treatments-xml/data/03/E4/E9/03E4E904CE60EC557EEEF985C4BFCACE.xml
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<mods:title id="8B6649822FE70F0D42455AF112ACEFD4">Chondrinidae taxonomy revisited: New synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata) 2539</mods:title>
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<mods:namePart id="14F2D39F0063DB83E3E220B8DF72CF2A">Kokshoorn, Bas</mods:namePart>
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<heading id="D0BAEF7ECE60EC4A7EEEF985C483CE00" bold="true" box="[151,898,1566,1592]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<taxonomicName id="4C4D2391CE60EC4A7EEEF985C5BECE00" ID-CoL="5DDYH" authority="Gittenberger, 1973" authorityName="Gittenberger" authorityYear="1973" box="[151,703,1566,1592]" class="Gastropoda" family="Chondrinidae" genus="Abida" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="subSpecies" species="secale" subSpecies="brongersmai">
<emphasis id="B9398400CE60EC4A7EEEF985C6CECE00" bold="true" box="[151,463,1566,1592]" italics="true" pageId="15" pageNumber="16">Abida secale brongersmai</emphasis>
<bibRefCitation id="EFDC25E3CE60EC4A7FAEF985C5BECE00" author="Gittenberger, E." box="[471,703,1566,1592]" pageId="15" pageNumber="16" pagination="1 - 267" refId="ref34929" refString="Gittenberger, E. (1973) Beitrage zur Kenntnis der Pupillacea III. Chondrininae. Zoologische Verhandelingen, 127, 1 - 267." type="journal article" year="1973">Gittenberger, 1973</bibRefCitation>
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(pl. 4 figs AE)
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<treatmentCitation id="0AEC7E03CE60EC4A7EEEF9DAC5B5CE60" author="Gittenberger, E." box="[151,692,1601,1624]" page="115" pageId="15" pageNumber="16" year="1973">
<taxonomicName id="4C4D2391CE60EC4A7EEEF9DAC5B5CE60" ID-CoL="5DDYH" authority="Gittenberger, 1973: 115" authorityName="Gittenberger" authorityPageNumber="115" authorityYear="1973" box="[151,692,1601,1624]" class="Gastropoda" family="Chondrinidae" genus="Abida" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="subSpecies" species="secale" subSpecies="brongersmai">
<emphasis id="B9398400CE60EC4A7EEEF9DAC6ACCE60" box="[151,429,1601,1624]" italics="true" pageId="15" pageNumber="16">Abida secale brongersmai</emphasis>
<bibRefCitation id="EFDC25E3CE60EC4A7FCDF9DAC5B5CE60" author="Gittenberger, E." box="[436,692,1601,1624]" pageId="15" pageNumber="16" pagination="1 - 267" refId="ref34929" refString="Gittenberger, E. (1973) Beitrage zur Kenntnis der Pupillacea III. Chondrininae. Zoologische Verhandelingen, 127, 1 - 267." type="journal article" year="1973">Gittenberger, 1973: 115</bibRefCitation>
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.
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Type locality:
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, Lérida, road to Alás, near Torres, right side of the river Segre, east of La Seu dUrgell.
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<materialsCitation id="3B25524FCE60EC4A7EBFF935C31ECEF0" box="[198,1055,1710,1736]" collectionCode="RMNH" location="National Museum of Natural History" pageId="15" pageNumber="16" specimenCount="1" typeStatus="holotype">
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:
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(
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), Leiden 54778.
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<paragraph id="8BF25812CE60EC4A7EBFF94EC6EECED7" blockId="15.[151,1436,1566,2007]" box="[198,495,1749,1775]" pageId="15" pageNumber="16">
Distribution.—See
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.
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Description (after
<bibRefCitation id="EFDC25E3CE60EC4A7FE7F967C572CF2E" author="Gittenberger, E." box="[414,627,1788,1814]" pageId="15" pageNumber="16" pagination="1 - 267" refId="ref34929" refString="Gittenberger, E. (1973) Beitrage zur Kenntnis der Pupillacea III. Chondrininae. Zoologische Verhandelingen, 127, 1 - 267." type="journal article" year="1973">Gittenberger 1973</bibRefCitation>
).—Shell slender cylindro-conical to fusiform, with 9¼11¼ weakly to moderately inflated whorls, sculptured with regularly placed, fine axial ribs. Initial teleoconch whorls often without sculpture. Body whorl obliquely flattened, slightly narrowed, with a distinct keel. Palatalis superior accompanied by a slight to moderately prominent indentation, infrapalatalis with conspicuous external indentation just behind apertural lip. Palatal side of the aperture vertical or slightly leaning backwards (in lateral view). Umbilicus moderately wide, circular. Shell height, 8.0
<quantity id="4CB5F5F7CE60EC4A7D08F826C4DDCFEF" box="[881,988,1981,2007]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="15" pageNumber="16" unit="mm" value="11.2">11.2 mm</quantity>
; width,
<quantity id="4CB5F5F7CE60EC4A7A40F826C3CBCFEF" box="[1081,1226,1981,2007]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="15" pageNumber="16" unit="mm" value="3.05" valueMax="3.3" valueMin="2.8">2.83.3 mm</quantity>
.
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Aperture roundish. Apertural lip strongly thickened, continuous across the parietal side, often protruding up to
<quantity id="4CB5F5F7CE7FEC557EA5FF25C624C8E0" box="[220,293,190,216]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="mm" value="1.0">1 mm</quantity>
. Angularis relatively high near the aperture but quickly decreasing in prominence, reaching past the front end of the parietalis, not or more or less clearly connected with spiralis. Subangularis well developed, not connected to angularis. Parietalis and infraparietalis well developed. Several weak to strong folds may be present on the apertural lip. Columellaris ending abruptly on the columella, clearly broader than the other lamellae there. Infracolumellaris, reaching further beyond the columella, often connected with a lamella on the apertural lip. Infrapalatalis, palatalis inferior and palatalis superior indented, reaching up to the edge of the aperture. At the anterodorsal palatal center a suturalis, an equally long or longer suprapalatalis and often a short basalis are present.
</paragraph>
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Genetic barcode. —GenBank accession numbers
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,
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.
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<paragraph id="8BF25812CE7FEC557EBFFD81C4BFCACE" blockId="16.[151,1436,152,758]" pageId="16" pageNumber="17">
Notes.—Both shell sculpture and apertural protrusion vary in this conspicuous subspecies. This variation is partly correlated with altitude (
<bibRefCitation id="EFDC25E3CE7FEC557C59FDDAC452CA63" author="Kokshoorn, B." box="[544,851,577,603]" pageId="16" pageNumber="17" refId="ref35737" refString="Kokshoorn, B. (2008) Resolving Riddles and Presenting New Puzzles in Chondrinidae Phylogenetics. PhD thesis, Leiden University, 192 pp." type="book" year="2008">Kokshoorn 2008: 99114</bibRefCitation>
). In the easternmost part of its present range
<emphasis id="B9398400CE7FEC557B11FDDAC62DCAB9" italics="true" pageId="16" pageNumber="17">A. s. brongersmai</emphasis>
occurs close to
<emphasis id="B9398400CE7FEC557F91FDF3C5ADCAB9" box="[488,684,616,641]" italics="true" pageId="16" pageNumber="17">A. s. margaridae</emphasis>
, but intermediate populations are not known. This might be due to the introduction of
<emphasis id="B9398400CE7FEC557FE2FD14C56CCA90" box="[411,621,655,680]" italics="true" pageId="16" pageNumber="17">A. s. brongersmai</emphasis>
in the area north of Pedra (
<collectionCode id="ED5CC0D7CE7FEC557DCCFD15C4D9CA90" box="[949,984,654,680]" pageId="16" pageNumber="17">SE</collectionCode>
of Bellver de Cerdanya). In that area, limestone rocks have been used for the construction of walls, creating an artificial habitat for
<taxonomicName id="4C4D2391CE7FEC557AADFD2EC23DCAF6" authorityName="Draparnaud" authorityYear="1801" box="[1236,1340,693,718]" class="Gastropoda" family="Chondrinidae" genus="Abida" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="secale">
<emphasis id="B9398400CE7FEC557AADFD2EC23DCAF6" box="[1236,1340,693,718]" italics="true" pageId="16" pageNumber="17">A. secale</emphasis>
</taxonomicName>
. Maybe the rocks originated from the region where
<emphasis id="B9398400CE7FEC557CEAFD47C462CACD" box="[659,867,732,757]" italics="true" pageId="16" pageNumber="17">A. s. brongersmai</emphasis>
occurs.
</paragraph>
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