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<document id="A2DB217F520A75F8D82300FDFE3B9108" ID-CLB-Dataset="196617" ID-DOI="10.11646/zootaxa.5213.2.6" ID-GBIF-Dataset="ce66e40a-3c1f-400d-bd74-0a441f98e93d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7354377" ID-ZooBank="F823FA04-77C4-469B-9BC3-A942303B9DBE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669280417862" checkinUser="plazi" docAuthor="Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul &amp; Robillard, Tony" docDate="2022" docId="03FAAD53FFA2FFCDFF045E87E37095B1" docLanguage="en" docName="zootaxa.5213.2.6.pdf" docOrigin="Zootaxa 5213 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ornebius lupus Tan &amp; Japir &amp; Chung &amp; Wahab &amp; Robillard 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="187" masterDocId="FFC3D52BFFAAFFC7FF935C35E2029174" masterDocTitle="Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore" masterLastPageNumber="189" masterPageNumber="177" pageNumber="185" updateTime="1699412075568" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B65C96C24C53296A6E13C9F24FB57635">Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore</mods:title>
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<mods:namePart id="08A25247EDC2947EC4C105368AE8E397">Tan, Ming Kai</mods:namePart>
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<mods:affiliation id="9579A60029126E480C29908D0D78BCDF">Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543, Republic of Singapore orthoptera. mingkai @ gmail. com; https: // orcid. org / 0000 - 0002 - 4324 - 6305</mods:affiliation>
<mods:nameIdentifier id="B6A9192471DE9E1C163A68FD5D5EE499" type="email">orthoptera.mingkai@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A73139D335936E305F34EB3A8D43F2F4">Japir, Razy</mods:namePart>
<mods:affiliation id="728FCCE380FD03297750F4B48B375271">Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia. &amp; razy. japir @ sabah. gov. my</mods:affiliation>
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<mods:namePart id="2132DCEBC5CB5554FE4211C6AC005FE1">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="812223FAC7E190ABE87AED239AFBD5F5">Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia. &amp; arthur. chung @ sabah. gov. my, https: // orcid / org / 0000 - 0002 - 9529 - 4114</mods:affiliation>
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<mods:namePart id="690CB97DBD8622EFCBA8D19B449C85AA">Wahab, Rodzay Bin Haji Abdul</mods:namePart>
<mods:affiliation id="6737B469B92F0E3B9A9DEBB4AD6C599D">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam. rodzay. wahab @ ubd. edu. bn; https: // orcid / org / 0000 - 0002 - 2151 - 7709</mods:affiliation>
<mods:nameIdentifier id="37B88569CFBFD8307060360F3FE87292" type="email">rodzay.wahab@ubd.edu.bn</mods:nameIdentifier>
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<mods:namePart id="416ABC6DAB2F0371599BD257A756968E">Robillard, Tony</mods:namePart>
<mods:nameIdentifier id="51855943A0671C77AE162BA58DE3CDF5" type="ORCID">0000-0002-2177-9549</mods:nameIdentifier>
<mods:affiliation id="13A66480C509EA39EE92C6F4D226C59D">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France. tony. robillard @ mnhn. fr; https: // orcid. org / 0000 - 0002 - 2177 - 9549</mods:affiliation>
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<mods:title id="64F35AC57F45FBF7D4BB680A894D75A9">Zootaxa</mods:title>
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<mods:date id="F23D029D281E4612622626ECD0854004">2022</mods:date>
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<treatment id="03FAAD53FFA2FFCDFF045E87E37095B1" ID-DOI="http://doi.org/10.5281/zenodo.7371349" ID-GBIF-Taxon="205139747" ID-Zenodo-Dep="7371349" LSID="urn:lsid:plazi:treatment:03FAAD53FFA2FFCDFF045E87E37095B1" httpUri="http://treatment.plazi.org/id/03FAAD53FFA2FFCDFF045E87E37095B1" lastPageId="10" lastPageNumber="187" pageId="8" pageNumber="185">
<subSubSection id="C3494FCEFFA2FFCFFF045E87E3F793B9" box="[151,501,690,717]" pageId="8" pageNumber="185" type="nomenclature">
<paragraph id="8BEC1C45FFA2FFCFFF045E87E3F793B9" blockId="8.[151,501,690,753]" box="[151,501,690,717]" pageId="8" pageNumber="185">
<heading id="D0A4AB29FFA2FFCFFF045E87E3F793B9" bold="true" box="[151,501,690,717]" fontSize="11" level="1" pageId="8" pageNumber="185" reason="1">
<emphasis id="B927C057FFA2FFCFFF045E87E3F793B9" bold="true" box="[151,501,690,717]" pageId="8" pageNumber="185">
<taxonomicName id="4C5367C6FFA2FFCFFF045E87E38993B9" authority="Tan" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab &amp; Robillard" authorityYear="2022" box="[151,395,690,717]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="B927C057FFA2FFCFFF045E87E35093B9" bold="true" box="[151,338,690,717]" italics="true" pageId="8" pageNumber="185">Ornebius lupus</emphasis>
Tan
</taxonomicName>
,
<taxonomicNameLabel id="A2147D2CFFA2FFCFFE0B5E86E3F793B9" box="[408,501,691,717]" pageId="8" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3494FCEFFA2FFCFFF045EE2E1ED9293" pageId="8" pageNumber="185" type="description">
<paragraph id="8BEC1C45FFA2FFCFFF045EE2E3689385" blockId="8.[151,501,690,753]" box="[151,362,727,753]" pageId="8" pageNumber="185">
(
<figureCitation id="136800C0FFA2FFCFFF0C5EE2E3289385" box="[159,298,727,753]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[184,1408,198,1900]" captionTargetId="figure-21@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 1. ♂ habitus of Cycloptiloides bimaculatus Tan et al., 2021(A), Ornebius pullus Ingrisch, 2006 (B) and Ornebius lupus sp. nov. (C, D) in dorsal (AC) and dorso-lateral view (D). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7354379" httpUri="https://zenodo.org/record/7354379/files/figure.png" pageId="8" pageNumber="185">Figs 1C, 1D</figureCitation>
,
<figureCitation id="136800C0FFA2FFCFFEA45EE2E3459385" box="[311,327,727,753]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="8" pageNumber="185">6</figureCitation>
,
<figureCitation id="136800C0FFA2FFCFFEC05EE2E3609385" box="[339,354,727,753]" captionStart="FIGURE 7" captionStartId="10.[151,250,1947,1971]" captionTargetBox="[156,1426,1252,1923]" captionTargetId="figure-467@10.[151,1436,1245,1924]" captionTargetPageId="10" captionText="FIGURE 7. Ornebius lupus sp. nov. ♂ holotype genitalia in dorsal (A, B) and lateral (C) views. es = external sclerite, is = internal sclerite, lv = lateral valve, mv = medial valve. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354402" httpUri="https://zenodo.org/record/7354402/files/figure.png" pageId="8" pageNumber="185">7</figureCitation>
)
</paragraph>
<paragraph id="8BEC1C45FFA2FFCFFF045F2EE1589247" blockId="8.[151,858,795,820]" box="[151,858,795,820]" pageId="8" pageNumber="185">
<taxonomicName id="4C5367C6FFA2FFCFFF045F2EE31B9247" box="[151,281,795,819]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B927C057FFA2FFCFFF045F2EE2F79247" box="[151,245,795,819]" italics="true" pageId="8" pageNumber="185">Ornebius</emphasis>
sp.
</taxonomicName>
<emphasis id="B927C057FFA2FFCFFE8A5F29E3329247" box="[281,304,796,819]" italics="true" pageId="8" pageNumber="185"></emphasis>
Tan
<emphasis id="B927C057FFA2FFCFFEC85F29E38A9247" box="[347,392,795,819]" italics="true" pageId="8" pageNumber="185">et al</emphasis>
. under review [calling songs from
<collectingCountry id="F3445CD5FFA2FFCFFD785F2EE1569247" box="[747,852,795,819]" name="Singapore" pageId="8" pageNumber="185">Singapore</collectingCountry>
]
</paragraph>
<paragraph id="8BEC1C45FFA2FFCFFF045F54E1DE92EB" blockId="8.[151,1437,865,2008]" pageId="8" pageNumber="185">
<emphasis id="B927C057FFA2FFCFFF045F54E395920F" bold="true" box="[151,407,865,891]" pageId="8" pageNumber="185">Specimens examined.</emphasis>
<materialsCitation id="3B3B1618FFA2FFCFFE0D5F54E1DE92EB" ID-GBIF-Occurrence="3970465301" collectingDate="2022-03-19" collectorName="M. K. Tan" country="Singapore" latitude="1.37805" location="Pasir Ris" longLatPrecision="1" longitude="103.95147" pageId="8" pageNumber="185" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B927C057FFA2FFCFFE0D5F54E016920F" bold="true" box="[414,532,865,891]" pageId="8" pageNumber="185">
<typeStatus id="54E8A2E7FFA2FFCFFE0D5F54E00D920F" box="[414,527,865,891]" pageId="8" pageNumber="185" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F3445CD5FFA2FFCFFD885F54E0BA920F" box="[539,696,865,891]" name="Singapore" pageId="8" pageNumber="185">SINGAPORE</collectingCountry>
<specimenCount id="9D55D7CCFFA2FFCFFD425F54E0F4920F" box="[721,758,865,891]" pageId="8" pageNumber="185" type="male">1♂</specimenCount>
;
<location id="8E8C4A9EFFA2FFCFFC905F54E165920F" LSID="urn:lsid:plazi:treatment:03FAAD53FFA2FFCDFF045E87E37095B1:8E8C4A9EFFA2FFCFFC905F54E165920F" box="[771,871,865,891]" country="Singapore" latitude="1.37805" longLatPrecision="1" longitude="103.95147" name="Pasir Ris" pageId="8" pageNumber="185">Pasir Ris</location>
mangrove boardwalk;
<geoCoordinate id="EE677A82FFA2FFCFFBFF5F54E6DC9208" box="[1132,1246,865,892]" degrees="1.37805" direction="north" orientation="latitude" pageId="8" pageNumber="185" precision="1" value="1.37805">N1.37805</geoCoordinate>
,
<geoCoordinate id="EE677A82FFA2FFCFFB7A5F54E7779208" box="[1257,1397,865,892]" degrees="103.95147" direction="east" orientation="longitude" pageId="8" pageNumber="185" precision="1" value="103.95147">E103.95147</geoCoordinate>
;
<date id="FFED3A85FFA2FFCFFA135F54E32092EB" pageId="8" pageNumber="185" value="2022-03-19">
<collectingDate id="EFA9C36DFFA2FFCFFA135F54E32092EB" pageId="8" pageNumber="185" value="2022-03-19">19 March 2022</collectingDate>
</date>
, night time; on foliage of mangrove tree; coll.
<collectorName id="26A67993FFA2FFCFFCBF5FB3E19992D4" box="[812,923,902,928]" pageId="8" pageNumber="185">M.K. Tan</collectorName>
; ZRC
</materialsCitation>
</paragraph>
<paragraph id="8BEC1C45FFA2FFCFFF545F9CE1ED9293" blockId="8.[151,1437,865,2008]" pageId="8" pageNumber="185">
<materialsCitation id="3B3B1618FFA2FFCFFF545F9CE1ED9293" ID-GBIF-Occurrence="3970465303" collectingDate="2022-05-02" collectorName="M. K. Tan" country="Singapore" latitude="1.37805" location="Pasir Ris" longLatPrecision="1" longitude="103.95147" pageId="8" pageNumber="185" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B927C057FFA2FFCFFF545F9CE34A92B7" bold="true" box="[199,328,937,963]" pageId="8" pageNumber="185">
<typeStatus id="54E8A2E7FFA2FFCFFF545F9CE34192B7" box="[199,323,937,963]" pageId="8" pageNumber="185" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F3445CD5FFA2FFCFFEC25F9CE3EC92B7" box="[337,494,937,963]" name="Singapore" pageId="8" pageNumber="185">SINGAPORE</collectingCountry>
<specimenCount id="9D55D7CCFFA2FFCFFD985F9CE02D92B7" box="[523,559,937,963]" pageId="8" pageNumber="185" type="male">1♂</specimenCount>
;
<location id="8E8C4A9EFFA2FFCFFDAC5F9CE0A792B7" LSID="urn:lsid:plazi:treatment:03FAAD53FFA2FFCDFF045E87E37095B1:8E8C4A9EFFA2FFCFFDAC5F9CE0A792B7" box="[575,677,937,963]" country="Singapore" latitude="1.37805" longLatPrecision="1" longitude="103.95147" name="Pasir Ris" pageId="8" pageNumber="185">Pasir Ris</location>
mangrove boardwalk;
<geoCoordinate id="EE677A82FFA2FFCFFC3D5F9CE62292B0" box="[942,1056,937,964]" degrees="1.37805" direction="north" orientation="latitude" pageId="8" pageNumber="185" precision="1" value="1.37805">N1.37805</geoCoordinate>
,
<geoCoordinate id="EE677A82FFA2FFCFFBBD5F9CE6BB92B0" box="[1070,1209,937,964]" degrees="103.95147" direction="east" orientation="longitude" pageId="8" pageNumber="185" precision="1" value="103.95147">E103.95147</geoCoordinate>
;
<date id="FFED3A85FFA2FFCFFB545F9CE75492B7" box="[1223,1366,937,964]" pageId="8" pageNumber="185" value="2022-05-02">
<collectingDate id="EFA9C36DFFA2FFCFFB545F9CE75492B7" box="[1223,1366,937,964]" pageId="8" pageNumber="185" value="2022-05-02">2 May 2022</collectingDate>
</date>
, night time; on foliage of mangrove tree; coll.
<collectorName id="26A67993FFA2FFCFFDC55FFBE0C6929C" box="[598,708,974,1000]" pageId="8" pageNumber="185">M.K. Tan</collectorName>
; MNHN-EO-ENSIF11343
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3494FCEFFA2FFCFFF545FC4E77897AF" pageId="8" pageNumber="185" type="diagnosis">
<paragraph id="8BEC1C45FFA2FFCFFF545FC4E77C9597" blockId="8.[151,1437,865,2008]" pageId="8" pageNumber="185">
<emphasis id="B927C057FFA2FFCFFF545FC4E340957F" bold="true" box="[199,322,1009,1035]" pageId="8" pageNumber="185">Diagnosis.</emphasis>
The new species has genitalia which appear most similar to those in the species group
<taxonomicName id="4C5367C6FFA2FFCFFAA75FC4E79E957F" authorityName="Guérin-Méneville" authorityYear="1844" box="[1332,1436,1009,1035]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis id="B927C057FFA2FFCFFAA75FC4E79E957F" box="[1332,1436,1009,1035]" italics="true" pageId="8" pageNumber="185">Ornebius</emphasis>
</taxonomicName>
(
<taxonomicName id="4C5367C6FFA2FFCFFF325820E315955B" authorityName="Ingrisch" authorityYear="1987" box="[161,279,1045,1071]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="rufonigrus">
<emphasis id="B927C057FFA2FFCFFF325820E315955B" box="[161,279,1045,1071]" italics="true" pageId="8" pageNumber="185">rufonigrus</emphasis>
</taxonomicName>
)
<bibRefCitation id="EFC261B4FFA2FFCFFEB65820E3C5955B" author="Ingrisch, S." box="[293,455,1045,1071]" pageId="8" pageNumber="185" pagination="163 - 185" refId="ref5553" refString="Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana Biologica, 68, 163 - 185." type="journal article" year="1987">Ingrisch, 1987</bibRefCitation>
, but differs by the dark-grey colouration; the maxillary palps being not so elongated (more similar to
<taxonomicName id="4C5367C6FFA2FFCFFE98580CE1D99527" authority="Tan, Ingrisch, Baroga-Barbecho &amp; Yap, 2019" authorityName="Tan, Ingrisch, Baroga-Barbecho &amp; Yap" authorityYear="2019" box="[267,987,1081,1108]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="alvarezi">
<emphasis id="B927C057FFA2FFCFFE98580CE3D69527" box="[267,468,1081,1107]" italics="true" pageId="8" pageNumber="185">Ornebius alvarezi</emphasis>
<bibRefCitation id="EFC261B4FFA2FFCFFE48580FE1D99527" author="Tan, M. K. &amp; Ingrisch, S. &amp; Baroga-Barbecho, J. B. &amp; Yap, S. A." box="[475,987,1081,1108]" pageId="8" pageNumber="185" pagination="166 - 176" refId="ref6085" refString="Tan, M. K., Ingrisch, S., Baroga-Barbecho, J. B. &amp; Yap, S. A. (2019) New species of Ornebius (Orthoptera; Mogoplistidae; Mogoplistinae) from Siargao Island of the Philippines. Zootaxa, 4590 (1), 166 - 176. https: // doi. org / 10.11646 / zootaxa. 4590.1.7" type="journal article" year="2019">Tan, Ingrisch, Baroga-Barbecho &amp; Yap, 2019</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="F3445CD5FFA2FFCFFBDF580CE6CC9527" box="[1100,1230,1081,1107]" name="Philippines" pageId="8" pageNumber="185">Philippines</collectingCountry>
; elongated oblong in
<taxonomicName id="4C5367C6FFA2FFCFFF275868E0259503" authority="Ingrisch, 2006" authorityName="Ingrisch" authorityYear="2006" box="[180,551,1117,1143]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="dumoga">
<emphasis id="B927C057FFA2FFCFFF275868E37C9503" box="[180,382,1117,1143]" italics="true" pageId="8" pageNumber="185">Ornebius dumoga</emphasis>
<bibRefCitation id="EFC261B4FFA2FFCFFE165868E0259503" author="Ingrisch, S." box="[389,551,1117,1143]" pageId="8" pageNumber="185" pagination="133 - 227" refId="ref5621" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006</bibRefCitation>
</taxonomicName>
from Sulawesi and in
<taxonomicName id="4C5367C6FFA2FFCFFCB55868E6BA9503" authority="Ingrisch, 2006" authorityName="Ingrisch" authorityYear="2006" box="[806,1208,1117,1143]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="consternus">
<emphasis id="B927C057FFA2FFCFFCB55868E60C9503" box="[806,1038,1117,1143]" italics="true" pageId="8" pageNumber="185">Ornebius consternus</emphasis>
<bibRefCitation id="EFC261B4FFA2FFCFFB855868E6BA9503" author="Ingrisch, S." box="[1046,1208,1117,1143]" pageId="8" pageNumber="185" pagination="133 - 227" refId="ref5621" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006</bibRefCitation>
</taxonomicName>
from Sulawesi), the FW colouration darker (orange in
<taxonomicName id="4C5367C6FFA2FFCFFD8058B7E08C95EF" authorityName="Ingrisch" authorityYear="2006" box="[531,654,1153,1180]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="dumoga">
<emphasis id="B927C057FFA2FFCFFD8058B7E08C95EF" box="[531,654,1153,1180]" italics="true" pageId="8" pageNumber="185">O. dumoga</emphasis>
</taxonomicName>
and in
<taxonomicName id="4C5367C6FFA2FFCFFD7358B4E66E95EF" authority="Ingrisch, 1987" authorityName="Ingrisch" authorityYear="1987" box="[736,1132,1153,1179]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="rufonigrus">
<emphasis id="B927C057FFA2FFCFFD7358B4E1C795EF" box="[736,965,1153,1179]" italics="true" pageId="8" pageNumber="185">Ornebius rufonigrus</emphasis>
<bibRefCitation id="EFC261B4FFA2FFCFFC5858B4E66E95EF" author="Ingrisch, S." box="[971,1132,1153,1179]" pageId="8" pageNumber="185" pagination="163 - 185" refId="ref5553" refString="Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana Biologica, 68, 163 - 185." type="journal article" year="1987">Ingrisch, 1987</bibRefCitation>
</taxonomicName>
from Indochina and Malay Peninsula) and less covered by the pronotum (compared to
<taxonomicName id="4C5367C6FFA2FFCFFCAB5893E1D695CB" authorityName="Ingrisch" authorityYear="2006" box="[824,980,1190,1216]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="consternus">
<emphasis id="B927C057FFA2FFCFFCAB5893E1D695CB" box="[824,980,1190,1216]" italics="true" pageId="8" pageNumber="185">O. consternus</emphasis>
</taxonomicName>
). The male genitalia of this new species differ from congeners from the species group by the internal sclerites at base being smaller instead of expanded.
</paragraph>
<paragraph id="8BEC1C45FFA2FFCFFF5458DBE6919753" blockId="8.[151,1437,865,2008]" pageId="8" pageNumber="185">
We also compared the new species with other species currently not placed in any species group (i.e., with unknown male genitalia). Our species resembles species with wholly dark/ black or smoky FWs, such as
<taxonomicName id="4C5367C6FFA2FFCFFAA75924E3F7943B" authority="(Chopard, 1930)" baseAuthorityName="Chopard" baseAuthorityYear="1930" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B927C057FFA2FFCFFAA75924E336943B" italics="true" pageId="8" pageNumber="185">Ornebius obscuripennis</emphasis>
(
<bibRefCitation id="EFC261B4FFA2FFCFFED05900E3EF943B" author="Chopard, L." box="[323,493,1333,1359]" pageId="8" pageNumber="185" pagination="1 - 42" refId="ref5367" refString="Chopard, L. (1930) The Gryllidae of Sarawak. The Sarawak Museum Journal, 4 (12), 1 - 42." type="journal article" year="1930">Chopard, 1930</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F3445CD5FFA2FFCFFDA95900E098943B" box="[570,666,1333,1359]" name="Malaysia" pageId="8" pageNumber="185">Sarawak</collectingCountry>
and
<taxonomicName id="4C5367C6FFA2FFCFFD425900E682943B" authority="(Chopard, 1929)" baseAuthorityName="Chopard" baseAuthorityYear="1929" box="[721,1152,1333,1359]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="B927C057FFA2FFCFFD425900E1BD943B" box="[721,959,1333,1359]" italics="true" pageId="8" pageNumber="185">Ornebius fuscipennis</emphasis>
(
<bibRefCitation id="EFC261B4FFA2FFCFFC5D5900E675943B" author="Chopard, L." box="[974,1143,1333,1359]" pageId="8" pageNumber="185" pagination="98 - 118" refId="ref5343" refString="Chopard, L. (1929) Spolia Mentawiensia: Gryllidae. Bulletin of the Raffles Museum, 2, 98 - 118." type="journal article" year="1929">Chopard, 1929</bibRefCitation>
)
</taxonomicName>
from Sumatra (including Padang, Pulau Sipura, Pulau Telo and Batu Islands). The new species differs from
<taxonomicName id="4C5367C6FFA2FFCFFBBD596FE6EF9407" baseAuthorityName="Chopard" baseAuthorityYear="1930" box="[1070,1261,1369,1396]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B927C057FFA2FFCFFBBD596FE6EF9407" box="[1070,1261,1369,1396]" italics="true" pageId="8" pageNumber="185">O. obscuripennis</emphasis>
</taxonomicName>
by the pronotal lateral lobe and lateral parts of the head being dark-grey (i.e., unicolorous with dorsal parts) instead of forming a yellow lateral band. It also differs from
<taxonomicName id="4C5367C6FFA2FFCFFDCB5997E0FB94CF" baseAuthorityName="Chopard" baseAuthorityYear="1929" box="[600,761,1441,1468]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="B927C057FFA2FFCFFDCB5997E0FB94CF" box="[600,761,1441,1468]" italics="true" pageId="8" pageNumber="185">O. fuscipennis</emphasis>
</taxonomicName>
by much smaller body size (BL 8.4
<emphasis id="B927C057FFA2FFCFFB005997E6B094CF" box="[1171,1202,1442,1467]" italics="true" pageId="8" pageNumber="185">vs.</emphasis>
<quantity id="4CABB1A0FFA2FFCFFB2A5994E72394C8" box="[1209,1313,1441,1468]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="185" unit="mm" value="10.5">10.5 mm</quantity>
; PL 2.8
<emphasis id="B927C057FFA2FFCFFAED5997E79E94CF" box="[1406,1436,1442,1467]" italics="true" pageId="8" pageNumber="185">vs.</emphasis>
<quantity id="4CABB1A0FFA2FFCFFF0459F0E2ED9494" box="[151,239,1477,1504]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="185" unit="mm" value="4.5">4.5 mm</quantity>
; TL 2.5
<emphasis id="B927C057FFA2FFCFFED959F3E36B94AB" box="[330,361,1478,1503]" italics="true" pageId="8" pageNumber="185">vs.</emphasis>
<quantity id="4CABB1A0FFA2FFCFFEFC59F0E3B09494" box="[367,434,1477,1504]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="185" unit="mm" value="2.0">2 mm</quantity>
; FIIIL 4.4
<emphasis id="B927C057FFA2FFCFFDBB59F3E04494AB" box="[552,582,1478,1503]" italics="true" pageId="8" pageNumber="185">vs.</emphasis>
<quantity id="4CABB1A0FFA2FFCFFDDF59F3E09394AB" box="[588,657,1477,1504]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="185" unit="mm" value="7.0">7 mm</quantity>
); by the FW more uniformly coloured instead of smoky with posterior margin blackish; by the apical segment of the maxillary palps longer than subapical and third; by the pronotal disc not covered with yellow-brown scales; and by the legs that are not unicolourous yellowish.
</paragraph>
<paragraph id="8BEC1C45FFA2FFCFFF545A04E77897AF" blockId="8.[151,1437,865,2008]" pageId="8" pageNumber="185">
Lastly, the new species also differs from a syntopic species,
<taxonomicName id="4C5367C6FFA2FFCFFCD75A04E707973F" authority="Tan &amp; Robillard, 2012" authorityName="Tan &amp; Robillard" authorityYear="2012" box="[836,1285,1585,1612]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="tampines">
<emphasis id="B927C057FFA2FFCFFCD75A04E60D973F" box="[836,1039,1585,1611]" italics="true" pageId="8" pageNumber="185">Ornebius tampines</emphasis>
<bibRefCitation id="EFC261B4FFA2FFCFFB875A07E707973F" author="Tan, M. K. &amp; Robillard, T." box="[1044,1285,1585,1612]" pageId="8" pageNumber="185" pagination="411 - 420" refId="ref6458" refString="Tan, M. K. &amp; Robillard, T. (2012) Two new cricket species (Orthoptera: Gryllidae and Mogoplistidae) from the mangrove areas of Singapore. The Raffles Bulletin of Zoology, 60 (2), 411 - 420." type="journal article" year="2012">Tan &amp; Robillard, 2012</bibRefCitation>
</taxonomicName>
by the overall body and FW colouration, the body shape is more typical of
<taxonomicName id="4C5367C6FFA2FFCFFCB95A60E18C971B" authorityName="Guérin-Méneville" authorityYear="1844" box="[810,910,1621,1647]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis id="B927C057FFA2FFCFFCB95A60E18C971B" box="[810,910,1621,1647]" italics="true" pageId="8" pageNumber="185">Ornebius</emphasis>
</taxonomicName>
, instead of slenderly built, the shape of the male genitalia and by the calling songs (see Tan
<emphasis id="B927C057FFA2FFCFFDFE5A4FE09C97E7" box="[621,670,1657,1683]" italics="true" pageId="8" pageNumber="185">et al</emphasis>
., under review). It is also somewhat similar to a sympatric species,
<taxonomicName id="4C5367C6FFA2FFCFFA115A4FE2DA97C3" authorityName="Ingrisch" authorityYear="2006" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="pullus">
<emphasis id="B927C057FFA2FFCFFA115A4FE2DA97C3" italics="true" pageId="8" pageNumber="185">O. pullus</emphasis>
</taxonomicName>
by the wholly black FWs, but differs by the lack of white margins along the lateral lobe and the posterior margin of the dorsal disc of the pronotum, by the shape of male genitalia and by the calling songs (see Tan
<emphasis id="B927C057FFA2FFCFFB315AF7E6D297AF" box="[1186,1232,1729,1755]" italics="true" pageId="8" pageNumber="185">et al</emphasis>
., under review)
</paragraph>
</subSubSection>
<subSubSection id="C3494FCEFFA2FFCFFF545AD0E7649674" box="[199,1382,1765,1792]" pageId="8" pageNumber="185" type="etymology">
<paragraph id="8BEC1C45FFA2FFCFFF545AD0E7649674" blockId="8.[151,1437,865,2008]" box="[199,1382,1765,1792]" pageId="8" pageNumber="185">
<emphasis id="B927C057FFA2FFCFFF545AD0E34C978B" bold="true" box="[199,334,1765,1791]" pageId="8" pageNumber="185">Etymology.</emphasis>
The species name refers to the dark-grey colouration, resembling the grey-wolf,
<taxonomicName id="4C5367C6FFA2FFCFFB495AD3E75D978B" authorityName="Linnaeus" authorityYear="1758" box="[1242,1375,1765,1792]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="8" pageNumber="185" phylum="Chordata" rank="species" species="lupus">
<emphasis id="B927C057FFA2FFCFFB495AD3E75D978B" box="[1242,1375,1765,1792]" italics="true" pageId="8" pageNumber="185">Canis lupus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3494FCEFFA2FFCDFF545B3CE01492D1" lastPageId="10" lastPageNumber="187" pageId="8" pageNumber="185" type="description">
<paragraph id="8BEC1C45FFA2FFCDFF545B3CE7389011" blockId="8.[151,1437,865,2008]" lastBlockId="10.[151,1437,151,1221]" lastPageId="10" lastPageNumber="187" pageId="8" pageNumber="185">
<emphasis id="B927C057FFA2FFCFFF545B3CE35B9657" bold="true" box="[199,345,1801,1827]" pageId="8" pageNumber="185">Description.</emphasis>
Small species; habitus of male as shown in
<figureCitation id="136800C0FFA2FFCFFCDD5B3CE1CD9657" box="[846,975,1801,1827]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[184,1408,198,1900]" captionTargetId="figure-21@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 1. ♂ habitus of Cycloptiloides bimaculatus Tan et al., 2021(A), Ornebius pullus Ingrisch, 2006 (B) and Ornebius lupus sp. nov. (C, D) in dorsal (AC) and dorso-lateral view (D). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7354379" httpUri="https://zenodo.org/record/7354379/files/figure.png" pageId="8" pageNumber="185">Fig 1C, 1D</figureCitation>
. Dorsum of head flattened, covered with dark-grey scales (
<figureCitation id="136800C0FFA2FFCFFEC95B18E3BC9633" box="[346,446,1837,1863]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="8" pageNumber="185">Figs. 6A</figureCitation>
). Frontal rostrum 2.0 times wider than scapus (
<figureCitation id="136800C0FFA2FFCFFC505B18E6199633" box="[963,1051,1837,1863]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="8" pageNumber="185">Fig. 6A</figureCitation>
). Scapus dark-grey (
<figureCitation id="136800C0FFA2FFCFFB6E5B18E74D9633" box="[1277,1359,1837,1863]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="8" pageNumber="185">Fig 6A</figureCitation>
); basal antennal segments black, thereafter brown with some segments black (
<figureCitation id="136800C0FFA2FFCFFC235B64E63E961F" box="[944,1084,1873,1899]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="8" pageNumber="185">Figs 6B, 6C</figureCitation>
). Frons and mouthparts of dark colour (
<figureCitation id="136800C0FFA2FFCFFF7D5B40E34496FB" box="[238,326,1909,1935]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="8" pageNumber="185">Fig. 6B</figureCitation>
). Maxillary palps pale grey; with apical (= fifth) segment longer than subapical (= fourth) segment, third segment of subequal length as subapical segment; apical segment pyriform (
<figureCitation id="136800C0FFA2FFCFFBA95BACE69696C7" box="[1082,1172,1945,1971]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="8" pageNumber="185">Fig. 6C</figureCitation>
). Lateral parts of head, including genae, of dark colour, without bands (
<figureCitation id="136800C0FFA2FFCFFD255B88E10D96A3" box="[694,783,1981,2007]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="8" pageNumber="185">Fig. 6C</figureCitation>
). Pronotal disc brown and wholly covered with dark-grey scales, about 1.3 times longer than wide, with anterior margin narrow and straight; lateral margin widening slightly posteriorly; covering only base of mirror of FW; posterior margin convex (
<figureCitation id="136800C0FFA0FFCDFC735C8EE63991A1" box="[992,1083,187,213]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="10" pageNumber="187">Fig. 6D</figureCitation>
). Lateral lobe of pronotum also wholly covered with dark-grey scales (
<figureCitation id="136800C0FFA0FFCDFDC35CEAE0AC918D" box="[592,686,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[184,1408,198,1900]" captionTargetId="figure-21@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 1. ♂ habitus of Cycloptiloides bimaculatus Tan et al., 2021(A), Ornebius pullus Ingrisch, 2006 (B) and Ornebius lupus sp. nov. (C, D) in dorsal (AC) and dorso-lateral view (D). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7354379" httpUri="https://zenodo.org/record/7354379/files/figure.png" pageId="10" pageNumber="187">Figs 1D</figureCitation>
,
<figureCitation id="136800C0FFA0FFCDFD285CEAE0E2918D" box="[699,736,223,249]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="10" pageNumber="187">6C</figureCitation>
). TI with internal tympanum small and oval; without external tympanum. FIII 1.41.5 times longer than TIII; TIII 2.6 times longer than MTaIII. Femora generally pale, covered with marmoration of dark-grey scales except in basal parts. Tibiae and tarsi generally grey. Abdominal tergites covered with dark-grey scales; sternites pale. Abdominal apex darkened. Cercus covered with grey scales.
</paragraph>
<caption id="DF2C4CCDFFA3FFCEFF045B29E1BB9608" ID-DOI="http://doi.org/10.5281/zenodo.7354397" ID-Zenodo-Dep="7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="9" pageNumber="186" startId="9.[151,250,1820,1844]" targetBox="[151,1436,189,1797]" targetPageId="9">
<paragraph id="8BEC1C45FFA3FFCEFF045B29E1BB9608" blockId="9.[151,1436,1820,1917]" pageId="9" pageNumber="186">
<emphasis id="B927C057FFA3FFCEFF045B29E3169640" bold="true" box="[151,276,1820,1844]" pageId="9" pageNumber="186">FIGURE 6.</emphasis>
<taxonomicName id="4C5367C6FFA3FFCEFE8F5B28E3BB9641" authority="Tan &amp; Japir &amp; Chung &amp; Wahab &amp; Robillard, 2022" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab &amp; Robillard" authorityYear="2022" box="[284,441,1821,1845]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="B927C057FFA3FFCEFE8F5B28E3BB9641" box="[284,441,1821,1845]" italics="true" pageId="9" pageNumber="186">Ornebius lupus</emphasis>
</taxonomicName>
<emphasis id="B927C057FFA3FFCEFE535B28E0109641" bold="true" box="[448,530,1821,1845]" pageId="9" pageNumber="186">
<taxonomicNameLabel id="A2147D2CFFA3FFCEFE535B28E0109641" box="[448,530,1821,1845]" pageId="9" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BEC1C45FFA0FFCDFF545D5AE1DF9261" blockId="10.[151,1437,151,1221]" pageId="10" pageNumber="187">
<emphasis id="B927C057FFA0FFCDFF545D5AE30990FD" bold="true" box="[199,267,367,393]" pageId="10" pageNumber="187">Male.</emphasis>
FW mostly infumated black, dorsal field darkest at posterior margin (
<figureCitation id="136800C0FFA0FFCDFBB45D5AE68090FD" box="[1063,1154,367,393]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="10" pageNumber="187">Fig. 6D</figureCitation>
); lateral field likewise in some areas with infumated white veins (
<figureCitation id="136800C0FFA0FFCDFDC75DA6E0A890D9" box="[596,682,403,429]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="10" pageNumber="187">Fig. 6E</figureCitation>
). Supra-anal plate with last abdominal tergite and epiproct distinctly separated by a transverse suture. Last abdominal tergite transverse, posterior margin abruptly indented in middle; with oval pale spot in middle (not obvious in dry-pinned specimen) (
<figureCitation id="136800C0FFA0FFCDFC015DEEE1E19081" box="[914,995,475,501]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="10" pageNumber="187">Fig. 6F</figureCitation>
<emphasis id="B927C057FFA0FFCDFC7A5DEEE60A9080" box="[1001,1032,475,500]" italics="true" pageId="10" pageNumber="187">vs.</emphasis>
6G); posterior margin on either side of the indentation with two patches of short and strong setae. Supra-anal plate less transverse than last abdominal tergite, black coloured, apex subtruncate (
<figureCitation id="136800C0FFA0FFCDFDE35E16E0F89349" box="[624,762,547,573]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="10" pageNumber="187">Figs 6F, 6G</figureCitation>
). Subgenital plate black (
<figureCitation id="136800C0FFA0FFCDFB895E16E6779349" box="[1050,1141,547,573]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="10" pageNumber="187">Fig. 6H</figureCitation>
). Paraproct process black, elongate, cylindrical and tapering into a subacute apex (
<figureCitation id="136800C0FFA0FFCDFC975E72E15F9315" box="[772,861,583,609]" captionStart="FIGURE 6" captionStartId="9.[151,250,1820,1844]" captionTargetBox="[151,1436,189,1797]" captionTargetId="figure-19@9.[151,1436,181,1797]" captionTargetPageId="9" captionText="FIGURE 6. Ornebius lupus sp. nov. ♂ holotype: head and pronotum in dorsal view (A); head in anterior (B) and lateral (C) views; FWs in dorsal (D) and lateral (E) views; abdominal apex in dorsal (F, G) and lateral (H) views. (F) shows the specimen preserved in ethanol and (G) shows the specimen dry-pinned. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354397" httpUri="https://zenodo.org/record/7354397/files/figure.png" pageId="10" pageNumber="187">Fig. 6H</figureCitation>
). Genitalia as in
<figureCitation id="136800C0FFA0FFCDFB845E72E6599315" box="[1047,1115,583,609]" captionStart="FIGURE 7" captionStartId="10.[151,250,1947,1971]" captionTargetBox="[156,1426,1252,1923]" captionTargetId="figure-467@10.[151,1436,1245,1924]" captionTargetPageId="10" captionText="FIGURE 7. Ornebius lupus sp. nov. ♂ holotype genitalia in dorsal (A, B) and lateral (C) views. es = external sclerite, is = internal sclerite, lv = lateral valve, mv = medial valve. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7354402" httpUri="https://zenodo.org/record/7354402/files/figure.png" pageId="10" pageNumber="187">Fig. 7</figureCitation>
. Central lobe of phallus with lateral valves (lv) elongate, apical area curved slightly ventrad in lateral view; tapering into obtuse apex, in apical area with inner margin curved in dorsal view; both sides together almost forming a dorsally and ventrally open tube. External sclerite (es) well developed, with external part more strongly sclerotized than inner part (in dorsal view). Medial valve (mv) membranous with base widened (in dorsal view) and upcurved (in lateral view). Internal sclerites (is) at base small but widened and curled, afterwards straight and elongate.
</paragraph>
<paragraph id="8BEC1C45FFA0FFCDFF545F2AE3A0924D" blockId="10.[151,1437,151,1221]" box="[199,418,799,825]" pageId="10" pageNumber="187">
<emphasis id="B927C057FFA0FFCDFF545F2AE327924D" bold="true" box="[199,293,799,825]" pageId="10" pageNumber="187">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BEC1C45FFA0FFCDFF545F76E01492D1" blockId="10.[151,1437,151,1221]" pageId="10" pageNumber="187">
<emphasis id="B927C057FFA0FFCDFF545F76E0109228" bold="true" box="[199,530,834,864]" pageId="10" pageNumber="187">
Measurements (
<specimenCount id="9D55D7CCFFA0FFCDFE155F71E3AB9214" box="[390,425,836,864]" pageId="10" pageNumber="187" type="male">2♂</specimenCount>
, in mm).
</emphasis>
<specimenCount id="9D55D7CCFFA0FFCDFD8A5F76E02C9229" box="[537,558,835,861]" pageId="10" pageNumber="187" type="male"></specimenCount>
<typeStatus id="54E8A2E7FFA0FFCDFDA75F76E0A39229" box="[564,673,835,861]" pageId="10" pageNumber="187" type="holotype">Holotype</typeStatus>
: BL = 8.4; PronL = 2.8; PronW = 2.2; FWL = 2.5; FWW = 2.3; FIIIL = 4.4; TIIIL = 3.1; MaTIIIL = 1.2.
<specimenCount id="9D55D7CCFFA0FFCDFD895F52E02D92F5" box="[538,559,871,897]" pageId="10" pageNumber="187" type="male"></specimenCount>
<typeStatus id="54E8A2E7FFA0FFCDFDA65F52E09892F5" box="[565,666,871,897]" pageId="10" pageNumber="187" type="paratype">paratype</typeStatus>
: BL = 8.4; PronL = 2.8; PronW = 2.2; FWL = 2.5; FWW = 2.3; FIIIL = 4.4; TIIIL = 2.9; MaTIIIL = 1.1.
</paragraph>
</subSubSection>
<subSubSection id="C3494FCEFFA0FFCDFF545F9AE3EB9299" pageId="10" pageNumber="187" type="biology_ecology">
<paragraph id="8BEC1C45FFA0FFCDFF545F9AE3EB9299" blockId="10.[151,1437,151,1221]" pageId="10" pageNumber="187">
<emphasis id="B927C057FFA0FFCDFF545F9AE32E92BD" bold="true" box="[199,300,943,969]" pageId="10" pageNumber="187">Ecology.</emphasis>
This species is found among foliage of mangrove trees, occurring in syntopy with another mangrovespecialist,
<taxonomicName id="4C5367C6FFA0FFCDFE9E5FE6E3E09298" authorityName="Tan &amp; Robillard" authorityYear="2012" box="[269,482,979,1005]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="187" phylum="Arthropoda" rank="species" species="tampines">
<emphasis id="B927C057FFA0FFCDFE9E5FE6E3E09298" box="[269,482,979,1005]" italics="true" pageId="10" pageNumber="187">Ornebius tampines</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3494FCEFFA0FFCDFF545FC3E0069565" box="[199,516,1014,1041]" pageId="10" pageNumber="187" type="distribution">
<paragraph id="8BEC1C45FFA0FFCDFF545FC3E0069565" blockId="10.[151,1437,151,1221]" box="[199,516,1014,1041]" pageId="10" pageNumber="187">
<emphasis id="B927C057FFA0FFCDFF545FC3E3639564" bold="true" box="[199,353,1014,1040]" pageId="10" pageNumber="187">Distribution.</emphasis>
<collectingCountry id="F3445CD5FFA0FFCDFEF45FC2E0069565" box="[359,516,1015,1041]" name="Singapore" pageId="10" pageNumber="187">SINGAPORE</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C3494FCEFFA0FFCDFF54582EE37095B1" pageId="10" pageNumber="187" type="materials_examined">
<paragraph id="8BEC1C45FFA0FFCDFF54582EE0F19541" blockId="10.[151,1437,151,1221]" box="[199,755,1050,1077]" pageId="10" pageNumber="187">
<emphasis id="B927C057FFA0FFCDFF54582EE3649540" bold="true" box="[199,358,1050,1077]" pageId="10" pageNumber="187">Type locality.</emphasis>
<collectingCountry id="F3445CD5FFA0FFCDFEFF582EE00E9541" box="[364,524,1051,1077]" name="Singapore" pageId="10" pageNumber="187">SINGAPORE</collectingCountry>
, Pasir Ris mangrove
</paragraph>
<paragraph id="8BEC1C45FFA0FFCDFF54580BE37095B1" blockId="10.[151,1437,151,1221]" pageId="10" pageNumber="187">
<emphasis id="B927C057FFA0FFCDFF54580BE364952D" bold="true" box="[199,358,1086,1113]" pageId="10" pageNumber="187">Calling song.</emphasis>
Described in Tan
<emphasis id="B927C057FFA0FFCDFDD3580AE077952D" box="[576,629,1087,1113]" italics="true" pageId="10" pageNumber="187">et al</emphasis>
. (under review). The calling song of this species is similar to that of
<taxonomicName id="4C5367C6FFA0FFCDFF045856E02C9509" authority="Ingrisch, 1987" authorityName="Ingrisch" authorityYear="1987" box="[151,558,1123,1149]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="187" phylum="Arthropoda" rank="species" species="rufonigrus">
<emphasis id="B927C057FFA0FFCDFF045856E37D9509" box="[151,383,1123,1149]" italics="true" pageId="10" pageNumber="187">Ornebius rufonigrus</emphasis>
<bibRefCitation id="EFC261B4FFA0FFCDFE1A5856E02C9509" author="Ingrisch, S." box="[393,558,1123,1149]" pageId="10" pageNumber="187" pagination="163 - 185" refId="ref5553" refString="Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana Biologica, 68, 163 - 185." type="journal article" year="1987">Ingrisch, 1987</bibRefCitation>
</taxonomicName>
from the same species group with regard to echeme duration and dominant frequency, although the new species produces echeme-sequences composed of triple-chirps instead of double-chrips as in
<taxonomicName id="4C5367C6FFA0FFCDFF41589EE36995B1" authorityName="Ingrisch" authorityYear="1987" box="[210,363,1195,1221]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="187" phylum="Arthropoda" rank="species" species="rufonigrus">
<emphasis id="B927C057FFA0FFCDFF41589EE36995B1" box="[210,363,1195,1221]" italics="true" pageId="10" pageNumber="187">O. rufonigrus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>