[345,459,1715,1739]" italics="true" pageId="18" pageNumber="19">Antankaria</emphasis>
<bibRefCitation id="EFDD4B64FF8CFFDE47FFF948FDA9F974" author="Distant, W. L." box="[466,611,1715,1739]" pageId="18" pageNumber="19" pagination="425 - 430" refId="ref68150" refString="Distant, W. L. (1904 b) Rhynchotal notes XXVIII. Annals and Magazine of Natural History, Series 7, 14, 425 - 430. https: // doi. org / 10.1080 / 03745480409443032" type="journal article" year="1904">Distant 1904b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4C4D16FF8CFFDE4440F948FCFBF974" authority="Sanborn, 2021" authorityName="Sanborn" authorityYear="2021" box="[621,817,1715,1739]" class="Insecta" family="Cicadidae" genus="Orientafroinsularis" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B938EA87FF8CFFDE4440F948FCFBF974" box="[621,817,1715,1739]" italics="true" pageId="18" pageNumber="19">Orientafroinsularis</emphasis>
</taxonomicName>
<emphasis id="B938EA87FF8CFFDE4515F948FCB5F974" bold="true" box="[824,895,1715,1739]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A20B57FCFF8CFFDE4515F948FCB5F974" box="[824,895,1715,1739]" pageId="18" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BF33695FF8CFFDF46BAF902FB5BFF6A" blockId="18.[151,1437,1785,2028]" lastBlockId="19.[151,1437,151,2013]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Remarks.—The position of the
<collectingCountry id="F35B7605FF8CFFDE47D5F902FDB5F8AC" box="[504,639,1785,1811]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
species of
<taxonomicName id="4C4C4D16FF8CFFDE44D1F902FC36F8AC" authority="Stal, 1870" authorityName="Stal" authorityYear="1870" box="[764,1020,1785,1812]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE44D1F902FC4EF8AC" box="[764,900,1785,1811]" italics="true" pageId="18" pageNumber="19">Chremistica</emphasis>
<bibRefCitation id="EFDD4B64FF8CFFDE45A6F902FC36F8AC" author="Stal, C." box="[907,1020,1785,1812]" pageId="18" pageNumber="19" pagination="607 - 776" refId="ref70935" refString="Stal, C. (1870) Hemiptera insularum Phillippinarum. Bidrag till Philippinska ˆ arnes Hemipter-fauna. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar, 27, 607 - 776. https: // doi. org / 10.5962 / bhl. title. 61898" type="journal article" year="1870">Stål, 1870</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C4C4D16FF8CFFDE421FF902FA84F8AC" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" box="[1074,1358,1785,1811]" class="Insecta" family="Cicadidae" genus="Antankaria" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE421FF902FB7AF8AC" box="[1074,1200,1785,1811]" italics="true" pageId="18" pageNumber="19">Antankaria</emphasis>
Distant, 1904
</taxonomicName>
within the
<taxonomicName id="4C4C4D16FF8CFFDE46EDF8E6FD8CF887" authority="Handlirsch, 1925" authorityName="Handlirsch" authorityYear="1925" box="[192,582,1821,1848]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="tribe" tribe="Cryptotympanini">
Cryptotympanini
<bibRefCitation id="EFDD4B64FF8CFFDE47ABF8E6FD8CF887" author="Handlirsch, A." box="[390,582,1821,1848]" pageId="18" pageNumber="19" pagination="1102 - 1126" refId="ref68808" refString="Handlirsch, A. (1925) Ordung: Homoptera (Latr.) Westw. (Homopterern) in Systemtische Ubersicht (Schluss.). Schroder's Handbuch der Entomologie, 3, 1102 - 1126." type="journal article" year="1925">Handlirsch, 1925</bibRefCitation>
</taxonomicName>
was questioned by the results of the five gene phylogenetic analysis of
<bibRefCitation id="EFDD4B64FF8CFFDE4348F8E5FEA7F8E3" author="Marshall, D. C. &amp; Moulds, M. S. &amp; Hill, K. B. R. &amp; Price, B. W. &amp; Wade, E. J. &amp; Owen, C. J. &amp; Goemans, G. &amp; Marathe, K. &amp; Sarkar, V. &amp; Cooley, J. R. &amp; Sanborn, A. F. &amp; Kunte, K. &amp; Villet, M. H. &amp; Simon, C." pageId="18" pageNumber="19" pagination="1 - 64" refId="ref69587" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018 b) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B938EA87FF8CFFDE46FEF8B9FECEF8E4" box="[211,260,1857,1883]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (2018b)
</bibRefCitation>
. The only
<collectingCountry id="F35B7605FF8CFFDE47CFF8BAFDA3F8E4" box="[482,617,1857,1883]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
species of
<taxonomicName id="4C4C4D16FF8CFFDE44C8F8BAFCA7F8E4" authorityName="Stal" authorityYear="1870" box="[741,877,1857,1883]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE44C8F8BAFCA7F8E4" box="[741,877,1857,1883]" italics="true" pageId="18" pageNumber="19">Chremistica</emphasis>
</taxonomicName>
included in the analysis formed part of a different clade than the Asian species of
<taxonomicName id="4C4C4D16FF8CFFDE442CF89EFD43F8C0" authorityName="Stal" authorityYear="1870" box="[513,649,1893,1919]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE442CF89EFD43F8C0" box="[513,649,1893,1919]" italics="true" pageId="18" pageNumber="19">Chremistica</emphasis>
</taxonomicName>
, separated at the first branch of the
<taxonomicName id="4C4C4D16FF8CFFDE4205F89EFB52F8C0" authorityName="Latreille" authorityYear="1802" box="[1064,1176,1893,1919]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Cicadinae">Cicadinae</taxonomicName>
clade and unrelated to
<taxonomicName id="4C4C4D16FF8CFFDE46BAF871FE74F81C" authority="Stal, 1861" authorityName="Stal" authorityYear="1861" box="[151,446,1929,1956]" class="Insecta" family="Cicadidae" genus="Cryptotympana" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE46BAF871FE8FF81B" box="[151,325,1930,1956]" italics="true" pageId="18" pageNumber="19">Cryptotympana</emphasis>
<bibRefCitation id="EFDD4B64FF8CFFDE4760F872FE74F81C" author="Stal, C." box="[333,446,1929,1956]" pageId="18" pageNumber="19" pagination="613 - 622" refId="ref70882" refString="Stal, C. (1861) Genera nonnulla nova Cicadinorum. Annales de la Societe Entomologique de France, Series 4, 1, 613 - 622." type="journal article" year="1861">Stål, 1861</bibRefCitation>
</taxonomicName>
and the other
<taxonomicName id="4C4C4D16FF8CFFDE4472F872FCD4F81C" authorityName="Handlirsch" authorityYear="1925" box="[607,798,1929,1955]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="tribe" tribe="Cryptotympanini">Cryptotympanini</taxonomicName>
included in the study (
<bibRefCitation id="EFDD4B64FF8CFFDE4232F872FADCF81B" author="Marshall, D. C. &amp; Moulds, M. S. &amp; Hill, K. B. R. &amp; Price, B. W. &amp; Wade, E. J. &amp; Owen, C. J. &amp; Goemans, G. &amp; Marathe, K. &amp; Sarkar, V. &amp; Cooley, J. R. &amp; Sanborn, A. F. &amp; Kunte, K. &amp; Villet, M. H. &amp; Simon, C." box="[1055,1302,1929,1956]" pageId="18" pageNumber="19" pagination="1 - 64" refId="ref69587" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018 b) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B938EA87FF8CFFDE42A1F871FB77F81C" box="[1164,1213,1929,1955]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. 2018b
</bibRefCitation>
). The major clade with the
<collectingCountry id="F35B7605FF8CFFDE4716F856FE08F878" box="[315,450,1965,1991]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
species of
<taxonomicName id="4C4C4D16FF8CFFDE446DF856FD02F878" authorityName="Stal" authorityYear="1870" box="[576,712,1965,1991]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE446DF856FD02F878" box="[576,712,1965,1991]" italics="true" pageId="18" pageNumber="19">Chremistica</emphasis>
</taxonomicName>
included the
<taxonomicName id="4C4C4D16FF8CFFDE4572F856FBC8F878" authorityName="Marshall &amp; Moulds" authorityYear="2018" box="[863,1026,1965,1991]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="tribe" tribe="Macrotristriini">Macrotristriini</taxonomicName>
in a separate divergent clade but the
<collectingCountry id="F35B7605FF8CFFDE46BAF82AFED4F854" box="[151,286,2001,2027]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
species of
<taxonomicName id="4C4C4D16FF8CFFDE478FF829FDF3F854" authorityName="Stal" authorityYear="1870" box="[418,569,2002,2027]" class="Insecta" family="Cicadidae" genus="Macrotristria" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE478FF829FDF3F854" box="[418,569,2002,2027]" italics="true" pageId="18" pageNumber="19">Macrotristria</emphasis>
</taxonomicName>
was not included in their analysis. The
<collectingCountry id="F35B7605FF8CFFDE4224F82AFB5AF854" box="[1033,1168,2001,2027]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
species of
<taxonomicName id="4C4C4D16FF8CFFDE4339F82AFA56F854" authorityName="Stal" authorityYear="1870" box="[1300,1436,2001,2027]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8CFFDE4339F82AFA56F854" box="[1300,1436,2001,2027]" italics="true" pageId="18" pageNumber="19">Chremistica</emphasis>
</taxonomicName>
were left in Crytoptympanini because
<bibRefCitation id="EFDD4B64FF8DFFDF4413FF6CFCF7FF0D" author="Marshall, D. C. &amp; Moulds, M. S. &amp; Hill, K. B. R. &amp; Price, B. W. &amp; Wade, E. J. &amp; Owen, C. J. &amp; Goemans, G. &amp; Marathe, K. &amp; Sarkar, V. &amp; Cooley, J. R. &amp; Sanborn, A. F. &amp; Kunte, K. &amp; Villet, M. H. &amp; Simon, C." box="[574,829,151,178]" pageId="19" pageNumber="20" pagination="1 - 64" refId="ref69587" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018 b) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B938EA87FF8DFFDF4485FF6CFD12FF0E" box="[680,728,151,177]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. (2018b)
</bibRefCitation>
lacked sufficient material to place the genera properly. A new tribe is provided here to incorporate these species along with species of
<taxonomicName id="4C4C4D16FF8DFFDF4221FF40FB40FF6A" authorityName="Distant" authorityYear="1904" box="[1036,1162,187,213]" class="Insecta" family="Cicadidae" genus="Antankaria" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF4221FF40FB40FF6A" box="[1036,1162,187,213]" italics="true" pageId="19" pageNumber="20">Antankaria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF33695FF8DFFDF46EAFF24FD2EFDDE" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
Diagnosis and distinguishing features.—The genitalia of the
<collectingCountry id="F35B7605FF8DFFDF45ACFF24FBC2FF46" box="[897,1032,223,249]" name="Madagascar" pageId="19" pageNumber="20">Madagascar</collectingCountry>
species of
<taxonomicName id="4C4C4D16FF8DFFDF42A0FF24FADFFF46" authorityName="Stal" authorityYear="1870" box="[1165,1301,223,249]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF42A0FF24FADFFF46" box="[1165,1301,223,249]" italics="true" pageId="19" pageNumber="20">Chremistica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D16FF8DFFDF437DFF24FF18FEA2" authorityName="Distant" authorityYear="1904" class="Insecta" family="Cicadidae" genus="Antankaria" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF437DFF24FF18FEA2" italics="true" pageId="19" pageNumber="20">Antankaria</emphasis>
</taxonomicName>
show distinct differences from the Asian species of
<taxonomicName id="4C4C4D16FF8DFFDF453AFEF8FC55FEA2" authorityName="Stal" authorityYear="1870" box="[791,927,259,285]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF453AFEF8FC55FEA2" box="[791,927,259,285]" italics="true" pageId="19" pageNumber="20">Chremistica</emphasis>
</taxonomicName>
including the pygofer distal shoulder curving to produce a dorsal point on either side of the base of the dorsal beak in species of the
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genera, the uncus of the Asian species of
<taxonomicName id="4C4C4D16FF8DFFDF47B1FEB0FDEEFEDA" authorityName="Stal" authorityYear="1870" box="[412,548,331,357]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF47B1FEB0FDEEFEDA" box="[412,548,331,357]" italics="true" pageId="19" pageNumber="20">Chremistica</emphasis>
</taxonomicName>
generally terminates in a much larger extension without a terminal bifurcation, the lateral branches of the uncus lack the broad base of the
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genera and may have finger-like extensions, the pygofer basal lobes are more lobate, separated from the pygofer margin, and often curving mediad or concealed by the mediad recurving lateral pygofer margin, the aedeagus lacks the terminal membrane, the head is more triangular, the eyes do not extend laterally beyond the anterior lateral pronotal collar, the pronotal collar is proportionately wider, the anterior terminus of the lateral pronotal collar is reflexed dorsally, the posterior cruciform elevation has a shallow smooth curved, the lateral margins of the abdomen are generally parallel, and female sternite VII has a very wide notch in the true species of
<taxonomicName id="4C4C4D16FF8DFFDF4478FDBCFD17FDDE" authorityName="Stal" authorityYear="1870" box="[597,733,583,609]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF4478FDBCFD17FDDE" box="[597,733,583,609]" italics="true" pageId="19" pageNumber="20">Chremistica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF33695FF8DFFDF46EAFD90FE8EF8F2" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
As a result of the significantly different genitalia and multiple morphological differences found in
<collectingCountry id="F35B7605FF8DFFDF4338FD90FA56FD3A" box="[1301,1436,619,645]" name="Madagascar" pageId="19" pageNumber="20">Madagascar</collectingCountry>
genera and the true species of
<taxonomicName id="4C4C4D16FF8DFFDF47C2FD74FCD8FD16" authority="Antankariini" authorityName=", Antankariini" authorityYear="2021" box="[495,786,655,681]" class="Insecta" family="Cicadidae" genus="Chremistica" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus" status="trib. nov.">
<emphasis id="B938EA87FF8DFFDF47C2FD74FDBDFD16" box="[495,631,655,681]" italics="true" pageId="19" pageNumber="20">Chremistica</emphasis>
, Antankariini
</taxonomicName>
<emphasis id="B938EA87FF8DFFDF4534FD74FCBBFD17" bold="true" box="[793,881,654,681]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A20B57FCFF8DFFDF4534FD74FCBBFD17" box="[793,881,654,681]" pageId="19" pageNumber="20" rank="tribe">n. tribe</taxonomicNameLabel>
</emphasis>
is here proposed with the
<typeStatus id="54F78837FF8DFFDF42B0FD74FB07FD16" box="[1181,1229,655,681]" pageId="19" pageNumber="20">type</typeStatus>
genus
<taxonomicName id="4C4C4D16FF8DFFDF4333FD74FE8BFD72" authority="Distant, 1904 b" authorityName="Distant" authorityYear="1904" class="Insecta" family="Cicadidae" genus="Antankaria" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF4333FD74FA56FD16" box="[1310,1436,655,681]" italics="true" pageId="19" pageNumber="20">Antankaria</emphasis>
<bibRefCitation id="EFDD4B64FF8DFFDF46BAFD48FE8BFD72" author="Distant, W. L." box="[151,321,691,717]" pageId="19" pageNumber="20" pagination="425 - 430" refId="ref68150" refString="Distant, W. L. (1904 b) Rhynchotal notes XXVIII. Annals and Magazine of Natural History, Series 7, 14, 425 - 430. https: // doi. org / 10.1080 / 03745480409443032" type="journal article" year="1904">Distant, 1904b</bibRefCitation>
</taxonomicName>
. The diagnosis of
<taxonomicName id="4C4C4D16FF8DFFDF4422FD48FD57FD72" authority="Sanborn, 2021" authorityName="Sanborn" authorityYear="2021" box="[527,669,691,717]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Antankariini">Antankariini</taxonomicName>
<emphasis id="B938EA87FF8DFFDF4488FD48FD34FD73" bold="true" box="[677,766,690,717]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A20B57FCFF8DFFDF4488FD48FD34FD73" box="[677,766,690,717]" pageId="19" pageNumber="20" rank="tribe">n. tribe</taxonomicNameLabel>
</emphasis>
is as follows: head about as wide as (
<taxonomicName id="4C4C4D16FF8DFFDF4283FD48FAE6FD72" authorityName="Distant" authorityYear="1904" box="[1198,1324,691,717]" class="Insecta" family="Cicadidae" genus="Antankaria" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF4283FD48FAE6FD72" box="[1198,1324,691,717]" italics="true" pageId="19" pageNumber="20">Antankaria</emphasis>
</taxonomicName>
) or wider than mesonotum (
<taxonomicName id="4C4C4D16FF8DFFDF4749FD2CFDF7FD4E" authority="Sanborn, 2021" authorityName="Sanborn" authorityYear="2021" box="[356,573,727,753]" class="Insecta" family="Cicadidae" genus="Orientafroinsularis" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B938EA87FF8DFFDF4749FD2CFDF7FD4E" box="[356,573,727,753]" italics="true" pageId="19" pageNumber="20">Orientafroinsularis</emphasis>
</taxonomicName>
<emphasis id="B938EA87FF8DFFDF4468FD2CFD5EFD4E" bold="true" box="[581,660,727,753]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A20B57FCFF8DFFDF4468FD2CFD5EFD4E" box="[581,660,727,753]" pageId="19" pageNumber="20" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
), supra-antennal plate reaching about half distance to eye, eyes protruding laterally from head, wider than anterior pronotal collar, lateral ocelli closer to each other than to eyes, higher than median ocellus in frontal view, lateral vertex narrower than eye, vertex at area of ocelli shorter than frons, epicranial suture deep and narrow between lateral ocelli, postclypeus extending beyond supra-antennal plates, postclypeus dorsal length not as long as dorsal width, not as long as dorsal vertex, postclypeus apex rounded in lateral view, convex in frontal view without central sulcus, pronotum shorter than mesonotum, paranota present but angled ventrally, pronotal collar lateral angle ampliate, wider than transverse pronotal collar, transverse pronotal collar not as wide as eye, mesonotum covering dorsal metanotum, metanotum extends laterally beyond wing groove, fore femora with three spines, tarsi three-segmented, hindcoxae lacking a large inner protuberance, meracanthus triangular generally curved mediad, extending over anterior male operculum or to middle of female sternite II, male operculum completely encapsulating meracanthus, with smoothly curved posterior margins, domed posterolateral to meracanthus, covering elongated tympanal cavity, extending to anterior of proportionately long sternite II, female operculum approximately rectangular, posterior margin sinuate, not reaching medial meracanthus or posterior sternite II, fore wings and hind wing hyaline, talc-like in some examples of
<taxonomicName id="4C4C4D16FF8DFFDF45EEFB50FB8BFB7A" authorityName="Distant" authorityYear="1904" box="[963,1089,1195,1221]" class="Insecta" family="Cicadidae" genus="Antankaria" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B938EA87FF8DFFDF45EEFB50FB8BFB7A" box="[963,1089,1195,1221]" italics="true" pageId="19" pageNumber="20">Antankaria</emphasis>
</taxonomicName>
, with eight and six apical cells respectively, infuscation on radial and radiomedial crossveins and distal veins between apical cells, pterostigma present, fore wing basal cell longer than broad, subrectangular, costal vein not expanded with shelf-like extension, radius anterior vein aligned closely with the subcostal vein for its length, fore wing cubitus anterior straight at base, base of median and cubitus anterior veins arising separately from basal cell, radius anterior and radius posterior arise from same location on node, cubitus anterior 1 divided by crossvein so distal portion shortest, radial and radiomedial crossveins slightly sinuate, parallel or almost parallel, ulnar cell 3 parallel to distal medial cell, abdomen longer than distance between apex of head and posterior of cruciform elevation, male abdomen fusiform, abdomen begins narrowing posteriorly to genitalia at tergite 4 or 5, second abdominal segment proportionately longer than other segments producing elongated tympanal cavity ventrally, timbal cover slightly inflated laterally, completely covering timbal, reaching metathorax anteriorly, timbal extending below wing base, tympana concealed by opercula, sternites lacking abdominal tubercles, male sternite VIII U-shaped when viewed from posterior with transverse posterior margin, female sternite VII with curved posterolateral margin and single V-shaped medial notch, pygofer distal shoulder smoothly curved with dorsally oriented point, dorsal beak broadly triangular, pygofer upper lobe absent, pygofer basal lobe well-developed extending about half pygofer length, adpressed to pygofer, pygofer lateral uncus lobes broad at base with narrow terminal extension, distally arching and curving mediad forming a canal to restrain aedeagus at apex, claspers absent, male aedeagus tubular with expanded terminal membrane and single vesicle process, restrained by medial extensions of uncus near its base, female abdominal segment 9 with dorsal beak, posterior margin sinuate.
</paragraph>
<paragraph id="8BF33695FF8DFFD846EAF8ACFF39FDA6" blockId="19.[151,1437,151,2013]" lastBlockId="20.[151,1437,151,609]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<taxonomicName id="4C4C4D16FF8DFFDF46EAF8ACFE9FF8CE" authority="Sanborn, 2021" authorityName="Sanborn" authorityYear="2021" box="[199,341,1879,1905]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Antankariini">Antankariini</taxonomicName>
<emphasis id="B938EA87FF8DFFDF4773F8ACFE73F8CF" bold="true" box="[350,441,1878,1905]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A20B57FCFF8DFFDF4773F8ACFE73F8CF" box="[350,441,1878,1905]" pageId="19" pageNumber="20" rank="tribe">n. tribe</taxonomicNameLabel>
</emphasis>
can be distinguished from the remaining
<taxonomicName id="4C4C4D16FF8DFFDF45B0F8ACFBC7F8CE" authorityName="Latreille" authorityYear="1802" box="[925,1037,1879,1905]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Cicadinae">Cicadinae</taxonomicName>
tribes by having in combination a supra-antennal plate reaching about half distance to eye, eyes protruding laterally from head, wider than anterior pronotal collar, lateral ocelli closer to each other than to eyes, higher than median ocellus in frontal view, epicranial suture deep and narrow between lateral ocelli, postclypeus dorsal length not as long as dorsal width, not as long as dorsal vertex, convex ventral postclypeus, lateral pronotum not ampliate, transverse pronotal collar not as wide as eye, hindcoxae lacking a large inner protuberance, male operculum completely encapsulating meracanthus, covering elongated tympanal cavity, costal vein not expanded with shelf-like extension, cubitus anterior 1 divided by crossvein so distal portion shortest, parallel or nealy parallel radial and radiomedial crossveins, abdomen longer than distance between apex of head and posterior of cruciform elevation, second male abdominal segment proportionately longer than other segments producing elongated tympanal cavity ventrally, timbal cover slightly inflated laterally, complete, reaching metathorax anteriorly, timbal extending below wing base, tympana concealed by opercula, sternites lacking abdominal tubercles, female sternite VII with single V-shaped medial notch, pygofer distal shoulder smoothly curved with dorsally oriented point, dorsal beak broadly triangular, pygofer lateral uncus lobes broad at base with narrow terminal extension, male aedeagus with expanded terminal membrane and single vesicle process.
</paragraph>
<paragraph id="8BF33695FF8AFFD846EAFDD8FC61FDDE" blockId="20.[151,1437,151,609]" pageId="20" pageNumber="21">
Distribution.—The included genera of the new tribe are restricted to
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and the
<collectingCountry id="F35B7605FF8AFFD842F5FDD8FF2FFDDE" name="Seychelles" pageId="20" pageNumber="21">Republic of Seychelles</collectingCountry>
(
<bibRefCitation id="EFDD4B64FF8AFFD846DEFDBCFE51FDDE" author="Metcalf, Z. P." box="[243,411,583,609]" pageId="20" pageNumber="21" pagination="1 - 585" refId="ref69829" refString="Metcalf, Z. P. (1963 a) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae. North Carolina State College Contribution, 1502, i-vii + 1 - 585." type="journal article" year="1963">Metcalf 1963a</bibRefCitation>
;
<bibRefCitation id="EFDD4B64FF8AFFD8478AFDBCFD38FDDE" author="Duffels, J. P. &amp; van der Laan, P. A." box="[423,754,582,609]" pageId="20" pageNumber="21" refId="ref68709" refString="Duffels, J. P. &amp; van der Laan, P. A. (1985) Catalogue of the Cicadoidea (Homoptera, Auchenorhyncha) 1956 - 1980. Series Entomologica 34. Dr. W. Junk Publishers, Dordrect, 414 pp." type="book" year="1985">Duffels &amp; van der Laan 1985</bibRefCitation>
;
<bibRefCitation id="EFDD4B64FF8AFFD844D3FDBCFC55FDDE" author="Sanborn, A. F." box="[766,927,583,609]" pageId="20" pageNumber="21" refId="ref70566" refString="Sanborn, A. F. (2013) Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet. Elsevier / Academic Press, San Diego, 1001 pp. https: // doi. org / 10.1016 / b 978 - 0 - 12 - 416647 - 9.00001 - 2" type="book" year="2013">Sanborn 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>