254 lines
35 KiB
XML
254 lines
35 KiB
XML
<document id="1B94BA59FA0840D08E9F5C67A36074A9" ID-CLB-Dataset="5440" ID-DOI="10.11865/zs.202006" ID-GBIF-Dataset="b121937a-8d53-4315-a3aa-18cd2d44dac8" ID-ISSN="2095-6827" ID-Zenodo-Dep="4617188" ID-ZooBank="DFC21F50-02A5-48B8-B4EC-18447FA75CDB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616070693767" checkinUser="carolina" docAuthor="Men, Qiulei" docDate="2020" docId="03F787C4A833FFF0FF5ACC6DFEC810B8" docLanguage="en" docName="ZoolSyst.45.1.59-68.pdf" docOrigin="Zoological Systematics 45 (1)" docSource="http://zoobank.org/21f2cc71-4be0-4ef9-8762-0e0348285e1b" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Tipula (Sivatipula) pianmaensis Men 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="63" masterDocId="FFCEFFBCA832FFF4FFD2CA02FFF2123A" masterDocTitle="Three new species of the subgenus Tipula (Sivatipula) from Yunnan and Tibet, China (Diptera: Tipulidae), with an updated key for Chinese species" masterLastPageNumber="68" masterPageNumber="59" pageNumber="60" updateTime="1698934182474" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E7462B96A0A8DC2B7AC0E27DEB1A650A">Three new species of the subgenus Tipula (Sivatipula) from Yunnan and Tibet, China (Diptera: Tipulidae), with an updated key for Chinese species</mods:title>
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<mods:namePart id="24ACD81FB4E3140DB7BFE99E540C2281">Men, Qiulei</mods:namePart>
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<mods:typeOfResource id="D1DA57275B96DC4B8F6C77E2447865AB">text</mods:typeOfResource>
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<mods:title id="F19B0BA84FA0F7876376D26621E3EABC">Zoological Systematics</mods:title>
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<mods:part id="901B237FAB3850399EFCA19978E30043">
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<mods:date id="EC7EBA54B3BD739B732480B2892CCFD5">2020</mods:date>
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<mods:number id="83C8390243551EC2A5EEAA7AC8304C43">2020-01-31</mods:number>
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<mods:classification id="024477E167E0E2862BF06EAB93A711E5">journal article</mods:classification>
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<mods:identifier id="EED513CD8D363BA3686465131D480BA2" type="CLB-Dataset">5440</mods:identifier>
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<mods:identifier id="E688685620B544C988B24F18ED3B6913" type="DOI">10.11865/zs.202006</mods:identifier>
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<mods:identifier id="587CA85A358EBF0C9D85B499379DFE47" type="ISSN">2095-6827</mods:identifier>
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<treatment id="03F787C4A833FFF0FF5ACC6DFEC810B8" ID-DOI="http://doi.org/10.5281/zenodo.5104883" ID-GBIF-Taxon="183543477" ID-Zenodo-Dep="5104883" LSID="urn:lsid:plazi:treatment:03F787C4A833FFF0FF5ACC6DFEC810B8" httpUri="http://treatment.plazi.org/id/03F787C4A833FFF0FF5ACC6DFEC810B8" lastPageId="4" lastPageNumber="63" pageId="1" pageNumber="60">
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<subSubSection id="C3446559A833FFF5FF5ACC6DFD2714BD" box="[136,725,1647,1671]" pageId="1" pageNumber="60" type="nomenclature">
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<paragraph id="8BE136D2A833FFF5FF5ACC6DFD2714BD" blockId="1.[136,725,1647,1671]" box="[136,725,1647,1671]" pageId="1" pageNumber="60">
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<heading id="D0A981BEA833FFF5FF5ACC6DFD2714BD" box="[136,725,1647,1671]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="60" reason="2">
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<emphasis id="B92AEAC0A833FFF5FF5ACC6DFDB614BD" bold="true" box="[136,580,1647,1671]" pageId="1" pageNumber="60">
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<taxonomicName id="4C5E4D51A833FFF5FF5ACC6DFE1A14BD" authority="Men, 2020" authorityName="Men" authorityYear="2020" box="[136,488,1647,1671]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="pianmaensis" status="sp. nov." subGenus="Sivatipula">
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<emphasis id="B92AEAC0A833FFF5FF5ACC6DFF2214BD" bold="true" box="[136,208,1647,1671]" italics="true" pageId="1" pageNumber="60">Tipula</emphasis>
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(
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<emphasis id="B92AEAC0A833FFF5FF0DCC6DFEBC14BD" bold="true" box="[223,334,1647,1671]" italics="true" pageId="1" pageNumber="60">Sivatipula</emphasis>
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)
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<emphasis id="B92AEAC0A833FFF5FE8CCC6DFE1A14BD" bold="true" box="[350,488,1647,1671]" italics="true" pageId="1" pageNumber="60">pianmaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21957BBA833FFF5FE3DCC6DFDB614BD" box="[495,580,1647,1671]" pageId="1" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="13652A57A833FFF5FDB3CC6DFD3F14BD" box="[609,717,1647,1671]" captionStart-0="Figures 1–6" captionStart-1="Figures 7–15" captionStartId-0="2.[155,227,2009,2031]" captionStartId-1="3.[155,227,1986,2008]" captionTargetBox-0="[189,1399,571,1946]" captionTargetId-0="figure-191@2.[189,1399,571,1969]" captionTargetPageId-0="2" captionText-0="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." captionText-1="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4617190" httpUri-0="https://zenodo.org/record/4617190/files/figure.png" pageId="1" pageNumber="60">Figs 1–15</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3446559A833FFF0FF6FCCA5FEC810B8" lastPageId="4" lastPageNumber="63" pageId="1" pageNumber="60" type="diagnosis">
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<paragraph id="8BE136D2A833FFF5FF6FCCA5FB651573" blockId="1.[136,1458,1703,2073]" pageId="1" pageNumber="60">Diagnosis. Body gray in general; thorax gray with three brownish-yellow stripes on prescutum, the median one divided by a broad gray vitta; antenna longer than body length; wing suffused with brownish-yellow, stigma light brown; abdomen brownish-yellow with brown lateral stripes, segments six and seven wholly black, hypopygium yellow, distinctly expanded; tergite nine with a pair of horn-shaped processes at posterior margin, ventral extension of tergite nine densely covered with small black teeth; inner gonostylus terminated into a long black beak; outer gonostylus Y-shaped.</paragraph>
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<paragraph id="8BE136D2A833FFF5FF6FCD56FC1015B5" blockId="1.[136,1458,1703,2073]" pageId="1" pageNumber="60">
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<materialsCitation id="3B363C8FA833FFF5FF6FCD56FC2C15B5" ID-GBIF-Occurrence="3322027303" collectingDate="2019-06-19" collectorName="Qiulei Men." country="China" county="Material" latitude="26.0" location="Pianma" longLatPrecision="1243" longitude="98.61667" municipality="Lushui" pageId="1" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
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<collectingCounty id="62804E5EA833FFF5FF6FCD56FEE51556" box="[189,279,1876,1900]" pageId="1" pageNumber="60">Material</collectingCounty>
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examined.
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<typeStatus id="54E58870A833FFF5FE43CD56FE061556" box="[401,500,1876,1900]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
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male,
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<collectingCountry id="F3497642A833FFF5FDE8CD56FD8F1556" box="[570,637,1876,1900]" name="China" pageId="1" pageNumber="60">China</collectingCountry>
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,
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<collectingRegion id="499AF830A833FFF5FD57CD56FD281551" box="[645,730,1876,1899]" country="China" name="Yunnan" pageId="1" pageNumber="60">Yunnan</collectingRegion>
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,
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<collectingMunicipality id="6B85ACA8A833FFF5FD30CD56FCDD1556" box="[738,815,1876,1900]" pageId="1" pageNumber="60">Lushui</collectingMunicipality>
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,
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<location id="8E816009A833FFF5FCEACD56FC791556" LSID="urn:lsid:plazi:treatment:03F787C4A833FFF0FF5ACC6DFEC810B8:8E816009A833FFF5FCEACD56FC791556" box="[824,907,1876,1900]" country="China" county="Material" latitude="26.0" longLatPrecision="1243" longitude="98.61667" municipality="Lushui" name="Pianma" pageId="1" pageNumber="60" stateProvince="Yunnan">Pianma</location>
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,
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<location id="8E816009A833FFF5FC46CD56FAB91551" LSID="urn:lsid:plazi:treatment:03F787C4A833FFF0FF5ACC6DFEC810B8:8E816009A833FFF5FC46CD56FAB91551" box="[916,1355,1876,1900]" country="China" county="Material" latitude="26.0" longLatPrecision="1243" longitude="98.61667" municipality="Lushui" name="Gaoligongshan National Natural Reserve" pageId="1" pageNumber="60" stateProvince="Yunnan">Gaoligongshan National Natural Reserve</location>
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,
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<geoCoordinate id="EE6A5015A833FFF5FA86CD56FA5C1556" box="[1364,1454,1876,1900]" degrees="26" direction="north" minutes="00" orientation="latitude" pageId="1" pageNumber="60" precision="925" value="26.0">26°00'N</geoCoordinate>
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,
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<geoCoordinate id="EE6A5015A833FFF5FF5ACD75FF2D15B5" box="[136,223,1911,1935]" degrees="98" direction="east" minutes="37" orientation="longitude" pageId="1" pageNumber="60" precision="925" value="98.61667">98°37'E</geoCoordinate>
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,
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<date id="FFE01012A833FFF5FF39CD75FE8215B5" box="[235,368,1911,1935]" pageId="1" pageNumber="60" value="2019-06-19">
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<collectingDate id="EFA4E9FAA833FFF5FF39CD75FE8215B5" box="[235,368,1911,1935]" pageId="1" pageNumber="60" value="2019-06-19">19 Jun 2019</collectingDate>
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</date>
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, leg.
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<collectorName id="26AB5304A833FFF5FE7ACD75FDD815B4" box="[424,554,1911,1935]" pageId="1" pageNumber="60">Qiulei Men.</collectorName>
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<typeStatus id="54E58870A833FFF5FDE2CD75FD6315B5" box="[560,657,1911,1935]" pageId="1" pageNumber="60" type="paratype">Paratype</typeStatus>
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.
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<specimenCount id="9D58FD5BA833FFF5FD48CD75FD1715B5" box="[666,741,1911,1935]" pageId="1" pageNumber="60" type="male">1 male</specimenCount>
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, same data as holotype
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BE136D2A833FFF5FF6FCD9BFF1915E9" blockId="1.[136,1458,1703,2073]" pageId="1" pageNumber="60">
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Description. Adult length (male,
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<emphasis id="B92AEAC0A833FFF5FDCBCD98FDD4158B" box="[537,550,1946,1969]" italics="true" pageId="1" pageNumber="60">n</emphasis>
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=2): body
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<quantity id="4CA69B37A833FFF5FD4BCD9BFCDD158B" box="[665,815,1945,1969]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" metricValueMax="1.47" metricValueMin="1.45" pageId="1" pageNumber="60" unit="mm" value="14.6" valueMax="14.7" valueMin="14.5">14.5–14.7 mm</quantity>
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(not including antenna), wing
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<quantity id="4CA69B37A833FFF5FBA1CD9BFAFF158B" box="[1139,1293,1945,1969]" metricMagnitude="-2" metricUnit="m" metricValue="1.6700000000000004" metricValueMax="1.6800000000000002" metricValueMin="1.6600000000000001" pageId="1" pageNumber="60" unit="mm" value="16.7" valueMax="16.8" valueMin="16.6">16.6–16.8 mm</quantity>
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, antenna
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<quantity id="4CA69B37A833FFF5FABDCD9BFF1515E9" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.66" metricValueMin="1.64" pageId="1" pageNumber="60" unit="mm" value="16.5" valueMax="16.6" valueMin="16.4">16.4– 16.6 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BE136D2A833FFF6FF6FCDDDFC0B131D" blockId="1.[136,1458,1703,2073]" lastBlockId="2.[136,1452,236,503]" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
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Head. Gray with rostrum darker in coloration laterally. Nasus black densely covered with black setae. Vertex gray, unpatterned (
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<figureCitation id="13652A57A833FFF5FECEC203FE931A23" box="[284,353,2049,2073]" captionStart="Figures 1–6" captionStartId="2.[155,227,2009,2031]" captionTargetBox="[189,1399,571,1946]" captionTargetId="figure-191@2.[189,1399,571,1969]" captionTargetPageId="2" captionText="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="1" pageNumber="60">Fig. 1</figureCitation>
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). Head including vertex and rostrum densely covered with black setae. Antenna 12-segmented, significantly longer than body; scape gray, pedicel and first two flagellomeres yellow, rest flagellomeres brown, each flagellomere subequal in length, cylindrical, with black verticils. Palpus dark gray.
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</paragraph>
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<paragraph id="8BE136D2A830FFF6FF6FCB30FC1813CD" blockId="2.[136,1452,236,503]" pageId="2" pageNumber="61">
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Thorax. Pronotum gray medially, darker in coloration laterally (
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<figureCitation id="13652A57A830FFF6FCA4CB30FC481370" box="[886,954,306,330]" captionStart="Figures 1–6" captionStartId="2.[155,227,2009,2031]" captionTargetBox="[189,1399,571,1946]" captionTargetId="figure-191@2.[189,1399,571,1969]" captionTargetPageId="2" captionText="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
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). Prescutum gray with three brownish-yellow stripes, the median one divided by a broad gray vitta (
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<figureCitation id="13652A57A830FFF6FD03CB56FCE01356" box="[721,786,340,364]" captionStart="Figures 1–6" captionStartId="2.[155,227,2009,2031]" captionTargetBox="[189,1399,571,1946]" captionTargetId="figure-191@2.[189,1399,571,1969]" captionTargetPageId="2" captionText="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="2" pageNumber="61">Fig. 3</figureCitation>
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). Scutum black with two gray triangular marks which are not connected to each other, scutellum and postnotum wholly gray (
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<figureCitation id="13652A57A830FFF6FC92CB75FC7013B5" box="[832,898,375,399]" captionStart="Figures 1–6" captionStartId="2.[155,227,2009,2031]" captionTargetBox="[189,1399,571,1946]" captionTargetId="figure-191@2.[189,1399,571,1969]" captionTargetPageId="2" captionText="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="2" pageNumber="61">Fig. 3</figureCitation>
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). Pleuron entirely gray (
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<figureCitation id="13652A57A830FFF6FB5FCB75FB2213B5" box="[1165,1232,375,399]" captionStart="Figures 1–6" captionStartId="2.[155,227,2009,2031]" captionTargetBox="[189,1399,571,1946]" captionTargetId="figure-191@2.[189,1399,571,1969]" captionTargetPageId="2" captionText="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
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). Legs very slender, coxae and trochanters gray, femora gray basally and gradually changed into dark brown, tibiae and tarsi dark brown. Halter black. Wing brownish-yellow, cell sc darker than ground color, stigma light brown (
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<figureCitation id="13652A57A830FFF6FBCFCBBEFB9313EE" box="[1053,1121,444,468]" captionStart="Figures 1–6" captionStartId="2.[155,227,2009,2031]" captionTargetBox="[189,1399,571,1946]" captionTargetId="figure-191@2.[189,1399,571,1969]" captionTargetPageId="2" captionText="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="2" pageNumber="61">Fig. 4</figureCitation>
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). Rs distinctly longer than R
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<subScript id="17DA3497A830FFF6FA4ECBC4FA5713EE" attach="left" box="[1436,1445,454,468]" fontSize="6" pageId="2" pageNumber="61">3</subScript>
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, petiole of cell m
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<subScript id="17DA3497A830FFF6FEE4CBEBFECD13CD" attach="left" box="[310,319,489,503]" fontSize="6" pageId="2" pageNumber="61">1</subScript>
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subequal in length to m-m, 1/3 the length of discal cell (
|
||
<figureCitation id="13652A57A830FFF6FC49CBDDFC2F13CD" box="[923,989,479,503]" captionStart="Figures 1–6" captionStartId="2.[155,227,2009,2031]" captionTargetBox="[189,1399,571,1946]" captionTargetId="figure-191@2.[189,1399,571,1969]" captionTargetPageId="2" captionText="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="2" pageNumber="61">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF21665AA830FFF6FF49CDDBFCAA1A35" ID-DOI="http://doi.org/10.5281/zenodo.4617190" ID-Zenodo-Dep="4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="2" pageNumber="61" startId="2.[155,227,2009,2031]" targetBox="[189,1399,571,1946]" targetPageId="2">
|
||
<paragraph id="8BE136D2A830FFF6FF49CDDBFCAA1A35" blockId="2.[155,1434,2009,2063]" pageId="2" pageNumber="61">
|
||
Figures 1–6.
|
||
<taxonomicName id="4C5E4D51A830FFF6FEC8CDDBFDBC15D4" authority="Men, 2020" authorityName="Men" authorityYear="2020" box="[282,590,2009,2030]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pianmaensis" status="sp. nov." subGenus="Sivatipula">
|
||
<emphasis id="B92AEAC0A830FFF6FEC8CDDBFEA515D4" box="[282,343,2009,2030]" italics="true" pageId="2" pageNumber="61">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B92AEAC0A830FFF6FEB4CDDBFE3515D4" box="[358,455,2009,2030]" italics="true" pageId="2" pageNumber="61">Sivatipula</emphasis>
|
||
)
|
||
<emphasis id="B92AEAC0A830FFF6FE04CDDBFDBC15D4" box="[470,590,2009,2030]" italics="true" pageId="2" pageNumber="61">pianmaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92AEAC0A830FFF6FD87CDDBFD5015D4" bold="true" box="[597,674,2009,2030]" pageId="2" pageNumber="61">
|
||
<taxonomicNameLabel id="A21957BBA830FFF6FD87CDDBFD5015D4" box="[597,674,2009,2030]" pageId="2" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE136D2A831FFF0FF6FCAEEFC44133F" blockId="3.[136,1452,236,468]" lastBlockId="4.[136,1452,236,642]" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
|
||
Abdomen. Segments one to five brownish-yellow with black lateral stripes, segments six and seven wholly black, tergite eight yellow, sternite eight mostly black (
|
||
<figureCitation id="13652A57A831FFF7FD9ACB0DFD7B131D" box="[584,649,271,295]" captionStart="Figures 1–6" captionStartId="2.[155,227,2009,2031]" captionTargetBox="[189,1399,571,1946]" captionTargetId="figure-191@2.[189,1399,571,1969]" captionTargetPageId="2" captionText="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4617190" httpUri="https://zenodo.org/record/4617190/files/figure.png" pageId="3" pageNumber="62">Fig. 5</figureCitation>
|
||
). Hypopygium yellow, distinctly enlarged (
|
||
<figureCitation id="13652A57A831FFF7FB8CCB0DFB33131D" box="[1118,1217,271,295]" captionStart-0="Figures 1–6" captionStart-1="Figures 7–15" captionStartId-0="2.[155,227,2009,2031]" captionStartId-1="3.[155,227,1986,2008]" captionTargetBox-0="[189,1399,571,1946]" captionTargetId-0="figure-191@2.[189,1399,571,1969]" captionTargetPageId-0="2" captionText-0="Figures 1–6. Tipula (Sivatipula) pianmaensis sp. nov. 1. Head, dorsal view. 2. Thorax, lateral view. 3. Thorax, dorsal view. 4. Wing. 5. Abdomen and hypopygiym, lateral view. 6. Hypopygium, lateral view." captionText-1="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4617190" httpUri-0="https://zenodo.org/record/4617190/files/figure.png" pageId="3" pageNumber="62">Figs 5–7</figureCitation>
|
||
). Tergite nine slightly protruding medially at posterior margin, terminating in a pair of horn-shaped processes at lateral corners, a pair of biforked lobes from ventral side of tergite nine, densely covered with black teeth apically (
|
||
<figureCitation id="13652A57A831FFF7FC38CB56FBBE1356" box="[1002,1100,340,364]" captionStart="Figures 7–15" captionStartId="3.[155,227,1986,2008]" captionText="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." pageId="3" pageNumber="62">Figs 7–9</figureCitation>
|
||
). Outer gonostylus, narrowed, Yshaped, with a finger-shaped process medially with numerous small black teeth at apex (
|
||
<figureCitation id="13652A57A831FFF7FB9FCB75FB6C13B5" box="[1101,1182,375,399]" captionStart="Figures 7–15" captionStartId="3.[155,227,1986,2008]" captionText="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." pageId="3" pageNumber="62">Fig. 10</figureCitation>
|
||
). Inner gonostylus broad basally, terminating in a long black beak, a small obtuse lobe and two finger-shaped processes arising from its mesal region of outer surface (
|
||
<figureCitation id="13652A57A831FFF7FE91CBBEFE6113EE" box="[323,403,444,468]" captionStart="Figures 7–15" captionStartId="3.[155,227,1986,2008]" captionText="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." pageId="3" pageNumber="62">Fig. 11</figureCitation>
|
||
). Sternite nine with a pair of small median processes which are densely covered with long setae (
|
||
<figureCitation id="13652A57A836FFF0FF42CAEEFF12133F" box="[144,224,236,261]" captionStart="Figures 7–15" captionStartId="3.[155,227,1986,2008]" captionText="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." pageId="4" pageNumber="63">Fig. 12</figureCitation>
|
||
). Adminiculum broad basally, gradually narrowed to apex (
|
||
<figureCitation id="13652A57A836FFF0FC88CAEEFC5B133F" box="[858,937,236,261]" captionStart="Figures 7–15" captionStartId="3.[155,227,1986,2008]" captionText="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." pageId="4" pageNumber="63">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF21665AA831FFF7FF49CDC0FB531A22" pageId="3" pageNumber="62">
|
||
<paragraph id="8BE136D2A831FFF7FF49CDC0FB531A22" blockId="3.[155,1433,1986,2072]" pageId="3" pageNumber="62">
|
||
Figures 7–15.
|
||
<taxonomicName id="4C5E4D51A831FFF7FEF8CDC0FD9315E2" authority="Men, 2020" authorityName="Men" authorityYear="2020" box="[298,609,1986,2008]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pianmaensis" status="sp. nov." subGenus="Sivatipula">
|
||
<emphasis id="B92AEAC0A831FFF7FEF8CDC0FE9515ED" box="[298,359,1986,2007]" italics="true" pageId="3" pageNumber="62">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B92AEAC0A831FFF7FEAACDC0FE2B15ED" box="[376,473,1986,2007]" italics="true" pageId="3" pageNumber="62">Sivatipula</emphasis>
|
||
)
|
||
<emphasis id="B92AEAC0A831FFF7FE3BCDC1FD9315E2" box="[489,609,1987,2008]" italics="true" pageId="3" pageNumber="62">pianmaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92AEAC0A831FFF7FDB8CDC1FD4B15E2" bold="true" box="[618,697,1987,2008]" pageId="3" pageNumber="62">
|
||
<taxonomicNameLabel id="A21957BBA831FFF7FDB8CDC1FD4B15E2" box="[618,697,1987,2008]" pageId="3" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE136D2A836FFF0FF6FCB0DFD821356" blockId="4.[136,1452,236,642]" pageId="4" pageNumber="63">
|
||
Semen pump with compressor apodeme V-shaped (
|
||
<figureCitation id="13652A57A836FFF0FD0DCB0DFCDC131D" box="[735,814,271,295]" captionStart="Figures 7–15" captionStartId="3.[155,227,1986,2008]" captionText="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." pageId="4" pageNumber="63">Fig. 14</figureCitation>
|
||
). Posterior immovable apodeme consisting of a median arm which curved dorsally (
|
||
<figureCitation id="13652A57A836FFF0FE5BCB30FE2B1373" box="[393,473,305,330]" captionStart="Figures 7–15" captionStartId="3.[155,227,1986,2008]" captionText="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." pageId="4" pageNumber="63">Fig. 15</figureCitation>
|
||
). Anterior immovable apodeme flattened. Aedeagus narrow, elongated, tubular, more than 5.0 times the length of semen pump (
|
||
<figureCitation id="13652A57A836FFF0FDC6CB56FD911356" box="[532,611,340,364]" captionStart="Figures 7–15" captionStartId="3.[155,227,1986,2008]" captionText="Figures 7–15. Tipula (Sivatipula) pianmaensis sp. nov. 7. Hypopygium, lateral view. 8. Tergite nine, dorsal view. 9. Tergite nine, posteroventral view. 10. Outer gonostylus, inner view. 11. Inner gonostylus, outer view. 12. Median processes on sternite nine, dorsal view. 13. Adminiculum, dorsal view. 14. Compressor apodeme, dorsal view. 15. Semen pump, lateral view." pageId="4" pageNumber="63">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE136D2A836FFF0FF6FCB75FE0F13B5" blockId="4.[136,1452,236,642]" box="[189,509,375,399]" pageId="4" pageNumber="63">
|
||
Distribution.
|
||
<collectingCountry id="F3497642A836FFF0FE9ECB75FE7E13B5" box="[332,396,375,399]" name="China" pageId="4" pageNumber="63">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="499AF830A836FFF0FE48CB75FE0013B5" box="[410,498,375,399]" country="China" name="Yunnan" pageId="4" pageNumber="63">Yunnan</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE136D2A836FFF0FF6FCB98FDB51006" blockId="4.[136,1452,236,642]" pageId="4" pageNumber="63">
|
||
Remarks. The new species is mostly similar to
|
||
<emphasis id="B92AEAC0A836FFF0FD7ECB98FD49138B" box="[684,699,410,433]" italics="true" pageId="4" pageNumber="63">T</emphasis>
|
||
.
|
||
<taxonomicName id="4C5E4D51A836FFF0FD15CB98FBF11388" authority="Alexander, 1964" authorityName="Alexander" authorityYear="1964" box="[711,1027,410,434]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="bhishma" subGenus="Sivatipula">
|
||
(
|
||
<emphasis id="B92AEAC0A836FFF0FD02CB98FD2F138B" box="[720,733,410,433]" italics="true" pageId="4" pageNumber="63">S</emphasis>
|
||
.)
|
||
<emphasis id="B92AEAC0A836FFF0FD20CB98FCBE1388" box="[754,844,410,434]" italics="true" pageId="4" pageNumber="63">bhishma</emphasis>
|
||
<bibRefCitation id="EFCF4B23A836FFF0FC83CB98FBF11388" author="Alexander" box="[849,1027,410,434]" pageId="4" pageNumber="63" pagination="77 - 130" refId="ref4367" refString="Alexander, C. P. 1964. New or little-known Tipulidae from eastern Asia (Diptera). LIII. Philippine Journal of Science, 93: 77 - 130." type="journal article" year="1964">Alexander, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
, a species from
|
||
<collectingRegion id="499AF830A836FFF0FB7BCB98FB061388" box="[1193,1268,410,434]" country="India" name="Assam" pageId="4" pageNumber="63">Assam</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3497642A836FFF0FB2CCB98FACA1388" box="[1278,1336,410,434]" name="India" pageId="4" pageNumber="63">India</collectingCountry>
|
||
, by similar structure of hypopygium and body color. It can be easily distinguished from
|
||
<emphasis id="B92AEAC0A836FFF0FC6CCBBFFC3F13EE" box="[958,973,445,468]" italics="true" pageId="4" pageNumber="63">T</emphasis>
|
||
.
|
||
<taxonomicName id="4C5E4D51A836FFF0FC08CBBEFB9313EE" authorityName="Alexander" authorityYear="1964" box="[986,1121,444,468]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="bhishma" subGenus="Sivatipula">
|
||
(
|
||
<emphasis id="B92AEAC0A836FFF0FC31CBBFFC0213EE" box="[995,1008,445,468]" italics="true" pageId="4" pageNumber="63">S</emphasis>
|
||
.)
|
||
<emphasis id="B92AEAC0A836FFF0FBD5CBBEFB9313EE" box="[1031,1121,444,468]" italics="true" pageId="4" pageNumber="63">bhishma</emphasis>
|
||
</taxonomicName>
|
||
by the outer gonostylus which is Y-shaped with a finger-shaped process (absent in
|
||
<emphasis id="B92AEAC0A836FFF0FD61CBE2FD3013CD" box="[691,706,480,503]" italics="true" pageId="4" pageNumber="63">T</emphasis>
|
||
.
|
||
<taxonomicName id="4C5E4D51A836FFF0FD02CBDDFCA413CD" authorityName="Alexander" authorityYear="1964" box="[720,854,479,503]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="bhishma" subGenus="Sivatipula">
|
||
(
|
||
<emphasis id="B92AEAC0A836FFF0FD0ACBDDFD1713CC" box="[728,741,479,502]" italics="true" pageId="4" pageNumber="63">S</emphasis>
|
||
.)
|
||
<emphasis id="B92AEAC0A836FFF0FD2ECBDDFCA413CD" box="[764,854,479,503]" italics="true" pageId="4" pageNumber="63">bhishma</emphasis>
|
||
</taxonomicName>
|
||
). The latter species has its outer gonostylus not bearing finger-shaped process. There is also a noticeable difference in the shape of process on tergite nine which is narrow in the new species, but broad in
|
||
<emphasis id="B92AEAC0A836FFF0FE4FC827FE5E1006" box="[413,428,549,572]" italics="true" pageId="4" pageNumber="63">T</emphasis>
|
||
.
|
||
<taxonomicName id="4C5E4D51A836FFF0FE6BC826FDB21006" authorityName="Alexander" authorityYear="1964" box="[441,576,548,572]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="bhishma" subGenus="Sivatipula">
|
||
(
|
||
<emphasis id="B92AEAC0A836FFF0FE10C827FE3D1006" box="[450,463,549,572]" italics="true" pageId="4" pageNumber="63">S</emphasis>
|
||
.)
|
||
<emphasis id="B92AEAC0A836FFF0FE34C826FDB21006" box="[486,576,548,572]" italics="true" pageId="4" pageNumber="63">bhishma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE136D2A836FFF0FF6FC845FEC810B8" blockId="4.[136,1452,236,642]" pageId="4" pageNumber="63">
|
||
Etymology. The specific epithet is a noun “
|
||
<emphasis id="B92AEAC0A836FFF0FD5CC84AFD2C1065" box="[654,734,584,607]" italics="true" pageId="4" pageNumber="63">pianma</emphasis>
|
||
” with Latin suffix “
|
||
<emphasis id="B92AEAC0A836FFF0FC65C84AFC1E1065" box="[951,1004,584,607]" italics="true" pageId="4" pageNumber="63">ensis</emphasis>
|
||
”, referring to the
|
||
<typeStatus id="54E58870A836FFF0FB7CC845FB2E1065" box="[1198,1244,583,607]" pageId="4" pageNumber="63">type</typeStatus>
|
||
locality of the new species, Pianma.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |