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<document id="E807E84F4B69537F69BD3136A793DCF0" ID-CLB-Dataset="60910" ID-DOI="10.5852/ejt.2022.813.1739" ID-GBIF-Dataset="9be7f121-e60d-4aed-8c95-e100f0950c9e" ID-ISSN="2118-9773" ID-Zenodo-Dep="6484118" ID-ZooBank="2113785B-CCA7-4412-AFE7-CCA1FF3AB3A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650912920744" checkinUser="felipe" docAuthor="Nolasco, Garduño &amp; Valdez-Mondragón, Alejandro" docDate="2022" docId="03FBE15DDC691A7B97F2B60288A0A293" docLanguage="en" docName="ejt-813_173-206.pdf" docOrigin="European Journal of Taxonomy 813" docSource="http://dx.doi.org/10.5852/ejt.2022.813.1739" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Physocyclus mariachi Nolasco &amp; Valdez-Mondragón 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="192" masterDocId="FFC29925DC651A68965FB0638906A30B" masterDocTitle="Four new species of the spider genus Physocyclus Simon, 1893 (Araneae: Pholcidae) from Mexico, with updated taxonomic identification keys" masterLastPageNumber="206" masterPageNumber="173" pageNumber="185" updateTime="1701997900573" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7A2D993113CCF83656631FEA0C993B55">Four new species of the spider genus Physocyclus Simon, 1893 (Araneae: Pholcidae) from Mexico, with updated taxonomic identification keys</mods:title>
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<mods:roleTerm id="C75D2B83C9220B1D2A6E9BFC681AB405">Author</mods:roleTerm>
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<mods:namePart id="14A95D7DBCDAF39A8FD8852AF9A4C1DD">Nolasco, Garduño</mods:namePart>
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<mods:affiliation id="88744CF80F4515BB5B91C96259F9F24E">Laboratory of Arachnology (LATLAX), Laboratorio Regional de Biodiversidad y Cultivo de Tejidos Vegetales (LBCTV), Instituto de Biología, Universidad Nacional Autónoma de Mexico (UNAM), sede Tlaxcala, Ex-Fábrica San Manuel, San Miguel Contla, 90640 Santa Cruz Tlaxcala, Tlaxcala, Mexico. Posgrado en Ciencias Biológicas (Doctorado), Centro Tlaxcala de Biología de la Conducta (CTBC), Universidad Autónoma de Tlaxcala (UATx), Carretera Federal Tlaxcala-Puebla, Km. 1.5, C. P. 90062, Tlaxcala, Mexico.</mods:affiliation>
<mods:nameIdentifier id="6EB8AEA1B20407CA4C93EA718F8ECA28" type="email">sam.zeppelin@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="EAFFF5D33FA87D36A1E414C2F52E2EBB">Valdez-Mondragón, Alejandro</mods:namePart>
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<mods:affiliation id="E9D7790A4FC7E18151ACB5A3BBEE3A72">Research Fellow. Laboratory of Arachnology (LATLAX), Laboratorio Regional de Biodiversidad y Cultivo de Tejidos Vegetales (LBCTV), Instituto de Biología, Universidad Nacional Autónoma de Mexico (UNAM), sede Tlaxcala, Ex-Fábrica San Manuel, San Miguel Contla, 90640 Santa Cruz Tlaxcala, Tlaxcala, Mexico.</mods:affiliation>
<mods:nameIdentifier id="211088615C28228DB9726CFC4A0CB1A0" type="email">lat_mactans@yahoo.com.mx</mods:nameIdentifier>
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<mods:title id="9F47DEC5464543D199984AA41AF98770">European Journal of Taxonomy</mods:title>
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<mods:date id="31A4991B2CBE34F34944036130E43E53">2022</mods:date>
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<treatment id="03FBE15DDC691A7B97F2B60288A0A293" ID-DOI="http://doi.org/10.5281/zenodo.6594553" ID-GBIF-Taxon="195701259" ID-Zenodo-Dep="6594553" LSID="urn:lsid:plazi:treatment:03FBE15DDC691A7B97F2B60288A0A293" httpUri="http://treatment.plazi.org/id/03FBE15DDC691A7B97F2B60288A0A293" lastPageId="19" lastPageNumber="192" pageId="12" pageNumber="185">
<subSubSection id="C34803C0DC691A6497F2B6028DD6A594" pageId="12" pageNumber="185" type="nomenclature">
<paragraph id="8BED504BDC691A6497F2B6028DD6A594" blockId="12.[356,1232,1633,1730]" pageId="12" pageNumber="185">
<heading id="D0A5E727DC691A6497F2B6028D80A570" box="[429,1158,1633,1660]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="185" reason="2">
<taxonomicName id="4C522BC8DC691A6497F2B6028BB0A577" ID-CoL="992RZ" authority="Nolasco &amp; Valdez-Mondragón, 2022" authorityName="Nolasco &amp; Valdez-Mondragón" authorityYear="2022" box="[429,694,1633,1660]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="185" phylum="Arthropoda" rank="species" species="mariachi" status="sp. nov.">
<emphasis id="B9268C59DC691A6497F2B6028BB0A577" bold="true" box="[429,694,1633,1660]" italics="true" pageId="12" pageNumber="185">Physocyclus mariachi</emphasis>
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<taxonomicNameLabel id="A2153122DC691A6494E2B6018A1EA577" box="[701,792,1634,1660]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
Nolasco &amp; Valdez-Mondragón
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urn:lsid:zoobank.org:act:
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<subSubSection id="C34803C0DC691A649487B6CB8A5DA5C9" box="[728,859,1704,1730]" pageId="12" pageNumber="185" type="description">
<paragraph id="8BED504BDC691A649487B6CB8A5DA5C9" blockId="12.[356,1232,1633,1730]" box="[728,859,1704,1730]" pageId="12" pageNumber="185">
<figureCitation id="13694CCEDC691A649487B6CB8A5DA5C9" box="[728,859,1704,1730]" captionStart-0="Figs 14–19" captionStart-1="Figs 20–23" captionStart-2="Figs 24–27" captionStart-3="Figs 28–33" captionStartId-0="14.[189,241,1928,1954]" captionStartId-1="16.[189,241,1963,1989]" captionStartId-2="17.[189,241,1928,1954]" captionStartId-3="18.[189,241,1892,1918]" captionTargetBox-0="[207,1395,683,1906]" captionTargetBox-1="[219,1384,641,1845]" captionTargetBox-2="[234,1356,687,1850]" captionTargetBox-3="[223,1394,613,1847]" captionTargetId-0="figure-94@14.[189,1398,682,1907]" captionTargetId-1="figure-112@16.[190,1399,612,1922]" captionTargetId-2="figure-72@17.[189,1398,641,1888]" captionTargetId-3="figure-73@18.[189,1398,610,1850]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." captionText-1="Figs 20–23. Physocyclus mariachi sp. nov. Male (holotype). 20–22. Left palp, prolateral, retrolateral and dorsal views, respectively. 23. Bulb of male palp, dorsal view. Scale bars: 0.5 mm." captionText-2="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." captionText-3="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6484125" figureDoi-1="http://doi.org/10.5281/zenodo.6484127" figureDoi-2="http://doi.org/10.5281/zenodo.6484129" figureDoi-3="http://doi.org/10.5281/zenodo.6484131" httpUri-0="https://zenodo.org/record/6484125/files/figure.png" httpUri-1="https://zenodo.org/record/6484127/files/figure.png" httpUri-2="https://zenodo.org/record/6484129/files/figure.png" httpUri-3="https://zenodo.org/record/6484131/files/figure.png" pageId="12" pageNumber="185">Figs 14–33</figureCitation>
</paragraph>
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<subSubSection id="C34803C0DC691A6596E2B68A8BEDA2D8" lastPageId="13" lastPageNumber="186" pageId="12" pageNumber="185" type="diagnosis">
<paragraph id="8BED504BDC691A6496E2B68A88CFA408" blockId="12.[189,457,1769,1795]" box="[189,457,1769,1795]" pageId="12" pageNumber="185">
<heading id="D0A5E727DC691A6496E2B68A88CFA408" bold="true" box="[189,457,1769,1795]" fontSize="11" level="3" pageId="12" pageNumber="185" reason="3">
<emphasis id="B9268C59DC691A6496E2B68A88CFA408" bold="true" box="[189,457,1769,1795]" pageId="12" pageNumber="185">Differential diagnosis</emphasis>
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<paragraph id="8BED504BDC691A6596E2B77B8BEDA2D8" blockId="12.[188,1402,1816,2018]" lastBlockId="13.[189,1400,265,467]" lastPageId="13" lastPageNumber="186" pageId="12" pageNumber="185">
Males of
<taxonomicName id="4C522BC8DC691A64976BB77B8B3BA439" authority="Nolasco &amp; Valdez-Mondragón, 2022" authorityName="Nolasco &amp; Valdez-Mondragón" authorityYear="2022" box="[308,573,1816,1842]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="185" phylum="Arthropoda" rank="species" species="mariachi" status="sp. nov.">
<emphasis id="B9268C59DC691A64976BB77B8B3BA439" box="[308,573,1816,1842]" italics="true" pageId="12" pageNumber="185">Physocyclus mariachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2153122DC691A64941AB77B8BA5A439" box="[581,675,1816,1842]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
resemble
<taxonomicName id="4C522BC8DC691A64957BB77B8AAFA439" authorityName="Valdez-Mondragon" authorityYear="2010" box="[804,937,1816,1842]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="185" phylum="Arthropoda" rank="species" species="paredesi">
<emphasis id="B9268C59DC691A64957BB77B8AAFA439" box="[804,937,1816,1842]" italics="true" pageId="12" pageNumber="185">P. paredesi</emphasis>
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in the shape of LAC, and by absence of sclerotized cones in frontal part of chelicerae (
<figureCitation id="13694CCEDC691A64957CB7588AA9A45E" box="[803,943,1851,1877]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="12" pageNumber="185">Figs 17–18</figureCitation>
,
<figureCitation id="13694CCEDC691A6495E1B7588AD9A45E" box="[958,991,1851,1877]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="12" pageNumber="185">24</figureCitation>
; VM 2010: figs: 204, 205). In
<taxonomicName id="4C522BC8DC691A64933EB75F882FA473" authority="Nolasco &amp; Valdez-Mondragón, 2022" authorityName="Nolasco &amp; Valdez-Mondragón" authorityYear="2022" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="185" phylum="Arthropoda" rank="species" species="mariachi" status="sp. nov.">
<emphasis id="B9268C59DC691A64933EB75F882FA473" italics="true" pageId="12" pageNumber="185">P. mariachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2153122DC691A64976AB73C8893A472" box="[309,405,1887,1913]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
(1) two longer and more sclerotized LAC on each male chelicera (
<figureCitation id="13694CCEDC691A6492BBB73D8C74A473" box="[1252,1394,1886,1912]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="12" pageNumber="185">Figs 17–19</figureCitation>
,
<figureCitation id="13694CCEDC691A6496E2B7E2880DA497" box="[189,267,1921,1948]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="12" pageNumber="185">24–25</figureCitation>
; VM 2010: figs 204–205;
<bibRefCitation id="EFC32DBADC691A649405B7E28A8FA490" author="Valdez-Mondragon A." box="[602,905,1921,1947]" pageId="12" pageNumber="185" pagination="202 - 220" refId="ref15200" refString="Valdez-Mondragon A. 2014. A reanalysis of the morphological phylogeny of the spider genus Physocyclus Simon, 1983 (Araneae: Pholcidae) with the description of a new species and description of the female of Physocyclus paredesi Valdez-Mondragon from Mexico. Zootaxa 3866 (2): 202 - 220. https: // doi. org / 10.11646 / zootaxa. 3866.2.2" type="journal article" year="2014">Valdez-Mondragón 2014</bibRefCitation>
: figs 21, 23–24); (2) LAC wider, more curved (
<figureCitation id="13694CCEDC691A649741B7C788AEA4B4" box="[286,424,1956,1983]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="12" pageNumber="185">Figs 17–18</figureCitation>
,
<figureCitation id="13694CCEDC691A6497ECB7C688D3A4B4" box="[435,469,1957,1983]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="12" pageNumber="185">24</figureCitation>
); (3) distal-frontal apophyses of chelicerae longer, with some sclerotized ones (
<figureCitation id="13694CCEDC691A64969AB7AB8854A4E9" box="[197,338,1992,2018]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="12" pageNumber="185">Figs 17–18</figureCitation>
,
<figureCitation id="13694CCEDC691A64973EB7AB8885A4E9" box="[353,387,1992,2018]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="12" pageNumber="185">24</figureCitation>
); (4) in retrolateral view, PR of male palp darker, straight (
<figureCitation id="13694CCEDC691A649200B7AB8DB8A4E9" box="[1119,1214,1992,2018]" captionStart="Figs 20–23" captionStartId="16.[189,241,1963,1989]" captionTargetBox="[219,1384,641,1845]" captionTargetId="figure-112@16.[190,1399,612,1922]" captionTargetPageId="16" captionText="Figs 20–23. Physocyclus mariachi sp. nov. Male (holotype). 20–22. Left palp, prolateral, retrolateral and dorsal views, respectively. 23. Bulb of male palp, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484127" httpUri="https://zenodo.org/record/6484127/files/figure.png" pageId="12" pageNumber="185">Figs 21</figureCitation>
,
<figureCitation id="13694CCEDC691A649292B7A48DE9A4E9" box="[1229,1263,1991,2018]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="12" pageNumber="185">26</figureCitation>
); (5) distal spine of PR harpoon-shaped; (6) in dorsal view, ES wide, sigmoid-shaped (
<figureCitation id="13694CCEDC681A659202B16A8DECA228" box="[1117,1258,265,291]" captionStart="Figs 20–23" captionStartId="16.[189,241,1963,1989]" captionTargetBox="[219,1384,641,1845]" captionTargetId="figure-112@16.[190,1399,612,1922]" captionTargetPageId="16" captionText="Figs 20–23. Physocyclus mariachi sp. nov. Male (holotype). 20–22. Left palp, prolateral, retrolateral and dorsal views, respectively. 23. Bulb of male palp, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484127" httpUri="https://zenodo.org/record/6484127/files/figure.png" pageId="13" pageNumber="186">Figs 22–23</figureCitation>
,
<figureCitation id="13694CCEDC681A6592A7B16A8C1CA228" box="[1272,1306,265,291]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="13" pageNumber="186">27</figureCitation>
); (7) in retrolateral view, E thinner but longer, with distal concavity between two small sclerotized spines (
<figureCitation id="13694CCEDC681A65969AB12C8855A262" box="[197,339,335,361]" captionStart="Figs 20–23" captionStartId="16.[189,241,1963,1989]" captionTargetBox="[219,1384,641,1845]" captionTargetId="figure-112@16.[190,1399,612,1922]" captionTargetPageId="16" captionText="Figs 20–23. Physocyclus mariachi sp. nov. Male (holotype). 20–22. Left palp, prolateral, retrolateral and dorsal views, respectively. 23. Bulb of male palp, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484127" httpUri="https://zenodo.org/record/6484127/files/figure.png" pageId="13" pageNumber="186">Figs 21–23</figureCitation>
,
<figureCitation id="13694CCEDC681A65973CB12C88B5A261" box="[355,435,335,362]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="13" pageNumber="186">26–27</figureCitation>
). Females of
<taxonomicName id="4C522BC8DC681A659401B1338BEDA262" authority="Nolasco &amp; Valdez-Mondragón, 2022" authorityName="Nolasco &amp; Valdez-Mondragón" authorityYear="2022" box="[606,747,335,361]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="186" phylum="Arthropoda" rank="species" species="mariachi" status="sp. nov.">
<emphasis id="B9268C59DC681A659401B1338BEDA262" box="[606,747,335,361]" italics="true" pageId="13" pageNumber="186">P. mariachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2153122DC681A6594A8B1338A51A261" box="[759,855,336,362]" pageId="13" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
with (1) triangular epigynum (
<figureCitation id="13694CCEDC681A659283B12C8C3FA262" box="[1244,1337,335,361]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="13" pageNumber="186">Fig. 31</figureCitation>
); (2) with small and conical apophyses on anterior part, located on two dark regions (
<figureCitation id="13694CCEDC681A6592D0B1118C1AA286" box="[1167,1308,370,397]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="13" pageNumber="186">Figs 31–32</figureCitation>
); (3) in dorsal view, PP long, thin, in convergent position to each other (
<figureCitation id="13694CCEDC681A659598B1F58D27A2BB" box="[967,1057,406,432]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="13" pageNumber="186">Fig. 33</figureCitation>
); PP oval, almost circular in
<taxonomicName id="4C522BC8DC681A6596E1B1D98844A2D8" authorityName="Valdez-Mondragon" authorityYear="2010" box="[190,322,441,467]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="186" phylum="Arthropoda" rank="species" species="paredesi">
<emphasis id="B9268C59DC681A6596E1B1D98844A2D8" box="[190,322,441,467]" italics="true" pageId="13" pageNumber="186">P. paredesi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC32DBADC681A65970CB1DA8B7BA2D8" author="Valdez-Mondragon A." box="[339,637,441,467]" pageId="13" pageNumber="186" pagination="202 - 220" refId="ref15200" refString="Valdez-Mondragon A. 2014. A reanalysis of the morphological phylogeny of the spider genus Physocyclus Simon, 1983 (Araneae: Pholcidae) with the description of a new species and description of the female of Physocyclus paredesi Valdez-Mondragon from Mexico. Zootaxa 3866 (2): 202 - 220. https: // doi. org / 10.11646 / zootaxa. 3866.2.2" type="journal article" year="2014">Valdez-Mondragón 2014</bibRefCitation>
: fig. 33).
</paragraph>
</subSubSection>
<caption id="DF2D00C3DC681A6596E2B7EB8C71A4E3" ID-DOI="http://doi.org/10.5281/zenodo.6484123" ID-Zenodo-Dep="6484123" httpUri="https://zenodo.org/record/6484123/files/figure.png" pageId="13" pageNumber="186" startId="13.[189,240,1928,1954]" targetBox="[191,1396,592,1906]" targetPageId="13">
<paragraph id="8BED504BDC681A6596E2B7EB8C71A4E3" blockId="13.[188,1399,1928,2025]" pageId="13" pageNumber="186">
<emphasis id="B9268C59DC681A6596E2B7EB8832A4A9" bold="true" box="[189,308,1928,1954]" pageId="13" pageNumber="186">Figs 8–13</emphasis>
. Typical habitats of the spider genus
<taxonomicName id="4C522BC8DC681A6594A7B7EB8D23A4A9" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[760,1061,1928,1954]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B9268C59DC681A6594A7B7EB8A82A4A9" box="[760,900,1928,1954]" italics="true" pageId="13" pageNumber="186">Physocyclus</emphasis>
Simon, 1893
</taxonomicName>
from Mexico.
<emphasis id="B9268C59DC681A659283B7EB8DECA4A9" bold="true" box="[1244,1258,1928,1954]" pageId="13" pageNumber="186">8</emphasis>
. Deciduous forests (Nayarit).
<emphasis id="B9268C59DC681A6597DEB7C888BCA4CE" bold="true" box="[385,442,1963,1990]" pageId="13" pageNumber="186">9–10</emphasis>
. Oasis in desert (Baja California Sur).
<emphasis id="B9268C59DC681A659536B7C88A82A4CE" bold="true" box="[873,900,1963,1989]" pageId="13" pageNumber="186">11</emphasis>
. Cave entrance located in a deciduous forest (Guerrero).
<emphasis id="B9268C59DC681A65971DB7AD8858A4E3" bold="true" box="[322,350,1998,2024]" pageId="13" pageNumber="186">12</emphasis>
. Thorny scrub with columnar cacti (Guanajuato).
<emphasis id="B9268C59DC681A6595D7B7AD8AA2A4E3" bold="true" box="[904,932,1998,2024]" pageId="13" pageNumber="186">13</emphasis>
. Xerophilous scrub (Baja California Sur).
</paragraph>
</caption>
<subSubSection id="C34803C0DC6B1A6696E2B16A8833A29C" pageId="14" pageNumber="187" type="etymology">
<paragraph id="8BED504BDC6B1A6696E2B16A8842A228" blockId="14.[189,324,265,291]" box="[189,324,265,291]" pageId="14" pageNumber="187">
<heading id="D0A5E727DC6B1A6696E2B16A8842A228" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="14" pageNumber="187" reason="3">
<emphasis id="B9268C59DC6B1A6696E2B16A8842A228" bold="true" box="[189,324,265,291]" pageId="14" pageNumber="187">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BED504BDC6B1A6696E2B1548833A29C" blockId="14.[189,1399,311,407]" pageId="14" pageNumber="187">
The species name is a noun in apposition, and it refers to a genre of regional and popular Mexican music commonly known as “Mariachi”, from the state of
<collectingRegion id="49969EA9DC6B1A669538B1398AB1A27F" box="[871,951,346,372]" country="Mexico" name="Jalisco" pageId="14" pageNumber="187">Jalisco</collectingRegion>
where the
<typeStatus id="54E9EEE9DC6B1A669264B1388D68A27E" box="[1083,1134,347,373]" pageId="14" pageNumber="187">type</typeStatus>
locality of the species is located.
</paragraph>
</subSubSection>
<subSubSection id="C34803C0DC6B1A6796E2B1DC883FA13C" lastPageId="15" lastPageNumber="188" pageId="14" pageNumber="187" type="materials_examined">
<paragraph id="8BED504BDC6B1A6696E2B1DC88ACA2D2" blockId="14.[189,426,447,473]" box="[189,426,447,473]" pageId="14" pageNumber="187">
<heading id="D0A5E727DC6B1A6696E2B1DC88ACA2D2" bold="true" box="[189,426,447,473]" fontSize="11" level="3" pageId="14" pageNumber="187" reason="3">
<emphasis id="B9268C59DC6B1A6696E2B1DC88ACA2D2" bold="true" box="[189,426,447,473]" pageId="14" pageNumber="187">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BED504BDC6B1A6696B5B18D8B06A17A" blockId="14.[189,1399,494,625]" pageId="14" pageNumber="187">
<materialsCitation id="3B3A5A16DC6B1A6696B5B18D8B06A17A" ID-GBIF-Occurrence="3797051306" collectingDate="2020-11-09" collectionCode="CNAN-T" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="1355" latitude="21.0879" location="Municipality of Hostotipaquillo" longLatPrecision="7" longitude="-104.0685" pageId="14" pageNumber="187" specimenCode="CNAN-T01471" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="B9268C59DC6B1A6696B5B18D885AA103" bold="true" box="[234,348,494,520]" pageId="14" pageNumber="187">
<typeStatus id="54E9EEE9DC6B1A6696B5B18D885AA103" box="[234,348,494,520]" pageId="14" pageNumber="187" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34510DBDC6B1A6696E2B2728837A120" box="[189,305,529,555]" name="Mexico" pageId="14" pageNumber="187">MEXICO</collectingCountry>
•
<specimenCount id="9D549BC2DC6B1A66970BB272886BA120" box="[340,365,529,555]" pageId="14" pageNumber="187" type="male">♂</specimenCount>
;
<collectingRegion id="49969EA9DC6B1A669720B27288D4A120" box="[383,466,529,555]" country="Mexico" name="Jalisco" pageId="14" pageNumber="187">Jalisco</collectingRegion>
,
<location id="8E8D0690DC6B1A6697BDB2728A63A120" LSID="urn:lsid:plazi:treatment:03FBE15DDC691A7B97F2B60288A0A293:8E8D0690DC6B1A6697BDB2728A63A120" box="[482,869,529,555]" country="Mexico" latitude="21.0879" longLatPrecision="7" longitude="-104.0685" name="Municipality of Hostotipaquillo" pageId="14" pageNumber="187" stateProvince="Jalisco">Municipality of Hostotipaquillo</location>
,
<locationDeviation id="9A5F4969DC6B1A669529B2728DDCA120" box="[886,1242,529,555]" location="Hostotipaquillo" pageId="14" pageNumber="187">
<quantity id="4CAAFDAEDC6B1A669529B2728AB0A120" box="[886,950,529,555]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="187" unit="km" value="2.0">2 km</quantity>
SW of Hostotipaquillo
</locationDeviation>
;
<geoCoordinate id="EE66368CDC6B1A6692B4B2728C74A120" box="[1259,1394,529,555]" degrees="21.0879" direction="north" orientation="latitude" pageId="14" pageNumber="187" precision="5" value="21.0879">21.0879° N</geoCoordinate>
,
<geoCoordinate id="EE66368CDC6B1A6696E2B257885EA145" box="[189,344,564,591]" degrees="104.0685" direction="west" orientation="longitude" pageId="14" pageNumber="187" precision="5" value="-104.0685">104.0685°W</geoCoordinate>
;
<elevation id="007FB778DC6B1A66973BB25788FBA145" box="[356,509,564,591]" metricMagnitude="3" metricUnit="m" metricValue="1.355" pageId="14" pageNumber="187" unit="m" value="1355.0">
<quantity id="4CAAFDAEDC6B1A66973BB25788BBA145" box="[356,445,564,591]" metricMagnitude="3" metricUnit="m" metricValue="1.355" pageId="14" pageNumber="187" unit="m" value="1355.0">1355 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEC768BDC6B1A669455B2578B9BA145" box="[522,669,564,590]" pageId="14" pageNumber="187" value="2020-11-09">
<collectingDate id="EFA88F63DC6B1A669455B2578B9BA145" box="[522,669,564,590]" pageId="14" pageNumber="187" value="2020-11-09">9 Nov. 2020</collectingDate>
</date>
;
<collectorName id="26A7359DDC6B1A6694F7B2578A18A145" box="[680,798,564,590]" pageId="14" pageNumber="187">A. Valdez</collectorName>
,
<collectorName id="26A7359DDC6B1A669575B2578AA3A145" box="[810,933,564,590]" pageId="14" pageNumber="187">I. Navarro</collectorName>
,
<collectorName id="26A7359DDC6B1A6695F0B2578D18A145" box="[943,1054,564,590]" pageId="14" pageNumber="187">A. Juárez</collectorName>
and
<collectorName id="26A7359DDC6B1A669208B2578DD0A145" box="[1111,1238,564,590]" pageId="14" pageNumber="187">S. Nolasco</collectorName>
leg.; daytime collection;
<specimenCode id="DBF4F830DC6B1A66971FB23488FAA17A" box="[320,508,599,625]" collectionCode="CNAN-T" pageId="14" pageNumber="187">CNAN-T01471</specimenCode>
.
</materialsCitation>
</paragraph>
<caption id="DF2D00C3DC6B1A6696E2B7EB8A60A4E2" ID-DOI="http://doi.org/10.5281/zenodo.6484125" ID-Zenodo-Dep="6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="14" pageNumber="187" startId="14.[189,241,1928,1954]" targetBox="[207,1395,683,1906]" targetPageId="14">
<paragraph id="8BED504BDC6B1A6696E2B7EB8A60A4E2" blockId="14.[189,1399,1928,2025]" pageId="14" pageNumber="187">
<emphasis id="B9268C59DC6B1A6696E2B7EB8841A4A9" bold="true" box="[189,327,1928,1954]" pageId="14" pageNumber="187">Figs 14–19</emphasis>
.
<taxonomicName id="4C522BC8DC6B1A669704B7EB8B65A4A9" authority="Nolasco &amp; Valdez-Mondragón, 2022" authorityName="Nolasco &amp; Valdez-Mondragón" authorityYear="2022" box="[347,611,1928,1954]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="187" phylum="Arthropoda" rank="species" species="mariachi" status="sp. nov.">
<emphasis id="B9268C59DC6B1A669704B7EB8B65A4A9" box="[347,611,1928,1954]" italics="true" pageId="14" pageNumber="187">Physocyclus mariachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2153122DC6B1A669430B7EB8BC9A4A9" box="[623,719,1928,1954]" pageId="14" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
Male (holotype).
<emphasis id="B9268C59DC6B1A6695EDB7EB8AFDA4A9" bold="true" box="[946,1019,1928,1954]" pageId="14" pageNumber="187">14–16</emphasis>
. Habitus in dorsal, lateral and ventral views, respectively.
<emphasis id="B9268C59DC6B1A669457B7C88B22A4CE" bold="true" box="[520,548,1963,1989]" pageId="14" pageNumber="187">17</emphasis>
. Carapace, frontal view.
<emphasis id="B9268C59DC6B1A66950FB7C88A6AA4CE" bold="true" box="[848,876,1963,1989]" pageId="14" pageNumber="187">18</emphasis>
,
<emphasis id="B9268C59DC6B1A669523B7C88A9FA4CE" bold="true" box="[892,921,1963,1989]" pageId="14" pageNumber="187">19</emphasis>
. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BED504BDC6A1A6796B5B16A8DE5A287" blockId="15.[189,1398,265,397]" pageId="15" pageNumber="188">
<materialsCitation id="3B3A5A16DC6A1A6796B5B16A8D1CA24D" ID-GBIF-Occurrence="3797051309" collectingDate="2020-11-09" collectionCode="CNAN-T" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="1355" latitude="21.0879" location="MEXICO" longLatPrecision="7" longitude="-104.0685" pageId="15" pageNumber="188" specimenCode="CNAN-T01472" specimenCount="1" specimenCount-female="1" stateProvince="MEXICO" typeStatus="paratype">
<emphasis id="B9268C59DC6A1A6796B5B16A886FA228" bold="true" box="[234,361,265,291]" pageId="15" pageNumber="188">
<typeStatus id="54E9EEE9DC6A1A6796B5B16A886FA228" box="[234,361,265,291]" pageId="15" pageNumber="188" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingRegion id="49969EA9DC6A1A6796E2B14F8837A24D" box="[189,305,300,326]" country="Mexico" name="Mexico" pageId="15" pageNumber="188">MEXICO</collectingRegion>
•
<specimenCount id="9D549BC2DC6A1A67970EB14F8887A24D" box="[337,385,300,326]" pageId="15" pageNumber="188" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBF4F830DC6A1A67953FB14F8D1CA24D" box="[864,1050,300,326]" collectionCode="CNAN-T" pageId="15" pageNumber="188">CNAN-T01472</specimenCode>
</materialsCitation>
•
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<quantity id="4CAAFDAEDC6A1A679263B12C8D8FA262" box="[1084,1161,335,362]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="15" pageNumber="188" unit="m" value="915.0">915 m</quantity>
a.s.l.
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;
<date id="FFEC768BDC6A1A679285B12C8C74A262" box="[1242,1394,335,361]" pageId="15" pageNumber="188" value="2020-11-09">
<collectingDate id="EFA88F63DC6A1A679285B12C8C74A262" box="[1242,1394,335,361]" pageId="15" pageNumber="188" value="2020-11-09">9 Nov. 2020</collectingDate>
</date>
;
<collectorName id="26A7359DDC6A1A6796E0B1118833A287" box="[191,309,370,396]" pageId="15" pageNumber="188">A. Valdez</collectorName>
,
<collectorName id="26A7359DDC6A1A67971FB11088BBA286" box="[320,445,371,397]" pageId="15" pageNumber="188">I. Navarro</collectorName>
,
<collectorName id="26A7359DDC6A1A679797B1118B3EA287" box="[456,568,370,396]" pageId="15" pageNumber="188">A. Juárez</collectorName>
and
<collectorName id="26A7359DDC6A1A67942DB1118BF4A287" box="[626,754,370,396]" pageId="15" pageNumber="188">S. Nolasco</collectorName>
leg.; daytime collection;
<specimenCode id="DBF4F830DC6A1A67927EB1118DD8A287" box="[1057,1246,370,396]" pageId="15" pageNumber="188">CNAN-T01473</specimenCode>
.
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</paragraph>
<paragraph id="8BED504BDC6A1A6796B5B1D088AEA2C6" blockId="15.[189,1399,435,567]" box="[234,424,435,461]" pageId="15" pageNumber="188">
<emphasis id="B9268C59DC6A1A6796B5B1D088AEA2C6" bold="true" box="[234,424,435,461]" pageId="15" pageNumber="188">Other material</emphasis>
</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B1B4883FA13C" blockId="15.[189,1399,435,567]" pageId="15" pageNumber="188">
<materialsCitation id="3B3A5A16DC6A1A6796E2B1B48D8CA2FA" ID-GBIF-Occurrence="3797051305" box="[189,1162,471,497]" collectionCode="AMNH" country="Mexico" location="Arroyo Santiago" municipality="Municipality of El Nayar" pageId="15" pageNumber="188" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nayarit">
<collectingCountry id="F34510DBDC6A1A6796E2B1B48837A2FA" box="[189,305,471,497]" name="Mexico" pageId="15" pageNumber="188">MEXICO</collectingCountry>
•
<specimenCount id="9D549BC2DC6A1A679715B1B4887EA2FA" box="[330,376,471,497]" pageId="15" pageNumber="188" type="male">1 ♂</specimenCount>
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;
<collectingRegion id="49969EA9DC6A1A67979FB1B48B1AA2FA" box="[448,540,471,497]" country="Mexico" name="Nayarit" pageId="15" pageNumber="188">Nayarit</collectingRegion>
,
<collectingMunicipality id="6B89CA31DC6A1A679477B1B48A52A2FA" box="[552,852,471,497]" pageId="15" pageNumber="188">Municipality of El Nayar</collectingMunicipality>
,
<location id="8E8D0690DC6A1A679501B1B48D20A2FA" LSID="urn:lsid:plazi:treatment:03FBE15DDC691A7B97F2B60288A0A293:8E8D0690DC6A1A679501B1B48D20A2FA" box="[862,1062,471,497]" country="Mexico" municipality="Municipality of El Nayar" name="Arroyo Santiago" pageId="15" pageNumber="188" stateProvince="Nayarit">Arroyo Santiago</location>
;
<collectionCode id="ED43C88EDC6A1A67926FB1B48D8CA2FA" box="[1072,1162,471,497]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="15" pageNumber="188">AMNH</collectionCode>
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•
<materialsCitation id="3B3A5A16DC6A1A6792FCB1B48BABA11F" ID-GBIF-Occurrence="3797051312" collectingDate="2020-11-09" collectionCode="LATLAX" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="1355" latitude="21.0879" location="Municipality of Hostotipaquillo" longLatPrecision="7" longitude="-104.0685" pageId="15" pageNumber="188" specimenCount="1" specimenCount-immature="1" stateProvince="Jalisco">
<specimenCount id="9D549BC2DC6A1A6792FCB1B48C28A2FA" box="[1187,1326,471,497]" pageId="15" pageNumber="188" type="immature">1 immature</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED43C88EDC6A1A679465B1998BABA11F" box="[570,685,506,532]" pageId="15" pageNumber="188">LATLAX</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B3A5A16DC6A1A679496B1998833A13C" ID-GBIF-Occurrence="3797051317" collectingDate="2020-11-09" collectionCode="LATLAX" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="1355" latitude="21.0879" location="Municipality of Hostotipaquillo" longLatPrecision="7" longitude="-104.0685" pageId="15" pageNumber="188" specimenCount="1" specimenCount-immature="1" stateProvince="Jalisco" typeStatus="paratype">
<specimenCount id="9D549BC2DC6A1A679496B1998A53A11F" box="[713,853,506,532]" pageId="15" pageNumber="188" type="immature">1 immature</specimenCount>
; same collection data as for second paratype;
<collectionCode id="ED43C88EDC6A1A6796E1B27E8833A13C" box="[190,309,541,567]" pageId="15" pageNumber="188">LATLAX</collectionCode>
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.
</paragraph>
</subSubSection>
<subSubSection id="C34803C0DC6A1A7996E2B23D8D29A2B1" lastPageId="17" lastPageNumber="190" pageId="15" pageNumber="188" type="description">
<paragraph id="8BED504BDC6A1A6796E2B23D8849A173" blockId="15.[189,335,606,632]" box="[189,335,606,632]" pageId="15" pageNumber="188">
<heading id="D0A5E727DC6A1A6796E2B23D8849A173" bold="true" box="[189,335,606,632]" fontSize="11" level="3" pageId="15" pageNumber="188" reason="3">
<emphasis id="B9268C59DC6A1A6796E2B23D8849A173" bold="true" box="[189,335,606,632]" pageId="15" pageNumber="188">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BED504BDC6A1A6796B5B2EE8B70A1AC" blockId="15.[188,1399,653,821]" box="[234,630,653,679]" pageId="15" pageNumber="188">
<emphasis id="B9268C59DC6A1A6796B5B2EE882CA1AC" bold="true" box="[234,298,653,679]" pageId="15" pageNumber="188">Male</emphasis>
(
<typeStatus id="54E9EEE9DC6A1A679765B2EE88A0A1AC" box="[314,422,653,679]" pageId="15" pageNumber="188" type="holotype">holotype</typeStatus>
, CNAN-T01471)
</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B2D2882FA03E" blockId="15.[188,1399,653,821]" pageId="15" pageNumber="188">MEASUREMENTS. Total length 2.5. Carapace 1.11 long, 1.3 wide. Clypeus 0.5 long. Diameter AME 0.08, ALE 0.12, PME 0.10, PLE 0.12. Distance ALE-PME 0.06, PME-PME 0.13. Leg lengths: I (total 23.5): femur 6.4/patella 0.5/tibia 6.7/metatarsus 8.5/tarsus 1.3; tibia II: 4.5; tibia III: 3.1; tibia IV: 5; tibia I L/d 40.5.</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B33F8BA2A79B" blockId="15.[188,1399,859,1168]" pageId="15" pageNumber="188">
PROSOMA. Carapace light beige with Y-shaped light gray pattern around fovea and posterior part of carapace. Fovea longitudinal. Carapace with three irregular spots on each side (
<figureCitation id="13694CCEDC6A1A67922AB31D8C07A092" box="[1141,1281,894,921]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="15" pageNumber="188">Figs 14–15</figureCitation>
). Clypeus wide, with dark brown irregular mark lengthwise (
<figureCitation id="13694CCEDC6A1A679552B3C18A62A0B7" box="[781,868,930,956]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="15" pageNumber="188">Fig. 17</figureCitation>
). Chelicerae pale, with two lateral apophyses on each side, with triangular tip and pointing down (
<figureCitation id="13694CCEDC6A1A679503B3A68AECA0D4" box="[860,1002,965,991]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="15" pageNumber="188">Figs 17–19</figureCitation>
,
<figureCitation id="13694CCEDC6A1A6795A3B3A68D4DA0EB" box="[1020,1099,965,992]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="15" pageNumber="188">24–25</figureCitation>
). Anterior apophyses of chelicerae wide, dark, sclerotized, with small cones, in lateral view looks like a sawn surface. Posterior apophyses of chelicera small and thinner than anterior apophyses, with 4–5 sclerotized cones on each side (
<figureCitation id="13694CCEDC6A1A679763B44D88C3A742" box="[316,453,1070,1097]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="15" pageNumber="188">Figs 17–19</figureCitation>
,
<figureCitation id="13694CCEDC6A1A67978DB44D8B24A742" box="[466,546,1070,1097]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="15" pageNumber="188">24–25</figureCitation>
). Chelicerae with stridulatory files. Sternum light beige, with small and inconspicuous light brown spots. Labium brown, wider than long. Endites brown prolaterally, beige retrolaterally, longer than wide (
<figureCitation id="13694CCEDC6A1A679461B4168B90A79B" box="[574,662,1141,1168]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="15" pageNumber="188">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B4D488E8A7FF" blockId="15.[189,1398,1206,1268]" pageId="15" pageNumber="188">
LEGS. All segments with beige coloration. Tibiae and femora with brown rings distally. Trochanters light brown (
<figureCitation id="13694CCEDC6A1A679709B4BA88E7A7FF" box="[342,481,1241,1268]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="15" pageNumber="188">Figs 14–16</figureCitation>
).
</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B5788A8BA652" blockId="15.[189,1398,1307,1369]" pageId="15" pageNumber="188">
OPISTHOSOMA. Globular, longer than wide, high, with numerous white, gray and dark spots (
<figureCitation id="13694CCEDC6A1A679353B57889DBA652" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="15" pageNumber="188">Figs 14– 15</figureCitation>
). Spinnerets dark brown, with short seta around (
<figureCitation id="13694CCEDC6A1A679578B55D8A79A652" box="[807,895,1342,1369]" captionStart="Figs 14–19" captionStartId="14.[189,241,1928,1954]" captionTargetBox="[207,1395,683,1906]" captionTargetId="figure-94@14.[189,1398,682,1907]" captionTargetPageId="14" captionText="Figs 14–19. Physocyclus mariachi sp. nov. Male (holotype). 14–16. Habitus in dorsal, lateral and ventral views, respectively. 17. Carapace, frontal view. 18, 19. Chelicerae in frontal and lateral views, respectively. Scale bars: 14–16 = 1 mm; 17–19 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484125" httpUri="https://zenodo.org/record/6484125/files/figure.png" pageId="15" pageNumber="188">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B5E38AB2A50F" blockId="15.[187,1398,1408,1540]" pageId="15" pageNumber="188">
PALP. DAP wide and dark (
<figureCitation id="13694CCEDC6A1A6797ACB5E38B4FA691" box="[499,585,1408,1434]" captionStart="Figs 20–23" captionStartId="16.[189,241,1963,1989]" captionTargetBox="[219,1384,641,1845]" captionTargetId="figure-112@16.[190,1399,612,1922]" captionTargetPageId="16" captionText="Figs 20–23. Physocyclus mariachi sp. nov. Male (holotype). 20–22. Left palp, prolateral, retrolateral and dorsal views, respectively. 23. Bulb of male palp, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484127" httpUri="https://zenodo.org/record/6484127/files/figure.png" pageId="15" pageNumber="188">Fig. 21</figureCitation>
). PR entirely dark, with notch in middle part, ending in long thin spine (
<figureCitation id="13694CCEDC6A1A67969CB5C0884BA6B6" box="[195,333,1443,1469]" captionStart="Figs 20–23" captionStartId="16.[189,241,1963,1989]" captionTargetBox="[219,1384,641,1845]" captionTargetId="figure-112@16.[190,1399,612,1922]" captionTargetPageId="16" captionText="Figs 20–23. Physocyclus mariachi sp. nov. Male (holotype). 20–22. Left palp, prolateral, retrolateral and dorsal views, respectively. 23. Bulb of male palp, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484127" httpUri="https://zenodo.org/record/6484127/files/figure.png" pageId="15" pageNumber="188">Figs 21–23</figureCitation>
,
<figureCitation id="13694CCEDC6A1A679707B5C0887CA6B5" box="[344,378,1443,1470]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="15" pageNumber="188">26</figureCitation>
). ES small and short, with oval shape in prolateral and retrolateral views (
<figureCitation id="13694CCEDC6A1A6792B8B5C08C76A6B6" box="[1255,1392,1443,1469]" captionStart="Figs 20–23" captionStartId="16.[189,241,1963,1989]" captionTargetBox="[219,1384,641,1845]" captionTargetId="figure-112@16.[190,1399,612,1922]" captionTargetPageId="16" captionText="Figs 20–23. Physocyclus mariachi sp. nov. Male (holotype). 20–22. Left palp, prolateral, retrolateral and dorsal views, respectively. 23. Bulb of male palp, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484127" httpUri="https://zenodo.org/record/6484127/files/figure.png" pageId="15" pageNumber="188">Figs 20–21</figureCitation>
,
<figureCitation id="13694CCEDC6A1A6796E2B5A589D8A6EA" box="[189,222,1478,1505]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="15" pageNumber="188">26</figureCitation>
), wide, long and sinuous in dorsal view (
<figureCitation id="13694CCEDC6A1A679487B5A58A65A6EB" box="[728,867,1478,1504]" captionStart="Figs 20–23" captionStartId="16.[189,241,1963,1989]" captionTargetBox="[219,1384,641,1845]" captionTargetId="figure-112@16.[190,1399,612,1922]" captionTargetPageId="16" captionText="Figs 20–23. Physocyclus mariachi sp. nov. Male (holotype). 20–22. Left palp, prolateral, retrolateral and dorsal views, respectively. 23. Bulb of male palp, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484127" httpUri="https://zenodo.org/record/6484127/files/figure.png" pageId="15" pageNumber="188">Figs 22–23</figureCitation>
,
<figureCitation id="13694CCEDC6A1A67952DB5A58A92A6EB" box="[882,916,1478,1504]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="15" pageNumber="188">27</figureCitation>
). E small and short in retrolateral view, ending in a small tip. SO positioned in distal part of E (
<figureCitation id="13694CCEDC6A1A679511B58A8AA1A50F" box="[846,935,1513,1540]" captionStart="Figs 24–27" captionStartId="17.[189,241,1928,1954]" captionTargetBox="[234,1356,687,1850]" captionTargetId="figure-72@17.[189,1398,641,1888]" captionTargetPageId="17" captionText="Figs 24–27. Physocyclus mariachi sp. nov. Male (holotype). 24–25. Detail of chelicerae, frontal and lateral views, respectively. 26. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view. 27. Detail of bulb, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="15" pageNumber="188">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="8BED504BDC6A1A6796B5B6488B97A54E" blockId="15.[189,657,1579,1640]" box="[234,657,1579,1605]" pageId="15" pageNumber="188">
<emphasis id="B9268C59DC6A1A6796B5B6488840A54E" bold="true" box="[234,326,1579,1605]" pageId="15" pageNumber="188">Female</emphasis>
(
<typeStatus id="54E9EEE9DC6A1A679709B64888C7A54E" box="[342,449,1579,1605]" pageId="15" pageNumber="188" type="paratype">paratype</typeStatus>
, CNAN-T01472)
</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B62D8B3DA563" blockId="15.[189,657,1579,1640]" box="[189,571,1614,1640]" pageId="15" pageNumber="188">Similar to the male, differences:</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B6F38BC3A418" blockId="15.[189,1399,1679,1811]" pageId="15" pageNumber="188">
PROSOMA. Carapace clearer, almost white, with darker dorsal pattern (
<figureCitation id="13694CCEDC6A1A679253B6EC8D61A5A2" box="[1036,1127,1679,1705]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="15" pageNumber="188">Fig. 28</figureCitation>
). Carapace with small protuberance on posterior part (
<figureCitation id="13694CCEDC6A1A679467B6D18B96A5C6" box="[568,656,1714,1741]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="15" pageNumber="188">Fig. 29</figureCitation>
, red arrow). Sternum brown, with beige region in middle part (
<figureCitation id="13694CCEDC6A1A679698B6B58827A5FB" box="[199,289,1750,1776]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="15" pageNumber="188">Fig. 30</figureCitation>
). Clypeus darker than in male, beige region in central part. Chelicerae with brown coloration, without LAC or sclerotized cones (
<figureCitation id="13694CCEDC6A1A679401B69A8BBEA418" box="[606,696,1785,1811]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="15" pageNumber="188">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="8BED504BDC6A1A6796E2B7598BD3A473" blockId="15.[187,1398,1850,1912]" pageId="15" pageNumber="188">
OPISTHOSOMA. With dorsal patch in anterior part (
<figureCitation id="13694CCEDC6A1A6794AEB7598A41A45F" box="[753,839,1850,1876]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="15" pageNumber="188">Fig. 28</figureCitation>
, red arrow). This structure might be a functional complex (with the protuberance of prosoma).
</paragraph>
<paragraph id="8BED504BDC6A1A7896E2B7C38A1DA262" blockId="15.[189,949,1951,1978]" lastBlockId="16.[189,1399,265,361]" lastPageId="16" lastPageNumber="189" pageId="15" pageNumber="188">
LEGS. Femora and tibiae with darker rings distally (
<figureCitation id="13694CCEDC6A1A679542B7FC8AAEA4B2" box="[797,936,1951,1977]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="15" pageNumber="188">Figs 28–30</figureCitation>
). EPIGYNUM. Wider than long, bell-shaped, with VAE small, conical, pointing forward, surrounded by dark spot, with clear stripe across epigynum longitudinally (
<figureCitation id="13694CCEDC751A789521B14F8D01A24D" box="[894,1031,300,326]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="16" pageNumber="189">Figs 31–32</figureCitation>
). PP small and elongated, ovalshaped, located in perpendicular position (
<figureCitation id="13694CCEDC751A7894EAB12C8A0BA262" box="[693,781,335,361]" captionStart="Figs 28–33" captionStartId="18.[189,241,1892,1918]" captionTargetBox="[223,1394,613,1847]" captionTargetId="figure-73@18.[189,1398,610,1850]" captionTargetPageId="18" captionText="Figs 28–33. Physocyclus mariachi sp. nov. Female (paratype). 28–30. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29). 31–33. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="16" pageNumber="189">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="8BED504BDC751A7896E2B1F28834A11F" blockId="16.[188,1399,401,532]" pageId="16" pageNumber="189">MEASUREMENTS. Total length 3.7. Carapace 1.2 long, 1.2 wide. Clypeus 0.4 long. Diameter AME 0.08, ALE 0.12, PME 0.11, PLE 0.11. Distance ALE-PME 0.06, PME-PME 0.12. Leg lengths: I (total 16.8): femur 4.4/patella 0.5/tibia 4.6/metatarsus 6.1/tarsus 1.1; tibia II: 3.3; tibia III: 2.4; tibia IV: 3.6; tibia I L/d: 24.6.</paragraph>
<caption id="DF2D00C3DC751A7896E2B7C88DB0A4E2" ID-DOI="http://doi.org/10.5281/zenodo.6484127" ID-Zenodo-Dep="6484127" httpUri="https://zenodo.org/record/6484127/files/figure.png" pageId="16" pageNumber="189" startId="16.[189,241,1963,1989]" targetBox="[219,1384,641,1845]" targetPageId="16">
<paragraph id="8BED504BDC751A7896E2B7C88DB0A4E2" blockId="16.[189,1399,1963,2025]" pageId="16" pageNumber="189">
<emphasis id="B9268C59DC751A7896E2B7C88842A4CE" bold="true" box="[189,324,1963,1989]" pageId="16" pageNumber="189">Figs 20–23</emphasis>
.
<taxonomicName id="4C522BC8DC751A78970AB7C88B5CA4CE" authority="Nolasco &amp; Valdez-Mondragón, 2022" authorityName="Nolasco &amp; Valdez-Mondragón" authorityYear="2022" box="[341,602,1963,1989]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="189" phylum="Arthropoda" rank="species" species="mariachi" status="sp. nov.">
<emphasis id="B9268C59DC751A78970AB7C88B5CA4CE" box="[341,602,1963,1989]" italics="true" pageId="16" pageNumber="189">Physocyclus mariachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2153122DC751A78943BB7CF8BC7A4CD" box="[612,705,1964,1990]" pageId="16" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
Male (holotype).
<emphasis id="B9268C59DC751A7895C3B7C88AE3A4CE" bold="true" box="[924,997,1963,1990]" pageId="16" pageNumber="189">20–22</emphasis>
. Left palp, prolateral, retrolateral and dorsal views, respectively.
<emphasis id="B9268C59DC751A789474B7AD8B4EA4E3" bold="true" box="[555,584,1998,2024]" pageId="16" pageNumber="189">23</emphasis>
. Bulb of male palp, dorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BED504BDC741A7996E2B16B8835A229" blockId="17.[189,307,264,290]" box="[189,307,264,290]" pageId="17" pageNumber="190">
<heading id="D0A5E727DC741A7996E2B16B8835A229" bold="true" box="[189,307,264,290]" fontSize="11" level="3" pageId="17" pageNumber="190" reason="3">
<emphasis id="B9268C59DC741A7996E2B16B8835A229" bold="true" box="[189,307,264,290]" pageId="17" pageNumber="190">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BED504BDC741A7996B5B1548857A25A" blockId="17.[189,1401,311,443]" box="[234,337,311,337]" pageId="17" pageNumber="190">
<emphasis id="B9268C59DC741A7996B5B1548857A25A" bold="true" box="[234,337,311,337]" pageId="17" pageNumber="190">Females</emphasis>
</paragraph>
<paragraph id="8BED504BDC741A7996E2B1398D29A2B1" blockId="17.[189,1401,311,443]" pageId="17" pageNumber="190">
Female collected in Plan de Barrancas is bigger than female collected at
<typeStatus id="54E9EEE9DC741A799274B1388D58A27E" box="[1067,1118,347,373]" pageId="17" pageNumber="190">type</typeStatus>
locality. With an ocher coloration. The coloration of the dorsal pattern of the carapace and chelicerae is darker in this female. (
<emphasis id="B9268C59DC741A799698B1C289DDA2B1" box="[199,219,417,442]" italics="true" pageId="17" pageNumber="190">N</emphasis>
=1): Tibia I: 5.7; tibia II: 3.9; tibia III: 2.9; tibia IV: 4.4; tibia I L/d: 26.2.
</paragraph>
</subSubSection>
<subSubSection id="C34803C0DC741A7996E2B18188EDA120" box="[189,491,482,555]" pageId="17" pageNumber="190" type="distribution">
<paragraph id="8BED504BDC741A7996E2B1818850A2F7" blockId="17.[189,342,482,508]" box="[189,342,482,508]" pageId="17" pageNumber="190">
<heading id="D0A5E727DC741A7996E2B1818850A2F7" bold="true" box="[189,342,482,508]" fontSize="11" level="3" pageId="17" pageNumber="190" reason="3">
<emphasis id="B9268C59DC741A7996E2B1818850A2F7" bold="true" box="[189,342,482,508]" pageId="17" pageNumber="190">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BED504BDC741A7996E2B27288EDA120" blockId="17.[189,491,529,555]" box="[189,491,529,555]" pageId="17" pageNumber="190">
<collectingCountry id="F34510DBDC741A7996E2B272881DA120" box="[189,283,529,555]" name="Mexico" pageId="17" pageNumber="190">Mexico</collectingCountry>
:
<collectingRegion id="49969EA9DC741A799779B2728870A120" box="[294,374,529,555]" country="Mexico" name="Jalisco" pageId="17" pageNumber="190">Jalisco</collectingRegion>
(
<figureCitation id="13694CCEDC741A7997D9B27288D8A120" box="[390,478,529,555]" captionStart="Fig" captionStartId="30.[189,230,1893,1919]" captionTargetBox="[189,1398,1000,1855]" captionTargetId="figure-195@30.[189,1398,1000,1855]" captionTargetPageId="30" captionText="Fig. 78. Known records of Physocyclus mariachi sp. nov., P. sikuapu sp. nov., P. lyncis sp. nov., and P. pocamadre sp. nov. from Western Mexico. Abbreviations: BC = Baja California; BCS = Baja California Sur; COL = Colima; DGO = Durango; GRO = Guerrero; JAL = Jalisco; MICH = Michoacan; NAY = Nayarit; SIN = Sinaloa; SON = Sonora." figureDoi="http://doi.org/10.5281/zenodo.6484149" httpUri="https://zenodo.org/record/6484149/files/figure.png" pageId="17" pageNumber="190">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2D00C3DC741A7996E2B7EB8AE0A4E2" ID-DOI="http://doi.org/10.5281/zenodo.6484129" ID-Zenodo-Dep="6484129" httpUri="https://zenodo.org/record/6484129/files/figure.png" pageId="17" pageNumber="190" startId="17.[189,241,1928,1954]" targetBox="[234,1356,687,1850]" targetPageId="17">
<paragraph id="8BED504BDC741A7996E2B7EB8AE0A4E2" blockId="17.[189,1398,1928,2025]" pageId="17" pageNumber="190">
<emphasis id="B9268C59DC741A7996E2B7EB884FA4A9" bold="true" box="[189,329,1928,1954]" pageId="17" pageNumber="190">Figs 24–27</emphasis>
.
<taxonomicName id="4C522BC8DC741A799701B7EB8B6EA4A9" authority="Nolasco &amp; Valdez-Mondragón, 2022" authorityName="Nolasco &amp; Valdez-Mondragón" authorityYear="2022" box="[350,616,1928,1954]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="190" phylum="Arthropoda" rank="species" species="mariachi" status="sp. nov.">
<emphasis id="B9268C59DC741A799701B7EB8B6EA4A9" box="[350,616,1928,1954]" italics="true" pageId="17" pageNumber="190">Physocyclus mariachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2153122DC741A799429B7EA8BDEA4A8" box="[630,728,1929,1955]" pageId="17" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
Male (holotype).
<emphasis id="B9268C59DC741A79959FB7EB8D0FA4A9" bold="true" box="[960,1033,1928,1955]" pageId="17" pageNumber="190">24–25</emphasis>
. Detail of chelicerae, frontal and lateral views, respectively.
<emphasis id="B9268C59DC741A799464B7C88B5EA4CE" bold="true" box="[571,600,1963,1989]" pageId="17" pageNumber="190">26</emphasis>
. Details of procursus, embolus and embolic sclerites of left palp, retrolateral view.
<emphasis id="B9268C59DC741A7997D4B7AD88AEA4E3" bold="true" box="[395,424,1998,2024]" pageId="17" pageNumber="190">27</emphasis>
. Detail of bulb, dorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C34803C0DC771A7A96E2B16A8D57A2B0" pageId="18" pageNumber="191" type="discussion">
<paragraph id="8BED504BDC771A7A96E2B16A8829A228" blockId="18.[189,303,265,291]" box="[189,303,265,291]" pageId="18" pageNumber="191">
<heading id="D0A5E727DC771A7A96E2B16A8829A228" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="18" pageNumber="191" reason="3">
<emphasis id="B9268C59DC771A7A96E2B16A8829A228" bold="true" box="[189,303,265,291]" pageId="18" pageNumber="191">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BED504BDC771A7A96E2B1548D57A2B0" blockId="18.[187,1400,311,443]" pageId="18" pageNumber="191">
This new species was previously recorded by W. J. Gertsch from Arroyo Santiago,
<collectingRegion id="49969EA9DC771A7A9227B1548DD5A25A" box="[1144,1235,311,337]" country="Mexico" name="Nayarit" pageId="18" pageNumber="191">Nayarit</collectingRegion>
,
<collectingCountry id="F34510DBDC771A7A9282B1548C3CA25A" box="[1245,1338,311,337]" name="Mexico" pageId="18" pageNumber="191">Mexico</collectingCountry>
, who tentatively named the species as
<taxonomicName id="4C522BC8DC771A7A9419B1398BD1A27F" authorityName="Simon" authorityYear="1893" box="[582,727,346,372]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B9268C59DC771A7A9419B1398BD1A27F" box="[582,727,346,372]" italics="true" pageId="18" pageNumber="191">Physocyclus</emphasis>
</taxonomicName>
“
<emphasis id="B9268C59DC771A7A94B4B1388A5CA27F" box="[747,858,347,372]" italics="true" pageId="18" pageNumber="191">nayaritus</emphasis>
”. The species was never described, however, Gertsch drew some sketches consulted by A. Valdez-Mondragón at the American Museum of Natural History (AMNH) (year 2008). Those sketches are shown herein (
<figureCitation id="13694CCEDC771A7A95E5B1C28D42A2B0" box="[954,1092,417,443]" captionStart="Fig" captionStartId="19.[189,230,1928,1954]" captionTargetBox="[209,1394,712,1884]" captionTargetId="figure-56@19.[189,1398,705,1889]" captionText="Fig. 34–37. Unpublished drawings of P. mariachi sp. nov. (= P. “nayaritus”) made by W.J. Gertsch, archived at AMNH. 34. Male palp in retrolateral view. 35–36. Left male chelicerae in frontal and retrolateral views, respectively. 37. Female epigynum, dorsal view" figureDoi="http://doi.org/10.5281/zenodo.6594494" httpUri="https://zenodo.org/record/6594494/files/figure.png" pageId="18" pageNumber="191">Figs 34–37</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2D00C3DC771A7A96E2B7078B82A4E2" ID-DOI="http://doi.org/10.5281/zenodo.6484131" ID-Zenodo-Dep="6484131" httpUri="https://zenodo.org/record/6484131/files/figure.png" pageId="18" pageNumber="191" startId="18.[189,241,1892,1918]" targetBox="[223,1394,613,1847]" targetPageId="18">
<paragraph id="8BED504BDC771A7A96E2B7078B82A4E2" blockId="18.[189,1399,1892,2025]" pageId="18" pageNumber="191">
<emphasis id="B9268C59DC771A7A96E2B7078843A474" bold="true" box="[189,325,1892,1919]" pageId="18" pageNumber="191">Figs 28–33</emphasis>
.
<taxonomicName id="4C522BC8DC771A7A9708B7068B5BA474" authority="Nolasco &amp; Valdez-Mondragón, 2022" authorityName="Nolasco &amp; Valdez-Mondragón" authorityYear="2022" box="[343,605,1893,1919]" class="Arachnida" family="Pholcidae" genus="Physocyclus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="191" phylum="Arthropoda" rank="species" species="mariachi" status="sp. nov.">
<emphasis id="B9268C59DC771A7A9708B7068B5BA474" box="[343,605,1893,1919]" italics="true" pageId="18" pageNumber="191">Physocyclus mariachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2153122DC771A7A9437B7068BC0A474" box="[616,710,1893,1919]" pageId="18" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
Female (paratype).
<emphasis id="B9268C59DC771A7A95E4B7068D02A474" bold="true" box="[955,1028,1893,1919]" pageId="18" pageNumber="191">28–30</emphasis>
. Habitus in dorsal, lateral and ventral views, respectively; red arrows indicate a dorsal patch in the opisthosoma (28) and the dorsal protuberance in carapace (29).
<emphasis id="B9268C59DC771A7A9470B7C88B71A4CE" bold="true" box="[559,631,1963,1990]" pageId="18" pageNumber="191">31–33</emphasis>
. Epigynum, ventral, lateral and dorsal views, respectively. Scale bars: 28–30 = 1 mm; 31–33 = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C34803C0DC761A7B96E2B16A88A0A293" pageId="19" pageNumber="192" type="biology_ecology">
<paragraph id="8BED504BDC761A7B96E2B16A8878A229" blockId="19.[189,382,264,291]" box="[189,382,264,291]" pageId="19" pageNumber="192">
<heading id="D0A5E727DC761A7B96E2B16A8878A229" bold="true" box="[189,382,264,291]" fontSize="11" level="3" pageId="19" pageNumber="192" reason="3">
<emphasis id="B9268C59DC761A7B96E2B16A8878A229" bold="true" box="[189,382,264,291]" pageId="19" pageNumber="192">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="8BED504BDC761A7B96E2B15488A0A293" blockId="19.[189,1399,311,408]" pageId="19" pageNumber="192">
The specimens were collected among boulders on the ground. The male
<typeStatus id="54E9EEE9DC761A7B9265B15B8D6BA259" box="[1082,1133,312,338]" pageId="19" pageNumber="192">type</typeStatus>
and
<specimenCount id="9D549BC2DC761A7B92F2B15B8C34A25A" box="[1197,1330,311,338]" pageId="19" pageNumber="192" type="female">one female</specimenCount>
were collected close together during copulation. The vegetation of the
<typeStatus id="54E9EEE9DC761A7B95E3B1388AE9A27E" box="[956,1007,347,373]" pageId="19" pageNumber="192">type</typeStatus>
locality was disturbed deciduous forest (
<figureCitation id="13694CCEDC761A7B974FB11E889FA293" box="[272,409,381,408]" captionStart="Figs 70–73" captionStartId="28.[189,241,1857,1883]" captionTargetBox="[195,1391,941,1810]" captionTargetId="figure-218@28.[189,1398,935,1814]" captionTargetPageId="28" captionText="Figs 70–73. Habitat and microhabitat of Physocyclus mariachi sp. nov. (70–71) and Physocyclus sikuapu sp. nov. (72–73). 70. Disturbed deciduous forest near Municipality of Hostotipaquillo (type locality), Jalisco, Mexico. 71. Big boulders on the ground where the specimens were collected (red arrows indicate the specific microhabitat). 72. Deciduous tropical forest, Municipality of Aquila, Costa Aquila Michoacan, Mexico. 73. Cave at the viewpoint of Costa Aquila (type locality)." figureDoi="http://doi.org/10.5281/zenodo.6484145" httpUri="https://zenodo.org/record/6484145/files/figure.png" pageId="19" pageNumber="192">Figs 70–71</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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