290 lines
40 KiB
XML
290 lines
40 KiB
XML
<document id="54B554AA6BE20FAFCA104842F389D839" ID="10.11646/zootaxa.4121.2.7" ID-CLB-Dataset="38916" ID-DOI="10.11646/zootaxa.4121.2.7" ID-GBIF-Dataset="43b4c678-f20c-4f05-b1db-8b876d39f8ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="265566" ID-ZooBank="9D2166F7-C1E0-4018-88D1-1FB2064F0EC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465368266281" checkinUser="plazi" docAuthor="Nguyen, Lien Thi Phuong" docDate="2016" docId="03F70157FF992A32FF14F8FB7590FD6E" docLanguage="en" docName="zootaxa.4121.2.7.pdf" docOrigin="Zootaxa 4121 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeleumenes burmanicus Bingham 1897" docType="treatment" docVersion="9" lastPageNumber="178" masterDocId="FFCE792FFF992A31FF83FFEC762FFFEB" masterDocTitle="Contribution to the taxonomy of the genus Coeleumenes van der Vecht, 1963 (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species" masterLastPageNumber="180" masterPageNumber="175" pageNumber="175" updateTime="1698662765855" updateUser="ExternalLinkService">
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<mods:title id="9854B6743A6FD1BEDBC4524B78FF0D77">Contribution to the taxonomy of the genus Coeleumenes van der Vecht, 1963 (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species</mods:title>
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<mods:name id="253D2E7AB19A34F75D79241B456F770F" type="personal">
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<mods:namePart id="A7C4C8800F8CE63F2A35FC093DF82FBD">Nguyen, Lien Thi Phuong</mods:namePart>
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<mods:title id="158001AB9048BF7CD63965C8DB7CCCFA">Zootaxa</mods:title>
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<mods:date id="D2573C224E56825F01F6D0EB59D485C6">2016</mods:date>
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<mods:number id="8D1F1099624F1C45007DC725B4D7AC34">4121</mods:number>
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<treatment id="03F70157FF992A32FF14F8FB7590FD6E" ID-DOI="http://doi.org/10.5281/zenodo.6088205" ID-GBIF-Taxon="120825340" ID-Zenodo-Dep="6088205" LSID="urn:lsid:plazi:treatment:03F70157FF992A32FF14F8FB7590FD6E" httpUri="http://treatment.plazi.org/id/03F70157FF992A32FF14F8FB7590FD6E" lastPageId="3" lastPageNumber="178" pageId="0" pageNumber="175">
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<subSubSection id="C344E3CAFF992A31FF14F8FB7724F8B8" pageId="0" pageNumber="175" type="nomenclature">
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<paragraph id="8BE1B041FF992A31FF14F8FB7489F8D9" blockId="0.[151,678,1815,1875]" box="[151,678,1815,1842]" pageId="0" pageNumber="175">
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<heading id="D0A9072DFF992A31FF14F8FB7489F8D9" bold="true" box="[151,678,1815,1842]" fontSize="11" level="1" pageId="0" pageNumber="175" reason="1">
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<taxonomicName id="4C5ECBC2FF992A31FF14F8FB7489F8D9" ID-CoL="5ZH5C" authority="Bingham, 1897" authorityName="Bingham" authorityYear="1897" box="[151,678,1815,1842]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="burmanicus">
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<emphasis id="B92A6C53FF992A31FF14F8FB7489F8D9" bold="true" box="[151,678,1815,1842]" pageId="0" pageNumber="175">
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<emphasis id="B92A6C53FF992A31FF14F8FB77FFF8DA" bold="true" box="[151,464,1815,1841]" italics="true" pageId="0" pageNumber="175">Coeleumenes burmanicus</emphasis>
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(
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<bibRefCitation id="EFCFCDB0FF992A31FE5CF8F474B1F8D9" author="Bingham" box="[479,670,1816,1842]" pageId="0" pageNumber="175" refString="Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera. Vol. 1. Wasps and Bees. Taylor and Francis, London, 579 pp." type="book" year="1897">Bingham, 1897</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE1B041FF992A31FF14F8D67724F8B8" blockId="0.[151,678,1815,1875]" box="[151,267,1850,1875]" pageId="0" pageNumber="175">
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(
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<figureCitation id="1365ACC4FF992A31FF1CF8D6772DF8B8" box="[159,258,1850,1875]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="0" pageNumber="175">Figs 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C344E3CAFF992A30FF14F86F7513FF00" lastPageId="1" lastPageNumber="176" pageId="0" pageNumber="175" type="reference_group">
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<paragraph id="8BE1B041FF992A31FF14F86F7708F85C" blockId="0.[151,1436,1922,2037]" pageId="0" pageNumber="175">
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<treatmentCitationGroup id="AB4E976FFF992A31FF14F86F7708F85C" pageId="0" pageNumber="175">
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<taxonomicName id="4C5ECBC2FF992A31FF14F86F7449F873" authority="Bingham, 1897: 350" authorityName="Bingham" authorityPageNumber="350" authorityYear="1897" box="[151,614,1922,1944]" class="Insecta" family="Eumenidae" genus="Montezumia" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="burmanica">
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<emphasis id="B92A6C53FF992A31FF14F86F77A5F873" box="[151,394,1922,1944]" italics="true" pageId="0" pageNumber="175">Montezumia burmanica</emphasis>
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<treatmentCitation id="0AFF9650FF992A31FE11F86E7449F873" author="Bingham" box="[402,614,1922,1944]" page="350" pageId="0" pageNumber="175" year="1897">
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<bibRefCitation id="EFCFCDB0FF992A31FE11F86E741CF873" author="Bingham" box="[402,563,1922,1944]" pageId="0" pageNumber="175" refString="Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera. Vol. 1. Wasps and Bees. Taylor and Francis, London, 579 pp." type="book" year="1897">Bingham, 1897</bibRefCitation>
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: 350
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</treatmentCitation>
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</taxonomicName>
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, fig. 101, ♀, ♂, “Tenasserim” (The Natural History Museum, London,
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<typeStatus id="54E50EE3FF992A31FACBF86F735BF873" box="[1352,1396,1923,1944]" pageId="0" pageNumber="175">type</typeStatus>
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no. 18.212a).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE1B041FF992A30FF14F82D74E8FF45" blockId="0.[151,1436,1922,2037]" lastBlockId="1.[151,1435,152,235]" lastPageId="1" lastPageNumber="176" pageId="0" pageNumber="175">
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<treatmentCitationGroup id="AB4E976FFF992A30FF14F82D74E8FF45" lastPageId="1" lastPageNumber="176" pageId="0" pageNumber="175">
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<taxonomicName id="4C5ECBC2FF992A31FF14F82D7460F83D" authority="Cameron, 1900: 535" authorityName="Cameron" authorityPageNumber="535" authorityYear="1900" box="[151,591,1984,2006]" class="Insecta" family="Eumenidae" genus="Montezumia" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="bisulcata">
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<emphasis id="B92A6C53FF992A31FF14F82D7758F83D" box="[151,375,1984,2006]" italics="true" pageId="0" pageNumber="175">Montezumia bisulcata</emphasis>
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<treatmentCitation id="0AFF9650FF992A31FEFEF82C7460F83D" author="Cameron" box="[381,591,1984,2006]" page="535" pageId="0" pageNumber="175" year="1900">
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<bibRefCitation id="EFCFCDB0FF992A31FEFEF82C7431F83D" author="Cameron" box="[381,542,1984,2006]" pageId="0" pageNumber="175" refString="Cameron, P. (1900) Descriptions of new genera and species of Hymenoptera. Annals and Magazine of Natural History, Series 7, 6, 530 - 539. http: // dx. doi. org / 10.1080 / 00222930008678422" type="journal article" year="1900">
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<collectingCountry id="F349F0D1FF992A31FEFEF82C77F3F83D" box="[381,476,1984,2006]" name="Cameroon" pageId="0" pageNumber="175">Cameron</collectingCountry>
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, 1900
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</bibRefCitation>
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: 535
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</treatmentCitation>
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</taxonomicName>
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, ♀, “Khasia Hills, Assam” (Oxford University Museum, Oxford).
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<taxonomicName id="4C5ECBC2FF992A31FF14F80C77A4F81F" box="[151,395,2014,2037]" class="Insecta" family="Eumenidae" genus="Montezumia" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="175" phylum="Arthropoda" rank="species" species="burmanica">
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<emphasis id="B92A6C53FF992A31FF14F80C77A4F81F" box="[151,395,2014,2037]" italics="true" pageId="0" pageNumber="175">Montezumia burmanica</emphasis>
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</taxonomicName>
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<taxonomicName id="4C5ECBC2FF992A31FE17F83374C5F81F" authority="Dover, 1931: 253" authorityName="Dover" authorityPageNumber="253" authorityYear="1931" box="[404,746,2014,2036]" pageId="0" pageNumber="175" rank="variety" variety="malayana">
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var.
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<emphasis id="B92A6C53FF992A31FE40F8327408F81F" box="[451,551,2014,2036]" italics="true" pageId="0" pageNumber="175">malayana</emphasis>
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||
<treatmentCitation id="0AFF9650FF992A31FDB2F83374C5F81F" author="Dover" box="[561,746,2014,2036]" page="253" pageId="0" pageNumber="175" year="1931">
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<bibRefCitation id="EFCFCDB0FF992A31FDB2F833749AF81F" author="Dover" box="[561,693,2014,2036]" pageId="0" pageNumber="175" refString="Dover, C. (1931 [" 1930 "]) The Vespidae in the Federated Malay State Museums. Journal of the Federated Malay State Museums. 16, 251 - 260." type="journal article" year="1931">Dover, 1931</bibRefCitation>
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: 253
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||
</treatmentCitation>
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||
</taxonomicName>
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||
, ♀, “Jor Camp, Perak,
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<collectingCountry id="F349F0D1FF992A31FC60F832721EF81F" box="[995,1073,2014,2036]" name="Malaysia" pageId="0" pageNumber="175">Malaya</collectingCountry>
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||
” (? The Natural History Museum, London). Synonymized by van der
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<treatmentCitation id="0AFF9650FF982A30FDAEFF7474E8FF45" author="Vecht" box="[557,711,152,174]" page="46" pageId="1" pageNumber="176" year="1963">
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||
<bibRefCitation id="EFCFCDB0FF982A30FDAEFF74748BFF45" author="Vecht" box="[557,676,152,174]" pageId="1" pageNumber="176" refString="Vecht, J. van der (1963) Studies on Indo-Australian and East-Asiatic Eumenidae (Hymenoptera, Vespoidea). Zoologische Verhandelingen, Leiden, 60, 3 - 113." type="journal article" year="1963">Vecht 1963</bibRefCitation>
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||
: 46
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</treatmentCitation>
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BE1B041FF982A30FF14FF5A7513FF00" blockId="1.[151,1435,152,235]" pageId="1" pageNumber="176">
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||
<treatmentCitationGroup id="AB4E976FFF982A30FF14FF5A7513FF00" pageId="1" pageNumber="176">
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||
<taxonomicName id="4C5ECBC2FF982A30FF14FF5A7494FF27" ID-CoL="5ZH5C" authority="van der Vecht, 1963: 46" authorityName="van der Vecht" authorityPageNumber="46" authorityYear="1963" box="[151,699,182,204]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="176" phylum="Arthropoda" rank="species" species="burmanicus">
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||
<emphasis id="B92A6C53FF982A30FF14FF5A778FFF27" box="[151,416,182,204]" italics="true" pageId="1" pageNumber="176">Coeleumenes burmanicus</emphasis>
|
||
: van der
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||
<treatmentCitation id="0AFF9650FF982A30FD8CFF5A7494FF27" author="Vecht" box="[527,699,182,204]" page="46" pageId="1" pageNumber="176" year="1963">
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<bibRefCitation id="EFCFCDB0FF982A30FD8CFF5A74BFFF27" author="Vecht" box="[527,656,182,204]" pageId="1" pageNumber="176" refString="Vecht, J. van der (1963) Studies on Indo-Australian and East-Asiatic Eumenidae (Hymenoptera, Vespoidea). Zoologische Verhandelingen, Leiden, 60, 3 - 113." type="journal article" year="1963">Vecht, 1963</bibRefCitation>
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||
: 46
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||
</treatmentCitation>
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</taxonomicName>
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, [Sikkim; Assam; Indo-China;
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<collectingCountry id="F349F0D1FF982A30FB8FFF5A726CFF27" box="[1036,1091,182,204]" name="Thailand" pageId="1" pageNumber="176">Siam</collectingCountry>
|
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;
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||
<collectingCountry id="F349F0D1FF982A30FBD1FF5A728EFF27" box="[1106,1185,182,204]" name="Malaysia" pageId="1" pageNumber="176">Malaya</collectingCountry>
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||
;
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||
<collectingCountry id="F349F0D1FF982A30FB33FF5A7329FF27" box="[1200,1286,182,204]" name="Vietnam" pageId="1" pageNumber="176">Vietnam</collectingCountry>
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(Than Moi)];
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<treatmentCitation id="0AFF9650FF982A30FF44FF3A778CFF00" author="Nguyen" box="[199,419,213,235]" page="9" pageId="1" pageNumber="176" year="2014">
|
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<bibRefCitation id="EFCFCDB0FF982A30FF44FF3A77A2FF00" author="Nguyen" box="[199,397,213,235]" pageId="1" pageNumber="176" refString="Nguyen, L. T. P., Dang, H. T., Kojima, J. & Carpenter, J. M. (2014) An annotated distributional checklist of solitary wasps of the subfamily Eumeninae (Hymenoptera: Vespidae) of Vietnam. Entomologica Americana, 120 (1), 7 - 17. http: // dx. doi. org / 10.1664 / 13 - RA- 010.1" type="journal article" year="2014">
|
||
Nguyen
|
||
<emphasis id="B92A6C53FF982A30FE9EFF3A777EFF00" box="[285,337,213,235]" italics="true" pageId="1" pageNumber="176">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
: 9
|
||
</treatmentCitation>
|
||
(
|
||
<collectingCountry id="F349F0D1FF982A30FE33FF397424FF00" box="[432,523,213,235]" name="Vietnam" pageId="1" pageNumber="176">Vietnam</collectingCountry>
|
||
: Bac Kan, Hoa Binh, Gia Lai)
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C344E3CAFF982A30FF14FEFB728FFE0E" pageId="1" pageNumber="176" type="materials_examined">
|
||
<paragraph id="8BE1B041FF982A30FF14FEFB728FFE0E" blockId="1.[151,1437,279,2033]" pageId="1" pageNumber="176">
|
||
<emphasis id="B92A6C53FF982A30FF14FEFB77AFFEDB" bold="true" box="[151,384,279,304]" pageId="1" pageNumber="176">Material examined.</emphasis>
|
||
<collectingCountry id="F349F0D1FF982A30FE04FEF47427FEDB" box="[391,520,280,304]" name="Vietnam" pageId="1" pageNumber="176">VIETNAM</collectingCountry>
|
||
[
|
||
<collectionCode id="ED4F2884FF982A30FD96FEF47471FEC4" box="[533,606,280,303]" pageId="1" pageNumber="176">IEBR</collectionCode>
|
||
]:
|
||
<emphasis id="B92A6C53FF982A30FDECFEFB753EFEDB" bold="true" box="[623,785,279,304]" pageId="1" pageNumber="176">Tuyen Quang</emphasis>
|
||
: 1 ♀, Na Hang natural rreserve, Na Hang, Son Phu ranger station, 22°21.2'07"N
|
||
<geoCoordinate id="EE6AD686FF982A30FE11FED07417FEBE" box="[402,568,316,341]" direction="east" orientation="longitude" pageId="1" pageNumber="176" precision="1" value="105.4263">105°25'34.7"E</geoCoordinate>
|
||
,
|
||
<quantity id="4CA61DA4FF982A30FDC6FED074A1FEBF" box="[581,654,316,341]" metricMagnitude="2" metricUnit="m" metricValue="2.6399999999999997" pageId="1" pageNumber="176" unit="m" value="264.0">264 m</quantity>
|
||
, LTP Nguyen, DD Nguyen & LX Truong,
|
||
<date id="FFE09681FF982A30FBFBFED072CBFEBE" box="[1144,1252,316,341]" pageId="1" pageNumber="176" value="2015-06-09">9.vi.2015</date>
|
||
;
|
||
<emphasis id="B92A6C53FF982A30FB72FED07376FEBE" bold="true" box="[1265,1369,316,341]" pageId="1" pageNumber="176">Bac Kan</emphasis>
|
||
: 1 ♀, Kim Hy NP, Vu Muon, Bach Thong,
|
||
<geoCoordinate id="EE6AD686FF982A30FDBDFEB374EDFE93" box="[574,706,351,376]" direction="north" orientation="latitude" pageId="1" pageNumber="176" precision="15" value="22.214167">22°12'51"N</geoCoordinate>
|
||
<geoCoordinate id="EE6AD686FF982A30FD4AFEB37574FE93" box="[713,859,351,376]" direction="east" orientation="longitude" pageId="1" pageNumber="176" precision="15" value="105.97833">105°58'42"E</geoCoordinate>
|
||
,
|
||
<date id="FFE09681FF982A30FCE4FEB375CCFE93" box="[871,995,351,376]" pageId="1" pageNumber="176" value="2012-08-05">5.viii.2012</date>
|
||
, J Kojima, H Nugroho & IED-c;
|
||
<emphasis id="B92A6C53FF982A30FADEFEB376EBFE76" bold="true" pageId="1" pageNumber="176">Lang Son</emphasis>
|
||
: Cai Kinh, Huu Lung,
|
||
<geoCoordinate id="EE6AD686FF982A30FE48FE687460FE76" box="[459,591,388,413]" direction="north" orientation="latitude" pageId="1" pageNumber="176" precision="15" value="21.861668">21°51'42"N</geoCoordinate>
|
||
<geoCoordinate id="EE6AD686FF982A30FDD4FE6874C9FE76" box="[599,742,388,413]" direction="east" orientation="longitude" pageId="1" pageNumber="176" precision="15" value="106.73167">106°43'54"E</geoCoordinate>
|
||
[1 ♀,
|
||
<date id="FFE09681FF982A30FCB3FE6875B3FE76" box="[816,924,388,413]" pageId="1" pageNumber="176" value="2014-11-02">2.xi.2014</date>
|
||
, LTP Nguyen, DD Nguyen & MP Nguyen; 1 ♀,
|
||
<specimenCount id="9D587BC8FF982A30FF3FFE4B76C4FE54" box="[188,235,423,448]" pageId="1" pageNumber="176" type="generic">1 ♂</specimenCount>
|
||
,
|
||
<date id="FFE09681FF982A30FF7AFE4B775CFE2B" box="[249,371,423,448]" pageId="1" pageNumber="176" value="2015-11-24">24.xi.2015</date>
|
||
, LTP Nguyen, DD Nguyen & NT Tran];
|
||
<emphasis id="B92A6C53FF982A30FCD1FE4B75E9FE2B" bold="true" box="[850,966,423,448]" pageId="1" pageNumber="176">Hoa Binh</emphasis>
|
||
: 1 ♀, Da Phuc, Yen Thuy,
|
||
<date id="FFE09681FF982A30FA85FE447346FE2B" box="[1286,1385,424,448]" pageId="1" pageNumber="176" value="2002-05-03">3.v.2002</date>
|
||
, TV Hoang;
|
||
<emphasis id="B92A6C53FF982A30FF73FE20777BFE0E" bold="true" box="[240,340,460,485]" pageId="1" pageNumber="176">Ha Tinh</emphasis>
|
||
: 1 ♀, Khe Van, Son Hong, Huong Son,
|
||
<quantity id="4CA61DA4FF982A30FC8DFE2075A4FE0F" box="[782,907,460,484]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="1" pageNumber="176" unit="m" value="150.0" valueMax="180.0" valueMin="120.0">120-180 m</quantity>
|
||
,
|
||
<date id="FFE09681FF982A30FC14FE207220FE0E" box="[919,1039,460,485]" pageId="1" pageNumber="176" value="2004-04-24">24.iv.2004</date>
|
||
, LX Truong.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C344E3CAFF982A32FF44FE037723FE46" lastPageId="3" lastPageNumber="178" pageId="1" pageNumber="176" type="description">
|
||
<paragraph id="8BE1B041FF982A30FF44FE037789FC53" blockId="1.[151,1437,279,2033]" pageId="1" pageNumber="176">
|
||
<emphasis id="B92A6C53FF982A30FF44FE03775BFDE3" bold="true" box="[199,372,495,520]" pageId="1" pageNumber="176">Redescription.</emphasis>
|
||
<emphasis id="B92A6C53FF982A30FEFEFE1C77FEFDE3" box="[381,465,496,520]" italics="true" pageId="1" pageNumber="176">Female</emphasis>
|
||
(n=6). Body length 13.0–15.0 mm; fore wing length
|
||
<quantity id="4CA61DA4FF982A30FBB9FE0372CEFDEC" box="[1082,1249,495,520]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.35" metricValueMin="1.15" pageId="1" pageNumber="176" unit="mm" value="12.5" valueMax="13.5" valueMin="11.5">11.5–13.5 mm</quantity>
|
||
. Head in frontal view subcircular, 1.2 × as wide as high (
|
||
<figureCitation id="1365ACC4FF982A30FDDFFDF874B2FDC7" box="[604,669,532,557]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="1" pageNumber="176">Fig. 1</figureCitation>
|
||
). Vertex with small cephalic foveae, with some pubescence, located close to each other, with distance between them about one-half distance between posterior ocelli; with shallow depression for cephalic foveae, which located inside faint V-shaped edge (
|
||
<figureCitation id="1365ACC4FF982A30FC55FDB07234FD9F" box="[982,1051,604,629]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="1" pageNumber="176">Fig. 2</figureCitation>
|
||
). Distance from posterior ocelli to apical margin of vertex 2.2 × distance from posterior ocelli to inner eye margin (
|
||
<figureCitation id="1365ACC4FF982A30FB9DFD93724BFD73" box="[1054,1124,639,664]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="1" pageNumber="176">Fig. 2</figureCitation>
|
||
). Gena in lateral view 0.5 × as wide as eye. Occipital carina developed only laterally. Inner eye margin slightly convergent ventrally, in frontal view slightly further apart from each other at vertex than at clypeus. Clypeus in lateral view gradually convex from base to basal half, then nearly straight to apical margin, in frontal view 1.2 × as high as wide (
|
||
<figureCitation id="1365ACC4FF982A30FB4FFD007320FCEF" box="[1228,1295,748,773]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="1" pageNumber="176">Fig. 1</figureCitation>
|
||
), with basal margin close to antennal sockets, apical margin deeply emarginated medially, forming sharp tooth on each lateral side, width of emargination about 1/3 width of clypeus between inner eye margins. Mandible with four prominent teeth, fourth tooth long and blunt apically. Scape more than 4 × as long as its maximum width; F1 2.3 × as long as wide, F2–5 longer than its width, F6 as long as wide, F7–9 wider than its length, terminal flagellomere bulletshaped, as long as wide.
|
||
</paragraph>
|
||
<paragraph id="8BE1B041FF982A30FF44FC29741CFB86" blockId="1.[151,1437,279,2033]" pageId="1" pageNumber="176">
|
||
Mesosoma dorsally longer than its width. Pronotal carina slightly raised, reaching ventral corner of pronotum. Mesoscutum weakly convex, shorter than its width between tegulae, about 0.9 × as long as wide; anterior margin broadly rounded. Disc of scutellum slightly convex, in profile at same level as mesoscutum. Metanotum flat. Propodeum without shelf, posterior surface not concave, with deep median groove (
|
||
<figureCitation id="1365ACC4FF982A30FBE9FBC372FEFBA3" box="[1130,1233,1071,1096]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="1" pageNumber="176">Fig. 3, 5</figureCitation>
|
||
), border between posterior and lateral surfaces angled.
|
||
</paragraph>
|
||
<paragraph id="8BE1B041FF982A30FF44FB9B750EFB33" blockId="1.[151,1437,279,2033]" pageId="1" pageNumber="176">
|
||
Metasomal segment 1 subpetiolate, shorter than segment 2. T1 dorsally 1.5 × as long as wide, with node at lateral half (
|
||
<figureCitation id="1365ACC4FF982A30FE9CFB70774DFB5F" box="[287,354,1180,1205]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="1" pageNumber="176">Fig. 3</figureCitation>
|
||
); T2 as wide as long (
|
||
<figureCitation id="1365ACC4FF982A30FDD9FB7074B2FB5F" box="[602,669,1180,1205]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="1" pageNumber="176">Fig. 3</figureCitation>
|
||
), with apical lamella short and slightly raised; S
|
||
<quantity id="4CA61DA4FF982A30FB39FB7072C9FB5E" box="[1210,1254,1180,1205]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="176" unit="in" value="2.0">2 in</quantity>
|
||
lateral view flat in basal half, angled and straight to apical margin (
|
||
<figureCitation id="1365ACC4FF982A30FD4CFB53753BFB33" box="[719,788,1215,1240]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="1" pageNumber="176">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE1B041FF982A30FF44FB0874A1F8D6" blockId="1.[151,1437,279,2033]" pageId="1" pageNumber="176">Body with short, silver setae except front, posterior part of head, lower part of propodeum, mesepisternum and S1 with long silver setae. Clypeus with large and deep punctures, border between punctures with small punctures, clypeus almost flat in middle of apical half, with sparse punctures and irregular striae. Mandible with several shallow small punctures. Frons with dense shallow large flat-bottomed punctures, vertex and gena with smaller and more clearly defined punctures. Pronotum with coarse punctures, spaces between them narrow, strongly raised, forming reticulation. Mesocutum with dense and coarse flat-bottomed punctures, punctures on anterior half similar to those on pronotum, posterior half with some smooth areas near apical margin and tegulae. Sculellum with punctures similar to those on pronotum, with median longitudinal faint carina on basal half, area along carina to apical margin smooth. Metanotum with punctures very coarse, with median longitudinal smooth area. Tegula sparsely punctate at base, remainder micropunctate. Mesepisternum with epicnemial carina, with bigger and coarser punctures posterodorsally than on pronotum, reticulated, barely punctured anteroventrally; border between posterodorsal and anteroventral parts distinct. Metapleuron almost smooth. Propodeum with posterior face irregularly rugose, lateral parts dull, with some punctures and transversely striate. T1 with deep, dense well defined punctures, T2 with sparser and shallower punctures than in T1, area between punctures smooth, punctures on T3– T4 much smaller and weaker than those on T1–T2, T5–T6 with minute punctures; punctures on S2 uniform and slightly deeper than those on T2, S1 with strong transverse striae throughout sternum, with narrow basal part smooth, punctures on S3–S5 small and deep.</paragraph>
|
||
<paragraph id="8BE1B041FF982A33FF44F8A4774EFF3E" blockId="1.[151,1437,279,2033]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="177" pageId="1" pageNumber="176">
|
||
<emphasis id="B92A6C53FF982A30FF44F8A47728F88B" box="[199,263,1864,1888]" italics="true" pageId="1" pageNumber="176">Color</emphasis>
|
||
. Black with some yellow markings as follows: spot at base of mandible, narrow short band along inner eye margin near clypeus, three small spots between antennal sockets (nearly united in two specimens), antennal scape beneath, narrow short band at vertex behind eye (sometimes absent), two narrow short bands along pronotal carina on dorsal part of pronotum, narrow band at apical margin of T1 (interrupted medially) extended to lateral sides with faint small spot above yellow part laterally, three short narrow bands at apical margin of T2, stripe on outer side of protibia. Propodeal valvulae and apical margin reddish-brown. Wings dark brown, strongly infuscate, veins dark brown.
|
||
</paragraph>
|
||
<caption id="DF21E0C9FF9B2A33FF14F8E1730FF8A9" httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="2" pageNumber="177" targetBox="[150,1436,267,1781]" targetPageId="2">
|
||
<paragraph id="8BE1B041FF9B2A33FF14F8E1730FF8A9" blockId="2.[151,1436,1805,1858]" pageId="2" pageNumber="177">
|
||
<emphasis id="B92A6C53FF9B2A33FF14F8E1771BF8C8" bold="true" box="[151,308,1805,1828]" pageId="2" pageNumber="177">FIGURES 1–7</emphasis>
|
||
.
|
||
<taxonomicName id="4C5ECBC2FF9B2A33FEC3F8E17469F8C8" box="[320,582,1805,1827]" class="Insecta" family="Eumenidae" genus="Coeleumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="species" species="burmanicus">
|
||
<emphasis id="B92A6C53FF9B2A33FEC3F8E17469F8C8" box="[320,582,1805,1827]" italics="true" pageId="2" pageNumber="177">Coeleumenes burmanicus</emphasis>
|
||
</taxonomicName>
|
||
. 1
|
||
<emphasis id="B92A6C53FF9B2A33FDDCF8E27443F8C8" bold="true" box="[607,620,1806,1827]" pageId="2" pageNumber="177">–</emphasis>
|
||
5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6
|
||
<emphasis id="B92A6C53FF9B2A33FCD9F8C17549F8A9" bold="true" box="[858,870,1837,1858]" pageId="2" pageNumber="177">–</emphasis>
|
||
7, ♂. 6. Head, frontal view, 7. Left antenna.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE1B041FF9B2A32FF44F88672DEFEAB" blockId="2.[151,1436,1897,2030]" lastBlockId="3.[151,1437,151,645]" lastPageId="3" lastPageNumber="178" pageId="2" pageNumber="177">
|
||
<emphasis id="B92A6C53FF9B2A33FF44F886772EF869" box="[199,257,1898,1922]" italics="true" pageId="2" pageNumber="177">Male</emphasis>
|
||
(n=1). Body length
|
||
<quantity id="4CA61DA4FF9B2A33FE6DF8857477F86A" box="[494,600,1897,1922]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="177" unit="mm" value="10.5">10.5 mm</quantity>
|
||
; fore wing length
|
||
<quantity id="4CA61DA4FF9B2A33FCADF88575A6F86A" box="[814,905,1897,1922]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="177" unit="mm" value="9.5">9.5 mm</quantity>
|
||
. Structures as in female, except follows: Area around occelli slightly swollen. Vertex without cephalic fovea. Distance from posterior ocelli to apical margin of vertex slightly more than 4 × distance from posterior ocelli to inner eye margin. Inner eye margins in frontal view 1.3 × further apart from each other at vertex than at clypeus. Clypeus in frontal view 1.1 × as high as wide, apical margin deeply emarginated medially, forming sharp triangle tooth on each lateral side (
|
||
<figureCitation id="1365ACC4FF9A2A32FB48FF7B7337FF5B" box="[1227,1304,151,176]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="3" pageNumber="178">Fig. 7</figureCitation>
|
||
), width of emargination nearly 1/2 width of clypeus between inner eye margins (
|
||
<figureCitation id="1365ACC4FF9A2A32FC3EFF50722BFF3F" box="[957,1028,188,213]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="3" pageNumber="178">Fig. 7</figureCitation>
|
||
). Mandible with four distinct teeth, first tooth with inner side sharply emarginated medially, forming two small triangle shape on each side of the tooth. Scape 3 × as long as its maximum width; F1 2 × as long as its maximum width, F2–9 longer than its width, F10 small, as long as wide, F11 slightly curve, slightly less than 2.5 × as long as its basal width (
|
||
<figureCitation id="1365ACC4FF9A2A32FB1CFECB72CCFEAB" box="[1183,1251,295,320]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="3" pageNumber="178">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE1B041FF9A2A32FF44FEA07723FE46" blockId="3.[151,1437,151,645]" pageId="3" pageNumber="178">
|
||
<emphasis id="B92A6C53FF9A2A32FF44FEA07728FE8F" box="[199,263,332,356]" italics="true" pageId="3" pageNumber="178">Color</emphasis>
|
||
. As in female, and longitudinal wide medial band of clypeus, spot at apical margin of femur and short strip at outer side of middle tibia yellow, mark on lateral margin of tegula brown; yellow band at apical margin of T1 absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C344E3CAFF9A2A32FF44FE5B7267FDB4" pageId="3" pageNumber="178" type="discussion">
|
||
<paragraph id="8BE1B041FF9A2A32FF44FE5B7267FDB4" blockId="3.[151,1437,151,645]" pageId="3" pageNumber="178">
|
||
<emphasis id="B92A6C53FF9A2A32FF44FE5B7714FE3B" bold="true" box="[199,315,439,464]" pageId="3" pageNumber="178">Remarks.</emphasis>
|
||
Symbiotic associations with mites were observed in four of seven specimens of this species. The mites usually occupy the posterior face of propodeum (
|
||
<figureCitation id="1365ACC4FF9A2A32FC81FE307569FE1E" box="[770,838,476,501]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1805,1828]" captionTargetBox="[150,1436,267,1781]" captionTargetId="figure@2.[150,1436,267,1781]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Coeleumenes burmanicus. 1 – 5, ♀. 1. Head, frontal view, 2. Vertex, dorsal view, 3. Propodeum and metasoma, dorsal view, 4. Metasoma, lateral view, 5. Propodeum with mites, ♂ 6 – 7, ♂. 6. Head, frontal view, 7. Left antenna." httpUri="https://zenodo.org/record/265567/files/figure.png" pageId="3" pageNumber="178">Fig. 5</figureCitation>
|
||
), sometimes at holes near metanotum and around the basal slit of the metasomal segment. This phenomenon is also reported by
|
||
<bibRefCitation id="EFCFCDB0FF9A2A32FC70FDEC7301FDF3" author="Kumar" box="[1011,1326,512,536]" pageId="3" pageNumber="178" refString="Kumar, G. & Carpenter, J. M. (2013) A taxonomic review of the genus Antodynerus de Saussure, 1855 (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent. Zootaxa, 3731 (2), 267 - 278. http: // dx. doi. org / 10.11646 / zootaxa. 3731.2.7" type="journal article" year="2013">Kumar & Carpenter (2013)</bibRefCitation>
|
||
, in potter wasp species of the genus
|
||
<taxonomicName id="4C5ECBC2FF9A2A32FE44FDC87501FDD6" authority="de Saussure, 1855" authorityName="de Saussure" authorityYear="1855" box="[455,814,548,573]" class="Insecta" family="Eumenidae" genus="Antodynerus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A6C53FF9A2A32FE44FDC8747BFDD7" box="[455,596,548,572]" italics="true" pageId="3" pageNumber="178">Antodynerus</emphasis>
|
||
de Saussure, 1855
|
||
</taxonomicName>
|
||
. According to
|
||
<bibRefCitation id="EFCFCDB0FF9A2A32FC5BFDC87328FDD7" author="Okabe" box="[984,1287,548,573]" pageId="3" pageNumber="178" refString="Okabe, K. & Makino, S. (2003) Life history of Kurosaia jiju (Acari: Winterschmidtiidae) symbiotic with a mason wasp, Anterhynchium flavomarginatum micado (Hymenoptera: Eumenidae). Annals of the Entomological Society of America, 96, 652 - 659. http: // dx. doi. org / 10.1603 / 0013 - 8746 (2003) 096 [0652: LHOKJA] 2.0. CO; 2" type="journal article" year="2003">Okabe and Makino (2003)</bibRefCitation>
|
||
, the mites of the family
|
||
<taxonomicName id="4C5ECBC2FF9A2A32FE93FDAB746BFD8B" authority="Acari" authorityName="Acari" box="[272,580,583,608]" class="Arachnida" family="Winterschmidtiidae" kingdom="Animalia" order="Astigmata" pageId="3" pageNumber="178" phylum="Arthropoda" rank="family">Winterschmidtiidae (Acari)</taxonomicName>
|
||
are known to be symbiotic with potter wasps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C344E3CAFF9A2A32FF44FD807590FD6E" box="[199,959,620,645]" pageId="3" pageNumber="178" type="distribution">
|
||
<paragraph id="8BE1B041FF9A2A32FF44FD807590FD6E" blockId="3.[151,1437,151,645]" box="[199,959,620,645]" pageId="3" pageNumber="178">
|
||
<emphasis id="B92A6C53FF9A2A32FF44FD80774FFD6E" bold="true" box="[199,352,620,645]" pageId="3" pageNumber="178">Distribution.</emphasis>
|
||
<collectingCountry id="F349F0D1FF9A2A32FEE4FD80778BFD6E" box="[359,420,620,645]" name="India" pageId="3" pageNumber="178">India</collectingCountry>
|
||
;
|
||
<collectingCountry id="F349F0D1FF9A2A32FE2CFD817430FD6E" box="[431,543,621,645]" name="Myanmar" pageId="3" pageNumber="178">Myanmar</collectingCountry>
|
||
;
|
||
<collectingCountry id="F349F0D1FF9A2A32FDA8FD8074BDFD6E" box="[555,658,620,645]" name="Thailand" pageId="3" pageNumber="178">Thailand</collectingCountry>
|
||
;
|
||
<collectingCountry id="F349F0D1FF9A2A32FD1EFD8174F8FD6F" box="[669,727,621,644]" name="Laos" pageId="3" pageNumber="178">Laos</collectingCountry>
|
||
;
|
||
<collectingCountry id="F349F0D1FF9A2A32FD60FD80756AFD6E" box="[739,837,620,645]" name="Vietnam" pageId="3" pageNumber="178">Vietnam</collectingCountry>
|
||
;
|
||
<collectingCountry id="F349F0D1FF9A2A32FCD3FD807595FD6E" box="[848,954,620,645]" name="Malaysia" pageId="3" pageNumber="178">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |