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<document id="5E2731F5F04B67B0DF9866A9AA585D49" ID-DOI="10.5281/zenodo.5354105" ID-ISSN="1942-1354" ID-Zenodo-Dep="5354105" ID-ZooBank="18BA3511-66C2-4EF9-AE74-040A88E15BC3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630213090854" checkinUser="felipe" docAuthor="Halstead, David G. H." docDate="2020" docId="03CDA22316478154FF39CD8BFE971F4C" docLanguage="en" docName="0842.pdf" docOrigin="Insecta Mundi 2020 (842)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Synobius Sharp 1899" docType="treatment" docVersion="4" lastPageNumber="3" masterDocId="FFF4DA5B16448150FFF9C97FFFDD1D09" masterDocTitle="New and little known Coleoptera (Silvanidae: Silvaninae) from Central and South America" masterLastPageNumber="37" masterPageNumber="1" pageNumber="2" updateTime="1699188192110" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="75D32D039718D6C58D303D0D81D94403">New and little known Coleoptera (Silvanidae: Silvaninae) from Central and South America</mods:title>
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<mods:namePart id="C1623A8209F94C307A1C694AC345B389">Halstead, David G. H.</mods:namePart>
<mods:affiliation id="70365D830DEC28DE63F7CF3FBAB1E193">57 Meadow Way, Old Windsor, Berkshire, SL 4 2 NY, UK</mods:affiliation>
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<mods:title id="E087FCB87F8E8234EC08213DC55BB5DF">Insecta Mundi</mods:title>
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<mods:date id="E87AF8F3D2AA8A7BAD91314E44ECD91C">2020</mods:date>
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<mods:identifier id="41C9A53397085A12CAC87983574AEE70" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03CDA22316478154FF39CD8BFE971F4C" ID-DOI="http://doi.org/10.5281/zenodo.5452321" ID-Zenodo-Dep="5452321" LSID="urn:lsid:plazi:treatment:03CDA22316478154FF39CD8BFE971F4C" httpUri="http://treatment.plazi.org/id/03CDA22316478154FF39CD8BFE971F4C" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<subSubSection id="C37E40BE16478153FF39CD8BFE59181B" box="[192,388,1268,1298]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8BDB133516478153FF39CD8BFE59181B" blockId="3.[192,388,1268,1298]" box="[192,388,1268,1298]" pageId="3" pageNumber="2">
<heading id="D093A45916478153FF39CD8BFE59181B" box="[192,388,1268,1298]" fontSize="12" level="2" pageId="3" pageNumber="2" reason="2">
<taxonomicName id="4C6468B616478153FF39CD8BFE59181B" authority="Sharp" authorityName="Sharp" authorityYear="1899" box="[192,388,1268,1298]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FF39CD8BFE59181B" bold="true" box="[192,388,1268,1298]" pageId="3" pageNumber="2">
<emphasis id="B910CF2716478153FF39CD8BFEEC1818" bold="true" box="[192,305,1268,1297]" italics="true" pageId="3" pageNumber="2">Synobius</emphasis>
Sharp
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E40BE16478153FF19CC60FE3B183E" box="[224,486,1311,1335]" pageId="3" pageNumber="2" type="reference_group">
<paragraph id="8BDB133516478153FF19CC60FE3B183E" blockId="3.[224,486,1311,1335]" box="[224,486,1311,1335]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB74341B16478153FF19CC60FE3B183E" box="[224,486,1311,1335]" pageId="3" pageNumber="2">
<treatmentCitation id="0AC5352416478153FF19CC60FE3F183E" author="Sharp D." box="[224,482,1311,1335]" page="558" pageId="3" pageNumber="2" year="1899">
<taxonomicName id="4C6468B616478153FF19CC60FE3F183E" authority="Sharp 1899: 558" authorityName="Sharp" authorityPageNumber="558" authorityYear="1899" box="[224,482,1311,1335]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FF19CC60FEE5183E" box="[224,312,1311,1335]" italics="true" pageId="3" pageNumber="2">Synobius</emphasis>
<bibRefCitation id="EFF56EC416478153FEC7CC60FE3F183E" author="Sharp D." box="[318,482,1311,1335]" pageId="3" pageNumber="2" pagination="499 - 563" refId="ref21758" refString="Sharp D. 1899. Cucujidae. Biologia Centrali-Americana, Insects, Coleoptera II (1): 499 - 563." type="journal article" year="1899">Sharp 1899: 558</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E40BE16478153FF39CC38FEF71A13" pageId="3" pageNumber="2" type="discussion">
<paragraph id="8BDB133516478153FF39CC38FD4B1B25" blockId="3.[190,1442,1351,1818]" pageId="3" pageNumber="2">
<emphasis id="B910CF2716478153FF39CC38FE9A1868" bold="true" box="[192,327,1351,1377]" pageId="3" pageNumber="2">Discussion.</emphasis>
<taxonomicName id="4C6468B616478153FEB6CC38FE241868" authority="Sharp" authorityName="Sharp" authorityYear="1899" box="[335,505,1351,1377]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FEB6CC38FE6C1868" box="[335,433,1351,1377]" italics="true" pageId="3" pageNumber="2">Synobius</emphasis>
Sharp
</taxonomicName>
is a little-known genus containing two described species,
<taxonomicName id="4C6468B616478153FB72CC38FF24188A" authority="Sharp, 1899" authorityName="Sharp" authorityYear="1899" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lobicollis">
<emphasis id="B910CF2716478153FB72CC38FA8E1868" box="[1163,1363,1351,1377]" italics="true" pageId="3" pageNumber="2">Synobius lobicollis</emphasis>
<bibRefCitation id="EFF56EC416478153FAA0CC38FF24188A" author="Sharp D." pageId="3" pageNumber="2" pagination="499 - 563" refId="ref21758" refString="Sharp D. 1899. Cucujidae. Biologia Centrali-Americana, Insects, Coleoptera II (1): 499 - 563." type="journal article" year="1899">Sharp, 1899</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="F37353A516478153FE49CC15FDD1188D" box="[432,524,1386,1412]" name="Panama" pageId="3" pageNumber="2">Panama</collectingCountry>
, and
<taxonomicName id="4C6468B616478153FDB9CC16FC1D188D" authority="(Grouvelle, 1896)" baseAuthorityName="Grouvelle" baseAuthorityYear="1896" box="[576,960,1385,1412]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lobatus">
<emphasis id="B910CF2716478153FDB9CC16FD2B188A" box="[576,758,1385,1411]" italics="true" pageId="3" pageNumber="2">Synobius lobatus</emphasis>
(
<bibRefCitation id="EFF56EC416478153FCFDCC16FC65188D" author="Grouvelle A." box="[772,952,1385,1412]" pageId="3" pageNumber="2" pagination="177 - 216" refId="ref21437" refString="Grouvelle A. 1896. Nitidulides, Colydiides, Cucujides et Parnides recoltes par M. E. Gounelle au Bresil et autres Clavicornes nouveaux d'Amerique. Annales de la Societe Entomologique de France 65: 177 - 216, 17 Figures." type="book chapter" year="1896">Grouvelle, 1896</bibRefCitation>
)
</taxonomicName>
, from
<collectingCountry id="F37353A516478153FBFCCC16FB94188A" box="[1029,1097,1385,1411]" name="Brazil" pageId="3" pageNumber="2">Brazil</collectingCountry>
. Grouvelle originally described his species in
<taxonomicName id="4C6468B616478153FEA0CCF4FE1B18AC" authorityName="Lentz" authorityYear="1857" box="[345,454,1419,1445]" class="Insecta" family="Silvanidae" genus="Nausibius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FEA0CCF4FE1B18AC" box="[345,454,1419,1445]" italics="true" pageId="3" pageNumber="2">Nausibius</emphasis>
</taxonomicName>
and, although differing from this genus in general facies, it does seem to be more closely related to it than any other in the family.
<bibRefCitation id="EFF56EC416478153FD75CCD2FC9418CE" author="Grouvelle A." box="[652,841,1453,1479]" pageId="3" pageNumber="2" pagination="313 - 386" refId="ref21480" refString="Grouvelle A. 1913. Notes sur les Silvanini. Synonymies et descriptions de genres nouveaux et de nouvelles especes. Annales de la Societe Entomologique de France 81: 313 - 386." type="journal article" year="1913">Grouvelle (1913)</bibRefCitation>
transferred his species to
<taxonomicName id="4C6468B616478153FB97CCD2FB0D18CE" authorityName="Sharp" authorityYear="1899" box="[1134,1232,1453,1479]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FB97CCD2FB0D18CE" box="[1134,1232,1453,1479]" italics="true" pageId="3" pageNumber="2">Synobius</emphasis>
</taxonomicName>
remarking that an examination of the figures of
<taxonomicName id="4C6468B616478153FDE9CCB0FD5418E1" authorityName="Sharp" authorityYear="1899" box="[528,649,1486,1513]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lobicollis">
<emphasis id="B910CF2716478153FDE9CCB0FDC118E0" box="[528,540,1487,1513]" italics="true" pageId="3" pageNumber="2">S</emphasis>
.
<emphasis id="B910CF2716478153FDD3CCB1FD5418E1" box="[554,649,1486,1512]" italics="true" pageId="3" pageNumber="2">lobicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6468B616478153FD3BCCB0FCF018E1" baseAuthorityName="Grouvelle" baseAuthorityYear="1896" box="[706,813,1486,1513]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lobatus">
<emphasis id="B910CF2716478153FD3BCCB0FD1218E0" box="[706,719,1487,1513]" italics="true" pageId="3" pageNumber="2">S</emphasis>
.
<emphasis id="B910CF2716478153FD27CCB1FCF018E1" box="[734,813,1486,1512]" italics="true" pageId="3" pageNumber="2">lobatus</emphasis>
</taxonomicName>
showed these species were very closely related. Indeed, one of the labels that he attached to his
<typeStatus id="54DFAD9716478153FD8CCC8FFD0A1B03" box="[629,727,1520,1546]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
following the establishment of the new genus by Sharp, suggests he thought that they might be conspecific.
</paragraph>
<paragraph id="8BDB133516478153FEF9CF4BFEF71A13" blockId="3.[190,1442,1351,1818]" pageId="3" pageNumber="2">
Sharp, when describing the genus, remarked that it was apparently allied to
<taxonomicName id="4C6468B616478153FB9CCF4BFB0D1B47" authorityName="Lentz" authorityYear="1857" box="[1125,1232,1588,1614]" class="Insecta" family="Silvanidae" genus="Nausibius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FB9CCF4BFB0D1B47" box="[1125,1232,1588,1614]" italics="true" pageId="3" pageNumber="2">Nausibius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6468B616478153FB27CF4BFA7D1B47" authority="Reiche" authorityName="Reiche" authorityYear="1854" box="[1246,1440,1588,1614]" class="Insecta" family="Silvanidae" genus="Cathartus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FB27CF4BFA911B47" box="[1246,1356,1588,1614]" italics="true" pageId="3" pageNumber="2">Cathartus</emphasis>
Reiche
</taxonomicName>
and
<taxonomicName id="4C6468B616478153FF08CF29FE791B79" authority="Reitter" authorityName="Reitter" authorityYear="1877" box="[241,420,1622,1648]" class="Insecta" family="Erotylidae" genus="Hapalips" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FF08CF29FE8F1B79" box="[241,338,1622,1648]" italics="true" pageId="3" pageNumber="2">Hapalips</emphasis>
Reitter
</taxonomicName>
(the last of these is now placed in the
<taxonomicName id="4C6468B616478153FCA9CF29FC151B78" authorityName="Latreille" authorityYear="1802" box="[848,968,1622,1649]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="family">Erotylidae</taxonomicName>
). He stated that ‘… the greatest peculiarity of the genus is the peculiar, glabrous, rigid antennae.’ and drew attention to the fact that the structure of the sterna is that normal in the ‘Silvanides’. In his accompanying species description, Sharp anticipated a myrmecophilous or termitophilous habit for this insect. The habitat data on the majority of specimens examined (see descriptions below) tends to confirm an inquiline relationship as they were found in nests of bees of the genus
<taxonomicName id="4C6468B616478153FAF9CFA1FEFB1A13" authority="Jurine" authorityName="Jurine" authorityYear="1807" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FAF9CFA1FA8B1BFE" box="[1280,1366,1758,1783]" italics="true" pageId="3" pageNumber="2">Trigona</emphasis>
Jurine (Apidae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E40BE16478154FF39CE57FE971F4C" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8BDB133516478154FF39CE57FC291C5E" blockId="3.[192,1442,1832,1927]" lastBlockId="4.[191,1441,214,581]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<emphasis id="B910CF2716478153FF39CE57FEE11A4B" bold="true" box="[192,316,1832,1858]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
This genus (
<figureCitation id="135F0FB016478153FE36CE57FDE11A4A" box="[463,572,1832,1859]" captionStart-0="Figures 1–6" captionStart-1="Figures 7–10" captionStart-2="Figures 11–17" captionStartId-0="6.[192,279,1554,1580]" captionStartId-1="7.[387,476,1092,1118]" captionStartId-2="8.[191,278,1384,1410]" captionTargetBox-0="[192,1440,172,1462]" captionTargetBox-1="[402,1212,236,1055]" captionTargetBox-2="[195,1422,226,1343]" captionTargetId-0="figure-116@6.[192,1440,161,1611]" captionTargetId-1="figure-326@7.[385,1248,176,1217]" captionTargetId-2="figure-209@8.[107,1445,176,1413]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figures 1–6. Synobius lobatus (Grouvelle), male. 1) Habitus. 2) Metatarsus. 3) Antenna, enlarged (1–3 holotype). 4–6) Genitalia. 4–5) Parameres and median lobes to show variation. 4) Narrower median lobe, specimen from Brazil, Est. Paraná, Rio Negro. 5) Broader median lobe, holotype specimen from Brazil, Rio Grande do Sol. 6) Paramere, enlarged apical region. Scale lines for Figure 1 = 2.0 mm, for Figures 4 and 5 = 0.2 mm." captionText-1="Figures 7–10. Synobius spp. pronota. 7–8) S. lobatus (Grouvelle). 7) Male, 8) Female. 9–10) S. lobicollis Sharp, both male. Scale line = 2.0 mm." captionText-2="Figures 11–17. Synobius lobicollis Sharp, male. 11) Habitus. 12) Head, anterior lateral region showing antennal base and eye. 13) Antenna, enlarged. 14–17) Genitalia and associated sternites. 14) Genitalia, without internal sac. 15) Paramere, enlargement of apical half. 16) Internal sac, apical third. 17) Sternites 8 and 9. Scale lines for Figure 11 = 2.0 mm, for Figures 14 and 16–17 = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5354107" figureDoi-1="http://doi.org/10.5281/zenodo.5354109" figureDoi-2="http://doi.org/10.5281/zenodo.5354111" httpUri-0="https://zenodo.org/record/5354107/files/figure.png" httpUri-1="https://zenodo.org/record/5354109/files/figure.png" httpUri-2="https://zenodo.org/record/5354111/files/figure.png" pageId="3" pageNumber="2">Fig. 1–17</figureCitation>
,
<figureCitation id="135F0FB016478153FDB3CE56FDA81A4A" box="[586,629,1833,1859]" captionStart="Figures 123–124" captionStartId="36.[192,279,1900,1926]" captionTargetBox="[213,1426,701,1865]" captionTargetId="figure-179@36.[192,1440,696,1869]" captionTargetPageId="36" captionText="Figures 123–124. Color habitus photographs.123) Synobius lobatus (Grouvelle). 124) Pensus hirtus, new species." figureDoi="http://doi.org/10.5281/zenodo.5354145" httpUri="https://zenodo.org/record/5354145/files/figure.png" pageId="3" pageNumber="2">123</figureCitation>
), as represented by the two known species, may be distinguished from
<taxonomicName id="4C6468B616478153FF39CE35FEF01A6D" authorityName="Lentz" authorityYear="1857" box="[192,301,1866,1892]" class="Insecta" family="Silvanidae" genus="Nausibius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716478153FF39CE35FEF01A6D" box="[192,301,1866,1892]" italics="true" pageId="3" pageNumber="2">Nausibius</emphasis>
</taxonomicName>
by its antennae, which have a very small, narrow terminal segment; the pronotum, which is without lateral teeth, has a strong indentation before the anterior angles that form rounded and in front slightly declivous lobes; and the elytra, which lack obvious striae (an unusual condition in
<taxonomicName id="4C6468B616408154FC09C9A9FB801DF9" authorityName="Lentz" authorityYear="1857" box="[1008,1117,214,240]" class="Insecta" family="Silvanidae" genus="Nausibius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716408154FC09C9A9FB801DF9" box="[1008,1117,214,240]" italics="true" pageId="4" pageNumber="3">Nausibius</emphasis>
</taxonomicName>
although it does occur). The tarsi in
<taxonomicName id="4C6468B616408154FEE3C987FEA11C1B" authorityName="Sharp" authorityYear="1899" box="[282,380,248,274]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716408154FEE3C987FEA11C1B" box="[282,380,248,274]" italics="true" pageId="4" pageNumber="3">Synobius</emphasis>
</taxonomicName>
are simple (
<figureCitation id="135F0FB016408154FDFCC987FD951C1B" box="[517,584,248,274]" captionStart="Figures 1–6" captionStartId="6.[192,279,1554,1580]" captionTargetBox="[192,1440,172,1462]" captionTargetId="figure-116@6.[192,1440,161,1611]" captionTargetPageId="6" captionText="Figures 1–6. Synobius lobatus (Grouvelle), male. 1) Habitus. 2) Metatarsus. 3) Antenna, enlarged (1–3 holotype). 4–6) Genitalia. 4–5) Parameres and median lobes to show variation. 4) Narrower median lobe, specimen from Brazil, Est. Paraná, Rio Negro. 5) Broader median lobe, holotype specimen from Brazil, Rio Grande do Sol. 6) Paramere, enlarged apical region. Scale lines for Figure 1 = 2.0 mm, for Figures 4 and 5 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5354107" httpUri="https://zenodo.org/record/5354107/files/figure.png" pageId="4" pageNumber="3">Fig. 2</figureCitation>
) and there are no antennal grooves on the head. Obvious secondary sexual characters have not been found although with larger individuals, more transverse pronota may be indicative of females, as is often the case in
<taxonomicName id="4C6468B616408154FDEBC843FDA21C5F" authorityName="Lentz" authorityYear="1857" box="[530,639,316,342]" class="Insecta" family="Silvanidae" genus="Nausibius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B910CF2716408154FDEBC843FDA21C5F" box="[530,639,316,342]" italics="true" pageId="4" pageNumber="3">Nausibius</emphasis>
</taxonomicName>
and some other silvanine genera.
</paragraph>
<paragraph id="8BDB133516408154FEF9C821FE971F4C" blockId="4.[191,1441,214,581]" pageId="4" pageNumber="3">
Apart from the
<typeStatus id="54DFAD9716408154FE4FC820FE2C1C70" box="[438,497,351,377]" pageId="4" pageNumber="3">types</typeStatus>
in the MNHN (
<taxonomicName id="4C6468B616408154FD48C821FCC91C71" box="[689,788,350,377]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lobatu">
<emphasis id="B910CF2716408154FD48C821FD631C71" box="[689,702,350,376]" italics="true" pageId="4" pageNumber="3">S</emphasis>
.
<emphasis id="B910CF2716408154FD37C821FCC91C71" box="[718,788,350,376]" italics="true" pageId="4" pageNumber="3">lobatu</emphasis>
</taxonomicName>
s) and NHML (
<taxonomicName id="4C6468B616408154FC30C821FB991C71" authorityName="Sharp" authorityYear="1899" box="[969,1092,350,377]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lobicollis">
<emphasis id="B910CF2716408154FC30C821FC0B1C71" box="[969,982,350,376]" italics="true" pageId="4" pageNumber="3">S</emphasis>
.
<emphasis id="B910CF2716408154FC1CC821FB991C71" box="[997,1092,350,376]" italics="true" pageId="4" pageNumber="3">lobicollis</emphasis>
</taxonomicName>
) the species are known to the author only from a small number of specimens from Nevermann’s collection made in the 1930s and now mainly in the USNM, but also NHML and MIZP (just
<taxonomicName id="4C6468B616408154FD21C8DCFC8C1CB5" authorityName="Sharp" authorityYear="1899" box="[728,849,418,445]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lobicollis">
<emphasis id="B910CF2716408154FD21C8DCFD391CB4" box="[728,740,419,445]" italics="true" pageId="4" pageNumber="3">S</emphasis>
.
<emphasis id="B910CF2716408154FD0BC8DDFC8C1CB5" box="[754,849,418,444]" italics="true" pageId="4" pageNumber="3">lobicollis</emphasis>
</taxonomicName>
in these last two depositories). Nevermann believed that his material represented two new species and labelled the specimens as such but did not publish descriptions. Comparison of Nevermann’s specimens with
<taxonomicName id="4C6468B616408154FD03C898FCBE1F09" baseAuthorityName="Grouvelle" baseAuthorityYear="1896" box="[762,867,486,513]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lobatus">
<emphasis id="B910CF2716408154FD03C898FCDA1F08" box="[762,775,487,513]" italics="true" pageId="4" pageNumber="3">S</emphasis>
.
<emphasis id="B910CF2716408154FCEDC899FCBE1F09" box="[788,867,486,512]" italics="true" pageId="4" pageNumber="3">lobatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6468B616408154FC61C898FBCC1F09" authorityName="Sharp" authorityYear="1899" box="[920,1041,486,513]" class="Insecta" family="Silvanidae" genus="Synobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lobicollis">
<emphasis id="B910CF2716408154FC61C898FC781F08" box="[920,933,487,513]" italics="true" pageId="4" pageNumber="3">S</emphasis>
.
<emphasis id="B910CF2716408154FC4BC899FBCC1F09" box="[946,1041,486,512]" italics="true" pageId="4" pageNumber="3">lobicollis</emphasis>
</taxonomicName>
has shown that they represent these two extremely similar and very obviously closely related taxa, which are maintained here as distinct species and redescribed.
</paragraph>
</subSubSection>
</treatment>
</document>
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