374 lines
51 KiB
XML
374 lines
51 KiB
XML
<document id="606B3E7A7FE1FB61B426AA82D48F69D8" ID-DOI="10.5281/zenodo.5374745" ID-ISSN="1638-9395" ID-Zenodo-Dep="5374745" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630276855747" checkinUser="marcus" docAuthor="Märss, Tiiu & Karatajūtē-Talimaa, Valentina" docDate="2002" docId="03FA0B64FF8CE024FC81AC649AD8BD6C" docLanguage="en" docName="Geodiversitas.2002.24.2.381-404.pdf" docOrigin="Geodiversitas 24 (2)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Shielia multispinata Märss & Karatajūtē-Talimaa 2002, n. sp." docType="treatment" docVersion="3" lastPageNumber="393" masterDocId="FFC3731CFF84E028FFC5A94F9879BC36" masterDocTitle="Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)" masterLastPageNumber="404" masterPageNumber="381" pageNumber="389" updateTime="1699210438067" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods id="1D68567DCDD44EB5C730D81C078F6B5C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="303A23FDBB6F4B6DF87424D883807379">
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<mods:title id="0F2690EC191C5ABE91A9815129A1B6B4">Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)</mods:title>
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</mods:titleInfo>
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<mods:name id="E4E8F1578F9A8487E9A96DA6574496D9" type="personal">
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<mods:role id="B95BD0767049EBF7D0DBE9B2FEE203FD">
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<mods:roleTerm id="74BD9BA10C400DC3094D07153B8F66C9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FC016113F6325418368E8021B9886AA0">Märss, Tiiu</mods:namePart>
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<mods:affiliation id="39E9E4AF4A711AE05AA946F539A8A813">Institute of Geology, Tallinn Technical University, 7 Estonia Avenue, Tallinn 10143 (Estonia) marss @ gi. ee</mods:affiliation>
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<mods:nameIdentifier id="E5EBA142778E4D1686D9586B890EE2AA" type="email">marss@gi.ee</mods:nameIdentifier>
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</mods:name>
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<mods:name id="3F0AE8E609745A1FDFD43E9681444709" type="personal">
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<mods:role id="BE353B84A7ED607A76BD96ED08A1D8E8">
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<mods:roleTerm id="F8C8169DD0FB1144C120A07495CE0A72">Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart id="8949F5D8D66CFE4BAB934E786C8948EF">Karatajūtē-Talimaa, Valentina</mods:namePart>
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<mods:affiliation id="D980F7FB4D91BABB5BCE9CA0A52B24E3">Institute of Geology of Lithuania, S ˘ evc enkos 13, Vilnius LR 2600 (Lithuania) vtalimaa @ takas. lt Six new and one earlier known thelodont from the Ordovician and Lower Silurian of Severnaya Zemlya Archipelago are described. Scales of the earliest dated thelodont Stroinolepis maenniki n. gen., n. sp. were obtained from the Strojnaya River section, October Revolution Island, upper Ozernaya and Strojnaya formations, Middle-Upper and Upper Ordovician, correspondingly. Loganellia matura n. sp. was found from the Srednii Island, Golomyannyj Formation, middle Llandovery. Paralogania klubovi n. sp. and Loganelliidae gen. et sp. indet. come from the middle (?) Llandovery of Pioneer Island. L. grossi Fredholm, 1990 in the Matusevich River section, Paralogania consimilis n. sp. and Thelodus calvus n. sp. in the Ushakov River section, and Shielia multispinata n. sp., in the Spokojnaya River section, are distributed in the Samojlovich Formation, Wenlock, Lower Silurian of October Revolution Island.</mods:affiliation>
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<mods:nameIdentifier id="1675A0446F1474A7B858E982137E9DFD" type="email">vtalimaa@takas.lt</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="6C06785FDC3373E2B525ECEE4CE80576">text</mods:typeOfResource>
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<mods:relatedItem id="E316EB59DDF738FF35BE779760FADBCA" type="host">
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<mods:titleInfo id="E27F8E4C7A26CD6E0ECC89E852E508E6">
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<mods:title id="605A2FBEA8FD33F3521D780A0575D033">Geodiversitas</mods:title>
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||
</mods:titleInfo>
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<mods:part id="464372CF8DFECF059987C1851670EC15">
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<mods:date id="68946FC2D55E00D37634FAF019A04ED1">2002</mods:date>
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||
<mods:detail id="B5E3CF793ED43EEA8341A8A3BED120EA" type="volume">
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<mods:number id="F1211BEFEDDC51C8DBBC315CB18AB76A">24</mods:number>
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||
</mods:detail>
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<mods:detail id="5E358BCE4EB87ED8DD6820466C5366A0" type="issue">
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<mods:number id="45E863A13EADE99B32D2C5DFCCC14852">2</mods:number>
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</mods:detail>
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<mods:extent id="1B7E2344AD558F0018A2E79AF0B4466E" unit="page">
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<mods:start id="5B631A9F1C0EDC27B333964EC85FFDA8">381</mods:start>
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<mods:end id="EFA8ACBC37C1082512D573BCD418559E">404</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="044D7CC2489E0667364EA7CAB812FE76">journal article</mods:classification>
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<mods:identifier id="BA93FC855A8D1DE2F47D863EAB645DFD" type="DOI">10.5281/zenodo.5374745</mods:identifier>
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<mods:identifier id="17CF588FD05DEF2D37286E0A4B16D8C0" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="E17489E48B01FA165A61BB7E0510CDB6" type="Zenodo-Dep">5374745</mods:identifier>
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</mods:mods>
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<treatment id="03FA0B64FF8CE024FC81AC649AD8BD6C" LSID="urn:lsid:plazi:treatment:03FA0B64FF8CE024FC81AC649AD8BD6C" httpUri="http://treatment.plazi.org/id/03FA0B64FF8CE024FC81AC649AD8BD6C" lastPageId="12" lastPageNumber="393" pageId="8" pageNumber="389">
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<subSubSection id="C349E9F9FF8CE020FC81AC649C41B955" pageId="8" pageNumber="389" type="nomenclature">
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<paragraph id="8BECBA72FF8CE020FC81AC649C41B955" blockId="8.[836,1126,1323,1379]" pageId="8" pageNumber="389">
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||
<taxonomicName id="4C53C1F1FF8CE020FC81AC649C5CB975" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[836,1061,1323,1347]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="multispinata" status="sp. nov.">
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<emphasis id="B9276660FF8CE020FC81AC649C5CB975" bold="true" box="[836,1061,1323,1347]" italics="true" pageId="8" pageNumber="389">Shielia multispinata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A214DB1BFF8CE020FBEEAC639C1FB975" box="[1067,1126,1324,1347]" pageId="8" pageNumber="389" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="1368A6F7FF8CE020FCBCAC049BC4B955" box="[889,957,1355,1379]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[208,1202,174,1484]" captionTargetId="figure-25@9.[206,1208,194,1502]" captionTargetPageId="9" captionText="FIG. 6. — Microstructure of the scales; A, B, Loganellia matura n. sp.; C-F, Shielia multispinata n. sp.; G, Paralogania consimilis n. sp.; A, Pi 7624; B, Pi 7625; C, Pi 7626; D, Pi 7627; E, Pi 7628; F, Pi 7629; G, Pi 7630; A, B, sample MF 3-8, Srednii Island, Golomyannyj Formation, middle part of Llandovery, Lower Silurian; C-F, sample MF 157-7, Spokojnaya River, upper part of Samojlovich Formation, Wenlock, Lower Silurian; G, sample MF 46-23, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; bc, canal in the base; cr, crown; dc, dentine canal; dt, dentine tubules; fpc, fan-shaped pulp canals; lac, lacunae of dentine canals; mpc, main pulp canal; n, neck; pc, pulp canal; pco, pulp canal opening; sp, spine. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374757" httpUri="https://zenodo.org/record/5374757/files/figure.png" pageId="8" pageNumber="389">Figs 6</figureCitation>
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C-F; 7A-S)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C349E9F9FF8CE020FD0FACC29C39B9E3" pageId="8" pageNumber="389" type="materials_examined">
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<paragraph id="8BECBA72FF8CE020FD0FACC29C39B9E3" blockId="8.[714,1247,1420,1493]" pageId="8" pageNumber="389">
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<materialsCitation id="3B3BB02FFF8CE020FD0FACC29C39B9E3" county="Lower Silurian." location="Pi" municipality="Wenlock" pageId="8" pageNumber="389" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E804D0FF8CE020FD0FACC29B43B997" box="[714,826,1421,1442]" pageId="8" pageNumber="389" type="holotype">HOLOTYPE</typeStatus>
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. —
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<location id="8E8CECA9FF8CE020FCA2ACC39B05B994" LSID="urn:lsid:plazi:treatment:03FA0B64FF8CE024FC81AC649AD8BD6C:8E8CECA9FF8CE020FCA2ACC39B05B994" box="[871,892,1420,1442]" county="Lower Silurian." municipality="Wenlock" name="Pi" pageId="8" pageNumber="389">Pi</location>
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6803 (
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<figureCitation id="1368A6F7FF8CE020FC07ACC39C6AB994" box="[962,1043,1420,1442]" captionStart="FIG" captionStartId="10.[151,162,1524,1541]" captionTargetBox="[203,1186,173,1467]" captionTargetId="figure-21@10.[203,1186,173,1467]" captionTargetPageId="10" captionText="FIG. 7. — A-S, Shielia multispinata n. sp.; A, head scale; B-D, F, transitional scales; E, G-P, trunk scales; Q-S, fin scales; A, B, scales in lateral view; C-N, P, Q, S, scales in crown view; O, a scale in oblique view; R, a scale in base view; A, Pi 7517; B, Pi 7519; C, Pi 7518; D, Pi 7520; E, Pi 7521; F, Pi 6804; G, Pi 7523; H, Pi 7522; I, Pi 7524; J, Pi 7525; K, holotype Pi 6803; L, Pi 6805; M, Pi 6801; N, O, Pi 7526; P, Pi 6802; Q, Pi 7527; R, Pi 7529; S, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; T-V, Thelodus? sp., scales in crown view; T, Pi 7516; U, LIG 35 908; V, Pi 6814; T, V, sample MF 46-23; U, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="8" pageNumber="389">Fig. 7K</figureCitation>
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), sample MF 157-7,
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<location id="8E8CECA9FF8CE020FD0FACEA9BE2B98D" LSID="urn:lsid:plazi:treatment:03FA0B64FF8CE024FC81AC649AD8BD6C:8E8CECA9FF8CE020FD0FACEA9BE2B98D" box="[714,923,1445,1467]" county="Lower Silurian." municipality="Wenlock" name="Spokojnaya River" pageId="8" pageNumber="389">Spokojnaya River</location>
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, upper part of
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<location id="8E8CECA9FF8CE020FB91ACEA9B4FB9E3" LSID="urn:lsid:plazi:treatment:03FA0B64FF8CE024FC81AC649AD8BD6C:8E8CECA9FF8CE020FB91ACEA9B4FB9E3" county="Lower Silurian." municipality="Wenlock" name="Samojlovich Formation" pageId="8" pageNumber="389">Samojlovich Formation</location>
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,
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<collectingMunicipality id="6B882008FF8CE020FC85ACF09BE5B9E3" box="[832,924,1471,1493]" pageId="8" pageNumber="389">Wenlock</collectingMunicipality>
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,
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<collectingCounty id="628DC2FEFF8CE020FC60ACF09C39B9E3" box="[933,1088,1471,1493]" pageId="8" pageNumber="389">Lower Silurian.</collectingCounty>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C349E9F9FF8CE020FD0FACA09C5FBA01" pageId="8" pageNumber="389" type="etymology">
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<paragraph id="8BECBA72FF8CE020FD0FACA09C5FBA01" blockId="8.[714,1247,1519,1591]" pageId="8" pageNumber="389">
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ETYMOLOGY. — From the Latin word
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<taxonomicName id="4C53C1F1FF8CE020FB9EACA09CA0BA32" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[1115,1241,1519,1540]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="multispinata">
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<emphasis id="B9276660FF8CE020FB9EACA09CA0BA32" box="[1115,1241,1519,1540]" italics="true" pageId="8" pageNumber="389">multispinata</emphasis>
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</taxonomicName>
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, to indicate that trunk scales of this species have several spines posteriorly of the crown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C349E9F9FF8CE023FD0FAF1D99EEBCC5" lastPageId="11" lastPageNumber="392" pageId="8" pageNumber="389" type="materials_examined">
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<paragraph id="8BECBA72FF8CE020FD0FAF1D9B93BAB7" blockId="8.[714,1247,1617,1665]" pageId="8" pageNumber="389">MATERIAL EXAMINED. — About 100 scales of good and fairly good preservation.</paragraph>
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<caption id="DF2CEAFAFF8DE021FF52ACBB9B4CBAB3" ID-DOI="http://doi.org/10.5281/zenodo.5374757" ID-Zenodo-Dep="5374757" httpUri="https://zenodo.org/record/5374757/files/figure.png" pageId="9" pageNumber="390" startId="9.[151,162,1524,1541]" targetBox="[208,1202,174,1484]" targetPageId="9">
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<paragraph id="8BECBA72FF8DE021FF52ACBB9B4CBAB3" blockId="9.[151,1248,1524,1669]" pageId="9" pageNumber="390">
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FIG. 6. — Microstructure of the scales;
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<emphasis id="B9276660FF8DE021FE2CACBB998FBA33" bold="true" box="[489,502,1524,1541]" pageId="9" pageNumber="390">A</emphasis>
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,
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<emphasis id="B9276660FF8DE021FDC7ACBB9A76BA33" bold="true" box="[514,527,1524,1541]" pageId="9" pageNumber="390">B</emphasis>
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,
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<taxonomicName id="4C53C1F1FF8DE021FDDEACBB9AC8BA33" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[539,689,1524,1541]" class="Pteraspidomorphi" family="Loganelliidae" genus="Loganellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="matura" status="sp. nov.">
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<emphasis id="B9276660FF8DE021FDDEACBB9AC8BA33" box="[539,689,1524,1541]" italics="true" pageId="9" pageNumber="390">Loganellia matura</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A214DB1BFF8DE021FD7DACBB9A95BA33" box="[696,748,1524,1541]" pageId="9" pageNumber="390" rank="species">n. sp.</taxonomicNameLabel>
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;
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<emphasis id="B9276660FF8DE021FD30ACBB9B6CBA33" bold="true" box="[757,789,1524,1541]" pageId="9" pageNumber="390">C -F</emphasis>
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,
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<taxonomicName id="4C53C1F1FF8DE021FCE4ACBB9BBFBA33" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[801,966,1524,1541]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="multispinata" status="sp. nov.">
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<emphasis id="B9276660FF8DE021FCE4ACBB9BBFBA33" box="[801,966,1524,1541]" italics="true" pageId="9" pageNumber="390">Shielia multispinata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A214DB1BFF8DE021FC09ACBB9C79BA33" box="[972,1024,1524,1541]" pageId="9" pageNumber="390" rank="species">n. sp.</taxonomicNameLabel>
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;
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<emphasis id="B9276660FF8DE021FBCCACBB9C6EBA33" bold="true" box="[1033,1047,1524,1541]" pageId="9" pageNumber="390">G</emphasis>
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,
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<taxonomicName id="4C53C1F1FF8DE021FBE6ACBB9CA6BA33" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[1059,1247,1524,1541]" class="Pteraspidomorphi" family="Loganelliidae" genus="Paralogania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="consimilis" status="sp. nov.">
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<emphasis id="B9276660FF8DE021FBE6ACBB9CA6BA33" box="[1059,1247,1524,1541]" italics="true" pageId="9" pageNumber="390">Paralogania consimilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A214DB1BFF8DE021FF52AF4698B2BA2C" box="[151,203,1545,1562]" pageId="9" pageNumber="390" rank="species">n. sp.</taxonomicNameLabel>
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;
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<emphasis id="B9276660FF8DE021FF11AF469898BA2C" bold="true" box="[212,225,1545,1562]" pageId="9" pageNumber="390">A</emphasis>
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, Pi 7624;
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<emphasis id="B9276660FF8DE021FEFFAF46993EBA2C" bold="true" box="[314,327,1545,1562]" pageId="9" pageNumber="390">B</emphasis>
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, Pi 7625;
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<emphasis id="B9276660FF8DE021FE64AF4699D6BA2C" bold="true" box="[417,431,1545,1562]" pageId="9" pageNumber="390">C</emphasis>
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, Pi 7626;
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<emphasis id="B9276660FF8DE021FDCCAF469A6EBA2C" bold="true" box="[521,535,1545,1562]" pageId="9" pageNumber="390">D</emphasis>
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, Pi 7627;
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<emphasis id="B9276660FF8DE021FDB4AF469A04BA2C" bold="true" box="[625,637,1545,1562]" pageId="9" pageNumber="390">E</emphasis>
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, Pi 7628;
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<emphasis id="B9276660FF8DE021FD12AF469A9BBA2C" bold="true" box="[727,738,1545,1562]" pageId="9" pageNumber="390">F</emphasis>
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, Pi 7629;
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<emphasis id="B9276660FF8DE021FCFEAF469B30BA2C" bold="true" box="[827,841,1545,1562]" pageId="9" pageNumber="390">G</emphasis>
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, Pi 7630;
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<emphasis id="B9276660FF8DE021FC66AF469BC9BA2C" bold="true" box="[931,944,1545,1562]" pageId="9" pageNumber="390">A</emphasis>
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,
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<emphasis id="B9276660FF8DE021FC78AF469BB3BA2C" bold="true" box="[957,970,1545,1562]" pageId="9" pageNumber="390">B</emphasis>
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, sample MF 3-8, Srednii Island, Golomyannyj Formation, middle part of Llandovery, Lower Silurian;
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<emphasis id="B9276660FF8DE021FD2BAF509B76BA06" bold="true" box="[750,783,1567,1584]" pageId="9" pageNumber="390">C -F</emphasis>
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, sample MF 157-7, Spokojnaya River, upper part of Samojlovich Formation, Wenlock, Lower Silurian;
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<emphasis id="B9276660FF8DE021FD85AF7B9A37BA73" bold="true" box="[576,590,1588,1605]" pageId="9" pageNumber="390">G</emphasis>
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, sample MF 46-23, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations:
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<emphasis id="B9276660FF8DE021FD7CAF069AA0BA6C" bold="true" box="[697,729,1609,1626]" pageId="9" pageNumber="390">asp</emphasis>
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, aspidine “pegs” in the base;
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<emphasis id="B9276660FF8DE021FC1EAF069B9FBA6C" bold="true" box="[987,998,1609,1626]" pageId="9" pageNumber="390">b</emphasis>
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, base;
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<emphasis id="B9276660FF8DE021FBE3AF069C45BA6C" bold="true" box="[1062,1084,1609,1626]" pageId="9" pageNumber="390">bc</emphasis>
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, canal in the base;
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<emphasis id="B9276660FF8DE021FF52AF1098D0BA46" bold="true" box="[151,169,1631,1648]" pageId="9" pageNumber="390">cr</emphasis>
|
||
, crown;
|
||
<emphasis id="B9276660FF8DE021FF35AF10997FBA46" bold="true" box="[240,262,1631,1648]" pageId="9" pageNumber="390">dc</emphasis>
|
||
, dentine canal;
|
||
<emphasis id="B9276660FF8DE021FE4FAF1099E5BA46" bold="true" box="[394,412,1631,1648]" pageId="9" pageNumber="390">dt</emphasis>
|
||
, dentine tubules;
|
||
<emphasis id="B9276660FF8DE021FDF5AF109A35BA46" bold="true" box="[560,588,1631,1648]" pageId="9" pageNumber="390">fpc</emphasis>
|
||
, fan-shaped pulp canals;
|
||
<emphasis id="B9276660FF8DE021FCE6AF109B44BA46" bold="true" box="[803,829,1631,1648]" pageId="9" pageNumber="390">lac</emphasis>
|
||
, lacunae of dentine canals;
|
||
<emphasis id="B9276660FF8DE021FBE0AF109C35BA46" bold="true" box="[1061,1100,1631,1648]" pageId="9" pageNumber="390">mpc</emphasis>
|
||
, main pulp canal;
|
||
<emphasis id="B9276660FF8DE021FF52AF3B98DBBAB3" bold="true" box="[151,162,1652,1669]" pageId="9" pageNumber="390">n</emphasis>
|
||
, neck;
|
||
<emphasis id="B9276660FF8DE021FF1AAF3B988CBAB3" bold="true" box="[223,245,1652,1669]" pageId="9" pageNumber="390">pc</emphasis>
|
||
, pulp canal;
|
||
<emphasis id="B9276660FF8DE021FEA5AF3B99FBBAB3" bold="true" box="[352,386,1652,1669]" pageId="9" pageNumber="390">pco</emphasis>
|
||
, pulp canal opening;
|
||
<emphasis id="B9276660FF8DE021FDF0AF3B9A33BAB3" bold="true" box="[565,586,1652,1669]" pageId="9" pageNumber="390">sp</emphasis>
|
||
, spine. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2CEAFAFF8EE022FF52ACBB9AB1BAB3" ID-DOI="http://doi.org/10.5281/zenodo.5374759" ID-Zenodo-Dep="5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="10" pageNumber="391" startId="10.[151,162,1524,1541]" targetBox="[203,1186,173,1467]" targetPageId="10">
|
||
<paragraph id="8BECBA72FF8EE022FF52ACBB9AB1BAB3" blockId="10.[151,1249,1524,1669]" pageId="10" pageNumber="391">
|
||
FIG. 7. —
|
||
<emphasis id="B9276660FF8EE022FF23ACBB997FBA33" bold="true" box="[230,262,1524,1541]" pageId="10" pageNumber="391">A -S</emphasis>
|
||
,
|
||
<taxonomicName id="4C53C1F1FF8EE022FED5ACBB99C9BA33" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[272,432,1524,1541]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="multispinata" status="sp. nov.">
|
||
<emphasis id="B9276660FF8EE022FED5ACBB99C9BA33" box="[272,432,1524,1541]" italics="true" pageId="10" pageNumber="391">Shielia multispinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214DB1BFF8EE022FE70ACBB999FBA33" box="[437,486,1524,1541]" pageId="10" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="B9276660FF8EE022FE28ACBB9983BA33" bold="true" box="[493,506,1524,1541]" pageId="10" pageNumber="391">A</emphasis>
|
||
, head scale;
|
||
<emphasis id="B9276660FF8EE022FDA2ACBB9AF3BA33" bold="true" box="[615,650,1524,1541]" pageId="10" pageNumber="391">B -D</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF8EE022FD51ACBB9AE6BA33" bold="true" box="[660,671,1524,1541]" pageId="10" pageNumber="391">F</emphasis>
|
||
, transitional scales;
|
||
<emphasis id="B9276660FF8EE022FC8DACBB9B2DBA33" bold="true" box="[840,852,1524,1541]" pageId="10" pageNumber="391">E</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF8EE022FC9BACBB9B06BA33" bold="true" box="[862,895,1524,1541]" pageId="10" pageNumber="391">G -P</emphasis>
|
||
, trunk scales;
|
||
<emphasis id="B9276660FF8EE022FC3DACBB9C63BA33" bold="true" box="[1016,1050,1524,1541]" pageId="10" pageNumber="391">Q -S</emphasis>
|
||
, fin scales;
|
||
<emphasis id="B9276660FF8EE022FBB9ACBB9CF0BA33" bold="true" box="[1148,1161,1524,1541]" pageId="10" pageNumber="391">A</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF8EE022FB56ACBB9CD9BA33" bold="true" box="[1171,1184,1524,1541]" pageId="10" pageNumber="391">B</emphasis>
|
||
, scales in lateral view;
|
||
<emphasis id="B9276660FF8EE022FED1AF46994EBA2C" bold="true" box="[276,311,1545,1562]" pageId="10" pageNumber="391">C -N</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF8EE022FE87AF469937BA2C" bold="true" box="[322,334,1545,1562]" pageId="10" pageNumber="391">P</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF8EE022FE9CAF469911BA2C" bold="true" box="[345,360,1545,1562]" pageId="10" pageNumber="391">Q</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF8EE022FEB6AF469906BA2C" bold="true" box="[371,383,1545,1562]" pageId="10" pageNumber="391">S</emphasis>
|
||
, scales in crown view;
|
||
<emphasis id="B9276660FF8EE022FD87AF469A28BA2C" bold="true" box="[578,593,1545,1562]" pageId="10" pageNumber="391">O</emphasis>
|
||
, a scale in oblique view;
|
||
<emphasis id="B9276660FF8EE022FCE0AF469B4BBA2C" bold="true" box="[805,818,1545,1562]" pageId="10" pageNumber="391">R</emphasis>
|
||
, a scale in base view;
|
||
<emphasis id="B9276660FF8EE022FC34AF469B87BA2C" bold="true" box="[1009,1022,1545,1562]" pageId="10" pageNumber="391">A</emphasis>
|
||
, Pi 7517;
|
||
<emphasis id="B9276660FF8EE022FB96AF469C19BA2C" bold="true" box="[1107,1120,1545,1562]" pageId="10" pageNumber="391">B</emphasis>
|
||
, Pi 7519;
|
||
<emphasis id="B9276660FF8EE022FB73AF469CBDBA2C" bold="true" box="[1206,1220,1545,1562]" pageId="10" pageNumber="391">C</emphasis>
|
||
, Pi 7518;
|
||
<emphasis id="B9276660FF8EE022FF0EAF5098A0BA06" bold="true" box="[203,217,1567,1584]" pageId="10" pageNumber="391">D</emphasis>
|
||
, Pi 7520;
|
||
<emphasis id="B9276660FF8EE022FEEBAF509943BA06" bold="true" box="[302,314,1567,1584]" pageId="10" pageNumber="391">E</emphasis>
|
||
, Pi 7521;
|
||
<emphasis id="B9276660FF8EE022FE4AAF5099E3BA06" bold="true" box="[399,410,1567,1584]" pageId="10" pageNumber="391">F</emphasis>
|
||
, Pi 6804;
|
||
<emphasis id="B9276660FF8EE022FE2AAF509984BA06" bold="true" box="[495,509,1567,1584]" pageId="10" pageNumber="391">G</emphasis>
|
||
, Pi 7523;
|
||
<emphasis id="B9276660FF8EE022FD97AF509A19BA06" bold="true" box="[594,608,1567,1584]" pageId="10" pageNumber="391">H</emphasis>
|
||
, Pi 7522;
|
||
<emphasis id="B9276660FF8EE022FD70AF509AC2BA06" bold="true" box="[693,699,1567,1584]" pageId="10" pageNumber="391">I</emphasis>
|
||
, Pi 7524;
|
||
<emphasis id="B9276660FF8EE022FCCAAF509B60BA06" bold="true" box="[783,793,1567,1584]" pageId="10" pageNumber="391">J</emphasis>
|
||
, Pi 7525;
|
||
<emphasis id="B9276660FF8EE022FCABAF509B02BA06" bold="true" box="[878,891,1567,1584]" pageId="10" pageNumber="391">K</emphasis>
|
||
, holotype Pi 6803;
|
||
<emphasis id="B9276660FF8EE022FBDAAF509C53BA06" bold="true" box="[1055,1066,1567,1584]" pageId="10" pageNumber="391">L</emphasis>
|
||
, Pi 6805;
|
||
<emphasis id="B9276660FF8EE022FBBAAF509CE9BA06" bold="true" box="[1151,1168,1567,1584]" pageId="10" pageNumber="391">M</emphasis>
|
||
, Pi 6801;
|
||
<emphasis id="B9276660FF8EE022FF52AF7B98DCBA73" bold="true" box="[151,165,1588,1605]" pageId="10" pageNumber="391">N</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF8EE022FF6AAF7B98C7BA73" bold="true" box="[175,190,1588,1605]" pageId="10" pageNumber="391">O</emphasis>
|
||
, Pi 7526;
|
||
<emphasis id="B9276660FF8EE022FED4AF7B9964BA73" bold="true" box="[273,285,1588,1605]" pageId="10" pageNumber="391">P</emphasis>
|
||
, Pi 6802;
|
||
<emphasis id="B9276660FF8EE022FEB4AF7B99F9BA73" bold="true" box="[369,384,1588,1605]" pageId="10" pageNumber="391">Q</emphasis>
|
||
, Pi 7527;
|
||
<emphasis id="B9276660FF8EE022FE16AF7B9999BA73" bold="true" box="[467,480,1588,1605]" pageId="10" pageNumber="391">R</emphasis>
|
||
, Pi 7529;
|
||
<emphasis id="B9276660FF8EE022FDF6AF7B9A46BA73" bold="true" box="[563,575,1588,1605]" pageId="10" pageNumber="391">S</emphasis>
|
||
, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian;
|
||
<emphasis id="B9276660FF8EE022FDB1AF069AEBBA6C" bold="true" box="[628,658,1609,1626]" pageId="10" pageNumber="391">T -V</emphasis>
|
||
,
|
||
<taxonomicName id="4C53C1F1FF8EE022FD58AF069A90BA6C" authorityName="Agassiz" authorityYear="1839" box="[669,745,1609,1626]" class="Pteraspidomorphi" family="Loganelliidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="10" pageNumber="391" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9276660FF8EE022FD58AF069A90BA6C" box="[669,745,1609,1626]" italics="true" pageId="10" pageNumber="391">Thelodus</emphasis>
|
||
</taxonomicName>
|
||
? sp., scales in crown view;
|
||
<emphasis id="B9276660FF8EE022FC12AF069B9BBA6C" bold="true" box="[983,994,1609,1626]" pageId="10" pageNumber="391">T</emphasis>
|
||
, Pi 7516;
|
||
<emphasis id="B9276660FF8EE022FBFDAF069C3FBA6C" bold="true" box="[1080,1094,1609,1626]" pageId="10" pageNumber="391">U</emphasis>
|
||
, LIG 35 908;
|
||
<emphasis id="B9276660FF8EE022FB7DAF069CBDBA6C" bold="true" box="[1208,1220,1609,1626]" pageId="10" pageNumber="391">V</emphasis>
|
||
, Pi 6814;
|
||
<emphasis id="B9276660FF8EE022FF08AF1098A1BA46" bold="true" box="[205,216,1631,1648]" pageId="10" pageNumber="391">T</emphasis>
|
||
,
|
||
<emphasis id="B9276660FF8EE022FF21AF109889BA46" bold="true" box="[228,240,1631,1648]" pageId="10" pageNumber="391">V</emphasis>
|
||
, sample MF 46-23;
|
||
<emphasis id="B9276660FF8EE022FE65AF1099D7BA46" bold="true" box="[416,430,1631,1648]" pageId="10" pageNumber="391">U</emphasis>
|
||
, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BECBA72FF8FE023FF52A9E499EEBCC5" blockId="11.[151,684,170,816]" pageId="11" pageNumber="392">OCCURRENCE. — Sample MF 157-7, Spokojnaya River, upper part of Samojlovich Formation, Wenlock, Lower Silurian.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349E9F9FF8FE023FF52A849992FBF06" pageId="11" pageNumber="392" type="diagnosis">
|
||
<paragraph id="8BECBA72FF8FE023FF52A849992FBF06" blockId="11.[151,684,170,816]" pageId="11" pageNumber="392">DIAGNOSIS. — Scales of medium size. Head scales rounded to oval with crenulated margins. Neck is rather distinct; base deep with vertical rootlets. Transitional scales are oval to rhomboidal with crenulation in the antero-lateral margins and one posterior crown apex. Diagnostic trunk scales bear a central part with a shallow groove, a pair of lateral ribs directed towards the posterior central crown apex, and two-three spines beneath the lateral ribs, one single spine situated below the central crown apex. Each more laterally positioned crown portion (ribs and spines) are downstepped. Anterior horizontal process of the base, sometimes very long, is present in all trunk scales. Neck is low but distinct. Pulp depression large. Opening of the main, medial, pulp canal is situated in the posterior portion of the base. Two to four pairs of lateral pulp canals may be present. Dentine canals start from the pulp depression or pulp canals; they are similar to the orthodentine canals. Fine dentine tubules occur in the crown but can also be found in the base. The crown surface is from a layer of enameloid.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349E9F9FF8FE024FF52AA259AE3BD15" lastPageId="12" lastPageNumber="393" pageId="11" pageNumber="392" type="description">
|
||
<paragraph id="8BECBA72FF8FE023FF52AA259951BFB4" blockId="11.[151,684,874,1667]" box="[151,296,874,900]" pageId="11" pageNumber="392">DESCRIPTION</paragraph>
|
||
<paragraph id="8BECBA72FF8FE023FF52AAC49968BF93" blockId="11.[151,684,874,1667]" box="[151,273,907,933]" pageId="11" pageNumber="392">
|
||
<emphasis id="B9276660FF8FE023FF52AAC49968BF93" box="[151,273,907,933]" italics="true" pageId="11" pageNumber="392">Morphology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF8FE023FF52AAE49B58BEB5" blockId="11.[151,684,874,1667]" lastBlockId="11.[714,1247,171,643]" pageId="11" pageNumber="392">
|
||
Only one sample (MF 157-7) yields the scales of this new species. In the set, comparatively large head and transitional scales are present in contrast to the small trunk and fin scales. The length of scale crowns range from 0.2 up to
|
||
<quantity id="4CAB1797FF8FE023FD8CAD649ADEB875" box="[585,679,1067,1091]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="392" unit="mm" value="0.5">0.5 mm</quantity>
|
||
. Elongated spiny trunk scales have an anterior horizontal basal spur up to
|
||
<quantity id="4CAB1797FF8FE023FE21AD249A42B8B5" box="[484,571,1131,1155]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="392" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long and another
|
||
<quantity id="4CAB1797FF8FE023FF31ADC49931B895" box="[244,328,1163,1187]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="392" unit="mm" value="0.5">0.5 mm</quantity>
|
||
makes the crown. The smallest scale is only
|
||
<quantity id="4CAB1797FF8FE023FEF7ADE499F3B8F5" box="[306,394,1195,1219]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="392" unit="mm" value="0.1">0.1 mm</quantity>
|
||
wide. Large head scales have a vertical base that is
|
||
<quantity id="4CAB1797FF8FE023FE7BAD849A68B8D5" box="[446,529,1227,1251]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="392" unit="mm" value="0.5">0.5 mm</quantity>
|
||
high at maximum. The
|
||
<typeStatus id="54E804D0FF8FE023FEDCADA499FAB935" box="[281,387,1259,1283]" pageId="11" pageNumber="392" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="1368A6F7FF8FE023FE50ADA49989B935" box="[405,496,1259,1283]" captionStart="FIG" captionStartId="10.[151,162,1524,1541]" captionTargetBox="[203,1186,173,1467]" captionTargetId="figure-21@10.[203,1186,173,1467]" captionTargetPageId="10" captionText="FIG. 7. — A-S, Shielia multispinata n. sp.; A, head scale; B-D, F, transitional scales; E, G-P, trunk scales; Q-S, fin scales; A, B, scales in lateral view; C-N, P, Q, S, scales in crown view; O, a scale in oblique view; R, a scale in base view; A, Pi 7517; B, Pi 7519; C, Pi 7518; D, Pi 7520; E, Pi 7521; F, Pi 6804; G, Pi 7523; H, Pi 7522; I, Pi 7524; J, Pi 7525; K, holotype Pi 6803; L, Pi 6805; M, Pi 6801; N, O, Pi 7526; P, Pi 6802; Q, Pi 7527; R, Pi 7529; S, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; T-V, Thelodus? sp., scales in crown view; T, Pi 7516; U, LIG 35 908; V, Pi 6814; T, V, sample MF 46-23; U, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="11" pageNumber="392">Fig. 7K</figureCitation>
|
||
) has a
|
||
<quantity id="4CAB1797FF8FE023FD83ADA49AD2B935" box="[582,683,1259,1283]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="392" unit="mm" value="0.25">0.25 mm</quantity>
|
||
long horizontal spur and
|
||
<quantity id="4CAB1797FF8FE023FE03AC449A64B915" box="[454,541,1291,1315]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="392" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long crown with width of
|
||
<quantity id="4CAB1797FF8FE023FE85AC6499DFB975" box="[320,422,1323,1347]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="392" unit="mm" value="0.25">0.25 mm</quantity>
|
||
. Head scales (
|
||
<figureCitation id="1368A6F7FF8FE023FD8EAC649ADAB975" box="[587,675,1323,1347]" captionStart="FIG" captionStartId="10.[151,162,1524,1541]" captionTargetBox="[203,1186,173,1467]" captionTargetId="figure-21@10.[203,1186,173,1467]" captionTargetPageId="10" captionText="FIG. 7. — A-S, Shielia multispinata n. sp.; A, head scale; B-D, F, transitional scales; E, G-P, trunk scales; Q-S, fin scales; A, B, scales in lateral view; C-N, P, Q, S, scales in crown view; O, a scale in oblique view; R, a scale in base view; A, Pi 7517; B, Pi 7519; C, Pi 7518; D, Pi 7520; E, Pi 7521; F, Pi 6804; G, Pi 7523; H, Pi 7522; I, Pi 7524; J, Pi 7525; K, holotype Pi 6803; L, Pi 6805; M, Pi 6801; N, O, Pi 7526; P, Pi 6802; Q, Pi 7527; R, Pi 7529; S, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; T-V, Thelodus? sp., scales in crown view; T, Pi 7516; U, LIG 35 908; V, Pi 6814; T, V, sample MF 46-23; U, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="11" pageNumber="392">Fig. 7A</figureCitation>
|
||
) have a round or oval crown and base configuration. The crown surface is flat and smooth with crenulated margins, with the neck in the shape of a shallow groove. The pulp cavity is placed in the middle of the scale and surrounded by vertical rootlets of the base. Transitional scales (
|
||
<figureCitation id="1368A6F7FF8FE023FF65AF44989FBA15" box="[160,230,1547,1571]" captionStart="FIG" captionStartId="10.[151,162,1524,1541]" captionTargetBox="[203,1186,173,1467]" captionTargetId="figure-21@10.[203,1186,173,1467]" captionTargetPageId="10" captionText="FIG. 7. — A-S, Shielia multispinata n. sp.; A, head scale; B-D, F, transitional scales; E, G-P, trunk scales; Q-S, fin scales; A, B, scales in lateral view; C-N, P, Q, S, scales in crown view; O, a scale in oblique view; R, a scale in base view; A, Pi 7517; B, Pi 7519; C, Pi 7518; D, Pi 7520; E, Pi 7521; F, Pi 6804; G, Pi 7523; H, Pi 7522; I, Pi 7524; J, Pi 7525; K, holotype Pi 6803; L, Pi 6805; M, Pi 6801; N, O, Pi 7526; P, Pi 6802; Q, Pi 7527; R, Pi 7529; S, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; T-V, Thelodus? sp., scales in crown view; T, Pi 7516; U, LIG 35 908; V, Pi 6814; T, V, sample MF 46-23; U, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="11" pageNumber="392">Fig. 7</figureCitation>
|
||
B-D, F) are oval or rhomboidal with crenulation or two notches antero-laterally of the crown. The base is becoming more horizontal. Trunk scales (
|
||
<figureCitation id="1368A6F7FF8FE023FEA7AF2499C1BAB5" box="[354,440,1643,1667]" captionStart="FIG" captionStartId="10.[151,162,1524,1541]" captionTargetBox="[203,1186,173,1467]" captionTargetId="figure-21@10.[203,1186,173,1467]" captionTargetPageId="10" captionText="FIG. 7. — A-S, Shielia multispinata n. sp.; A, head scale; B-D, F, transitional scales; E, G-P, trunk scales; Q-S, fin scales; A, B, scales in lateral view; C-N, P, Q, S, scales in crown view; O, a scale in oblique view; R, a scale in base view; A, Pi 7517; B, Pi 7519; C, Pi 7518; D, Pi 7520; E, Pi 7521; F, Pi 6804; G, Pi 7523; H, Pi 7522; I, Pi 7524; J, Pi 7525; K, holotype Pi 6803; L, Pi 6805; M, Pi 6801; N, O, Pi 7526; P, Pi 6802; Q, Pi 7527; R, Pi 7529; S, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; T-V, Thelodus? sp., scales in crown view; T, Pi 7516; U, LIG 35 908; V, Pi 6814; T, V, sample MF 46-23; U, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="11" pageNumber="392">Fig. 7E</figureCitation>
|
||
, G-P) are rather flattened. A longitudinal shallow groove is placed on the medial elongated portion of the crown surface. On both sides of it a slightly downstepped rib starts from the base and is directed towards the posterior medial crown apex; lateral ribs do not join with the latter and a split occurs between them. The second pair of downstepped ribs can be relatively long or consist of up to three spines. The lower side of the posterior crown portion exhibits five ridges. The base has an anterior spur-like process. The pulp depression is large, and openings of pulp canals have a posterior position. Fin scales (
|
||
<figureCitation id="1368A6F7FF8FE023FBEDAB649C12BE75" box="[1064,1131,555,579]" captionStart="FIG" captionStartId="10.[151,162,1524,1541]" captionTargetBox="[203,1186,173,1467]" captionTargetId="figure-21@10.[203,1186,173,1467]" captionTargetPageId="10" captionText="FIG. 7. — A-S, Shielia multispinata n. sp.; A, head scale; B-D, F, transitional scales; E, G-P, trunk scales; Q-S, fin scales; A, B, scales in lateral view; C-N, P, Q, S, scales in crown view; O, a scale in oblique view; R, a scale in base view; A, Pi 7517; B, Pi 7519; C, Pi 7518; D, Pi 7520; E, Pi 7521; F, Pi 6804; G, Pi 7523; H, Pi 7522; I, Pi 7524; J, Pi 7525; K, holotype Pi 6803; L, Pi 6805; M, Pi 6801; N, O, Pi 7526; P, Pi 6802; Q, Pi 7527; R, Pi 7529; S, Pi 7528; sample MF 157-7, Spokojnaya River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian; T-V, Thelodus? sp., scales in crown view; T, Pi 7516; U, LIG 35 908; V, Pi 6814; T, V, sample MF 46-23; U, sample 32a-208, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: A-I, K-M, P-V, 0.1 mm; J, N, O, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5374759" httpUri="https://zenodo.org/record/5374759/files/figure.png" pageId="11" pageNumber="392">Fig. 7</figureCitation>
|
||
Q-S) have the same shape as trunk scales but are much smaller.
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF8FE023FD0FABE49BD0BEF3" blockId="11.[714,1247,682,1251]" box="[714,937,682,709]" pageId="11" pageNumber="392">
|
||
<emphasis id="B9276660FF8FE023FD0FABE49BD0BEF3" box="[714,937,682,709]" italics="true" pageId="11" pageNumber="392">
|
||
Histology (
|
||
<figureCitation id="1368A6F7FF8FE023FCFDABE59B0FBEF3" box="[824,886,682,709]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[208,1202,174,1484]" captionTargetId="figure-25@9.[206,1208,194,1502]" captionTargetPageId="9" captionText="FIG. 6. — Microstructure of the scales; A, B, Loganellia matura n. sp.; C-F, Shielia multispinata n. sp.; G, Paralogania consimilis n. sp.; A, Pi 7624; B, Pi 7625; C, Pi 7626; D, Pi 7627; E, Pi 7628; F, Pi 7629; G, Pi 7630; A, B, sample MF 3-8, Srednii Island, Golomyannyj Formation, middle part of Llandovery, Lower Silurian; C-F, sample MF 157-7, Spokojnaya River, upper part of Samojlovich Formation, Wenlock, Lower Silurian; G, sample MF 46-23, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; bc, canal in the base; cr, crown; dc, dentine canal; dt, dentine tubules; fpc, fan-shaped pulp canals; lac, lacunae of dentine canals; mpc, main pulp canal; n, neck; pc, pulp canal; pco, pulp canal opening; sp, spine. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374757" httpUri="https://zenodo.org/record/5374757/files/figure.png" pageId="11" pageNumber="392">Fig. 6</figureCitation>
|
||
C-F)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF8FE023FD0FAB849CF5B8D5" blockId="11.[714,1247,682,1251]" pageId="11" pageNumber="392">
|
||
The pulp cavity of the young head and transitional scales can be rather deep and wide, but is narrower in adult scales. The pulp depression of trunk scales is large; from the posterior part of it three to nine forked or fan-shaped pulp canals (a medial and four pairs of lateral) enter into three to nine posterior spines and go far posteriorly (
|
||
<figureCitation id="1368A6F7FF8FE023FD17AAE49B28BFF5" box="[722,849,939,963]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[208,1202,174,1484]" captionTargetId="figure-25@9.[206,1208,194,1502]" captionTargetPageId="9" captionText="FIG. 6. — Microstructure of the scales; A, B, Loganellia matura n. sp.; C-F, Shielia multispinata n. sp.; G, Paralogania consimilis n. sp.; A, Pi 7624; B, Pi 7625; C, Pi 7626; D, Pi 7627; E, Pi 7628; F, Pi 7629; G, Pi 7630; A, B, sample MF 3-8, Srednii Island, Golomyannyj Formation, middle part of Llandovery, Lower Silurian; C-F, sample MF 157-7, Spokojnaya River, upper part of Samojlovich Formation, Wenlock, Lower Silurian; G, sample MF 46-23, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; bc, canal in the base; cr, crown; dc, dentine canal; dt, dentine tubules; fpc, fan-shaped pulp canals; lac, lacunae of dentine canals; mpc, main pulp canal; n, neck; pc, pulp canal; pco, pulp canal opening; sp, spine. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374757" httpUri="https://zenodo.org/record/5374757/files/figure.png" pageId="11" pageNumber="392">Fig. 6C, D</figureCitation>
|
||
). The openings of pulp canals can form a longitudinal row in the base (
|
||
<figureCitation id="1368A6F7FF8FE023FBBEAA849CABBFD5" box="[1147,1234,971,995]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[208,1202,174,1484]" captionTargetId="figure-25@9.[206,1208,194,1502]" captionTargetPageId="9" captionText="FIG. 6. — Microstructure of the scales; A, B, Loganellia matura n. sp.; C-F, Shielia multispinata n. sp.; G, Paralogania consimilis n. sp.; A, Pi 7624; B, Pi 7625; C, Pi 7626; D, Pi 7627; E, Pi 7628; F, Pi 7629; G, Pi 7630; A, B, sample MF 3-8, Srednii Island, Golomyannyj Formation, middle part of Llandovery, Lower Silurian; C-F, sample MF 157-7, Spokojnaya River, upper part of Samojlovich Formation, Wenlock, Lower Silurian; G, sample MF 46-23, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; bc, canal in the base; cr, crown; dc, dentine canal; dt, dentine tubules; fpc, fan-shaped pulp canals; lac, lacunae of dentine canals; mpc, main pulp canal; n, neck; pc, pulp canal; pco, pulp canal opening; sp, spine. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374757" httpUri="https://zenodo.org/record/5374757/files/figure.png" pageId="11" pageNumber="392">Fig. 6F</figureCitation>
|
||
). Fine, nearly parallel dentine canals go out from the pulp canals or from the base (
|
||
<figureCitation id="1368A6F7FF8FE023FBB4AD449CA8B815" box="[1137,1233,1035,1059]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[208,1202,174,1484]" captionTargetId="figure-25@9.[206,1208,194,1502]" captionTargetPageId="9" captionText="FIG. 6. — Microstructure of the scales; A, B, Loganellia matura n. sp.; C-F, Shielia multispinata n. sp.; G, Paralogania consimilis n. sp.; A, Pi 7624; B, Pi 7625; C, Pi 7626; D, Pi 7627; E, Pi 7628; F, Pi 7629; G, Pi 7630; A, B, sample MF 3-8, Srednii Island, Golomyannyj Formation, middle part of Llandovery, Lower Silurian; C-F, sample MF 157-7, Spokojnaya River, upper part of Samojlovich Formation, Wenlock, Lower Silurian; G, sample MF 46-23, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine “pegs” in the base; b, base; bc, canal in the base; cr, crown; dc, dentine canal; dt, dentine tubules; fpc, fan-shaped pulp canals; lac, lacunae of dentine canals; mpc, main pulp canal; n, neck; pc, pulp canal; pco, pulp canal opening; sp, spine. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5374757" httpUri="https://zenodo.org/record/5374757/files/figure.png" pageId="11" pageNumber="392">Fig. 6E</figureCitation>
|
||
). Dentine canals are relatively straight and numerous. They resemble more the canals in the orthodentine than in the mesodentine. Dentine tubules straight. In the base, fine tubules of Sharpey’s fibres, perpendicular to the base surface, and aspidine “pegs” are distributed.
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF8FE023FD0FAC449B20B914" blockId="11.[714,1247,1291,1667]" box="[714,857,1291,1317]" pageId="11" pageNumber="392">COMPARISON</paragraph>
|
||
<paragraph id="8BECBA72FF8FE024FD0FAC649AFCBCD5" blockId="11.[714,1247,1291,1667]" lastBlockId="12.[151,684,171,227]" lastPageId="12" lastPageNumber="393" pageId="11" pageNumber="392">
|
||
Head and transitional scales of
|
||
<taxonomicName id="4C53C1F1FF8FE023FBFAAC649B5DB955" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="multispinata" status="sp. nov.">
|
||
<emphasis id="B9276660FF8FE023FBFAAC649B5DB955" italics="true" pageId="11" pageNumber="392">Shielia multispinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214DB1BFF8FE023FCEAAC039B0DB955" box="[815,884,1356,1379]" pageId="11" pageNumber="392" rank="species">n. sp.</taxonomicNameLabel>
|
||
differ from those in
|
||
<taxonomicName id="4C53C1F1FF8FE023FB4FAC039B09B9B5" authority="(Stetson, 1931)" baseAuthorityName="Stetson" baseAuthorityYear="1931" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="taiti">
|
||
<emphasis id="B9276660FF8FE023FB4FAC039CA6B955" box="[1162,1247,1356,1379]" italics="true" pageId="11" pageNumber="392">S. taiti</emphasis>
|
||
(Stetson, 1931)
|
||
</taxonomicName>
|
||
in being larger and more crenulated while in
|
||
<taxonomicName id="4C53C1F1FF8FE023FCA1ACC39BD2B995" baseAuthorityName="Stetson" baseAuthorityYear="1931" box="[868,939,1420,1443]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="taiti">
|
||
<emphasis id="B9276660FF8FE023FCA1ACC39BD2B995" box="[868,939,1420,1443]" italics="true" pageId="11" pageNumber="392">S. taiti</emphasis>
|
||
</taxonomicName>
|
||
they are smaller (
|
||
<taxonomicName id="4C53C1F1FF8FE023FBABACC39CCCB995" baseAuthorityName="Stetson" baseAuthorityYear="1931" box="[1134,1205,1420,1443]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="taiti">
|
||
<emphasis id="B9276660FF8FE023FBABACC39CCCB995" box="[1134,1205,1420,1443]" italics="true" pageId="11" pageNumber="392">S. taiti</emphasis>
|
||
</taxonomicName>
|
||
has scales up to
|
||
<quantity id="4CAB1797FF8FE023FC98ACE49BCDB9F5" box="[861,948,1451,1475]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="392" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long), simpler anteriorly and with two wing-like structures laterally. The new species has trunk scales with lateral ribs directed towards the posterior central crown apex; in
|
||
<taxonomicName id="4C53C1F1FF8FE023FCE2AF639B16BA75" baseAuthorityName="Stetson" baseAuthorityYear="1931" box="[807,879,1580,1603]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="taiti">
|
||
<emphasis id="B9276660FF8FE023FCE2AF639B16BA75" box="[807,879,1580,1603]" italics="true" pageId="11" pageNumber="392">S. taiti</emphasis>
|
||
</taxonomicName>
|
||
they start from the neck, tend to run more parallel to the central portion and end with free spines posteriorly.
|
||
<taxonomicName id="4C53C1F1FF8FE023FBD1AF239C27BAB5" baseAuthorityName="Stetson" baseAuthorityYear="1931" box="[1044,1118,1644,1667]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="taiti">
|
||
<emphasis id="B9276660FF8FE023FBD1AF239C27BAB5" box="[1044,1118,1644,1667]" italics="true" pageId="11" pageNumber="392">S. taiti</emphasis>
|
||
</taxonomicName>
|
||
has deeper grooves between the ribs. In
|
||
<taxonomicName id="4C53C1F1FF88E024FE3CA9E39AD5BCF5" authority="Märss & Karatajūtē-Talimaa, 2002" authorityName="Märss & Karatajūtē-Talimaa" authorityYear="2002" box="[505,684,171,195]" class="Pteraspidomorphi" family="Loganelliidae" genus="Shielia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="12" pageNumber="393" phylum="Chordata" rank="species" species="multispinata" status="sp. nov.">
|
||
<emphasis id="B9276660FF88E024FE3CA9E39AD5BCF5" box="[505,684,171,195]" italics="true" pageId="12" pageNumber="393">S. multispinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214DB1BFF88E024FF52A98398ABBCD5" box="[151,210,204,227]" pageId="12" pageNumber="393" rank="species">n. sp.</taxonomicNameLabel>
|
||
the spines start beneath the lateral ribs.
|
||
</paragraph>
|
||
<paragraph id="8BECBA72FF88E024FF6CA8449AE3BD15" blockId="12.[169,666,267,291]" box="[169,666,267,291]" pageId="12" pageNumber="393">
|
||
Genus
|
||
<taxonomicName id="4C53C1F1FF88E024FF3DA8449904BD15" authorityName="Karatajute-Talimaa" authorityYear="1997" box="[248,381,267,291]" class="Pteraspidomorphi" family="Loganelliidae" genus="Paralogania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="12" pageNumber="393" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9276660FF88E024FF3DA8449904BD15" box="[248,381,267,291]" italics="true" pageId="12" pageNumber="393">Paralogania</emphasis>
|
||
</taxonomicName>
|
||
Karatajūtē- Talimaa, 1997
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349E9F9FF88E024FF52A80A9AD8BD6C" box="[151,673,324,346]" pageId="12" pageNumber="393" type="materials_examined">
|
||
<paragraph id="8BECBA72FF88E024FF52A80A9AD8BD6C" blockId="12.[151,673,324,346]" box="[151,673,324,346]" pageId="12" pageNumber="393">
|
||
<collectionCode id="ED4222B7FF88E024FF52A80A9974BD6C" box="[151,269,325,346]" pageId="12" pageNumber="393">TYPE SPECIES</collectionCode>
|
||
. —
|
||
<taxonomicName id="4C53C1F1FF88E024FEFDA80A9AE5BD6C" authority="(Gross, 1967)" baseAuthorityName="Gross" baseAuthorityYear="1967" box="[312,668,324,346]" class="Pteraspidomorphi" family="Loganelliidae" genus="Paralogania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phlebolepidiformes" pageId="12" pageNumber="393" phylum="Chordata" rank="species" species="kummerowi">
|
||
<emphasis id="B9276660FF88E024FEFDA80A9A62BD6C" box="[312,539,325,346]" italics="true" pageId="12" pageNumber="393">Paralogania kummerowi</emphasis>
|
||
(Gross, 1967)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |