172 lines
16 KiB
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172 lines
16 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3854816" ID-GBIF-Dataset="40effc56-2cc7-40ae-9592-c7baafeccef1" ID-GBIF-Taxon="163976987" ID-ISSN="1313–2970" ID-Zenodo-Dep="3854816" approvalRequired="53" approvalRequired_for_bibRefs="1" approvalRequired_for_textStreams="51" approvalRequired_for_treatments="1" checkinTime="1587997431956" checkinUser="plazi" docAuthor="Lourenco, Wilson & Duhem, Bernard" docDate="2009" docId="03CD87CC650FC134FF16FD41FBAABA8B" docLanguage="en" docName="ZK_article_2004.pdf" docOrigin="ZooKeys 14" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@18c8ecf5" docStyleName="zookeys.2008.journal_article" docTitle="Pantobuthus Lourenco & Duhem 2009, gen. n." docType="treatment" docVersion="6" lastPageId="8" lastPageNumber="45" masterDocId="FFF4FFB46508C13CFF9AFFF7FFFEBB75" masterDocTitle="Saharo-Sindian buthid scorpions; description of two new genera and species from Occidental Sahara and Afghanistan" masterLastPageNumber="54" masterPageNumber="37" pageId="7" pageNumber="44" updateTime="1636548053746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Saharo-Sindian buthid scorpions; description of two new genera and species from Occidental Sahara and Afghanistan</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lourenco, Wilson</mods:namePart>
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<mods:affiliation>Museum national d Histoire naturelle, Paris, France</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Duhem, Bernard</mods:namePart>
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<mods:affiliation>Muséum national d'Histoire naturelle, Paris ,, France</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-07-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>14</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>37</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.14.212</mods:identifier>
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<mods:identifier type="GBIF-Dataset">40effc56-2cc7-40ae-9592-c7baafeccef1</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576470</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3854816" ID-GBIF-Taxon="163976987" ID-Zenodo-Dep="3854816" LSID="urn:lsid:plazi:treatment:03CD87CC650FC134FF16FD41FBAABA8B" httpUri="http://treatment.plazi.org/id/03CD87CC650FC134FF16FD41FBAABA8B" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44">
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<subSubSection box="[140,462,694,721]" pageId="7" pageNumber="44" type="nomenclature">
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<paragraph blockId="7.[140,968,694,757]" box="[140,462,694,721]" pageId="7" pageNumber="44">
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<heading bold="true" box="[140,462,694,721]" fontSize="11" level="2" pageId="7" pageNumber="44" reason="0">
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<emphasis bold="true" box="[140,462,694,721]" pageId="7" pageNumber="44">
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Genus
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<taxonomicName authority="Lourenco & Duhem, 2009" authorityName="Lourenco & Duhem" authorityYear="2009" box="[223,374,694,721]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[223,374,694,721]" italics="true" pageId="7" pageNumber="44">Pantobuthus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[381,462,695,721]" pageId="7" pageNumber="44" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,968,730,757]" pageId="7" pageNumber="44" type="description">
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<paragraph blockId="7.[140,968,694,757]" box="[140,968,730,757]" pageId="7" pageNumber="44">
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<uri box="[140,968,730,757]" pageId="7" pageNumber="44">
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urn:lsid:zoobank.org:act:
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<uuid box="[421,968,730,757]" pageId="7" pageNumber="44">3C2F50F7-DD92-4854-8584-CBE4FF62B084</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSection pageId="7" pageNumber="44" type="multiple">
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<subSubSection pageId="7" pageNumber="44" type="diagnosis">
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<paragraph blockId="7.[140,1108,800,1672]" pageId="7" pageNumber="44">
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<emphasis bold="true" box="[140,258,800,826]" pageId="7" pageNumber="44">Diagnosis</emphasis>
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. Large scorpions, 66.3 mm in total length (excluding telson). Coloration generally pale yellow with regions of very light grey. Carapace carinae moderately marked; ‘lyra’ configuration rather incomplete; median ocular tubercle at the centre of carapace; three pairs of lateral eyes situated in a posterior position in relation to the ante-
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</paragraph>
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<paragraph blockId="7.[140,1108,800,1672]" pageId="7" pageNumber="44">
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rior edge; third pair totally vestigial. Sternum triangular, wider than long. Pedipalps very short and bulky; chela strongly globular; dentate margins on fixed and movable fingers of pedipalp chela composed of 9-10 oblique rows of granules. Outer and inner accessory granules present on both fingers, strongly marked. Pectinal tooth count 24-23; fulcra present; basal middle lamellae not dilated. Chelicerae with one basal denticle on the fixed finger; basal denticles of the movable finger strongly reduced. Metasomal segments II to IV with strong spinoid granules on ventral carinae; telson with a strongly globular vesicle; aculeus much shorter than vesicle and strongly curved; subaculear tooth absent. Legs: tarsi with two series of short spine-like setae; internal series with 5-7 setae, external reduced to 1-3 setae or totally absent; tibial spur strong on legs III and IV; pedal spurs moderate to strong on legs I to IV. Trichobothrial pattern A-β (beta), orthobothriotaxy. Trichobothrium
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<emphasis bold="true" box="[327,358,1328,1354]" pageId="7" pageNumber="44">db</emphasis>
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of chela totally displaced over the internal face of fixed finger.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="44" type="etymology">
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<paragraph blockId="7.[140,1108,800,1672]" pageId="7" pageNumber="44">Derivatio nominis: generic name refers to the multiple characteristics of buthid genera present in the new genus.</paragraph>
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</subSubSection>
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<subSubSection box="[185,676,1433,1460]" pageId="7" pageNumber="44" type="type_taxon">
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<paragraph blockId="7.[140,1108,800,1672]" box="[185,676,1433,1460]" pageId="7" pageNumber="44">
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<emphasis bold="true" box="[185,332,1433,1460]" pageId="7" pageNumber="44">
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<typeStatus box="[185,243,1434,1460]" pageId="7" pageNumber="44">Type</typeStatus>
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species
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</emphasis>
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<taxonomicName authority="Lourenco & Duhem, 2009" authorityName="Lourenco & Duhem" authorityYear="2009" box="[340,607,1434,1460]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
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<emphasis box="[340,607,1434,1460]" italics="true" pageId="7" pageNumber="44">Pantobuthus complicatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[615,676,1434,1460]" pageId="7" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44" type="discussion">
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<paragraph blockId="7.[140,1108,800,1672]" pageId="7" pageNumber="44">
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<emphasis bold="true" box="[185,507,1469,1495]" pageId="7" pageNumber="44">Affinities of the new genus.</emphasis>
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The new genus
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<taxonomicName authorityName="Lourenco & Duhem" authorityYear="2009" box="[689,822,1469,1495]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis box="[689,822,1469,1495]" italics="true" pageId="7" pageNumber="44">Pantobuthus</emphasis>
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</taxonomicName>
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shows affinities with several genera, such as
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<taxonomicName authorityName="Vachon" authorityYear="1950" box="[376,529,1504,1530]" class="Arachnida" family="Buthidae" genus="Odontobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis box="[376,529,1504,1530]" italics="true" pageId="7" pageNumber="44">Odontobuthus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Leach" authorityYear="1815" box="[548,623,1504,1530]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis box="[548,623,1504,1530]" italics="true" pageId="7" pageNumber="44">Buthus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Vachon" authorityYear="1950" box="[641,767,1504,1530]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis box="[641,767,1504,1530]" italics="true" pageId="7" pageNumber="44">Mesobuthus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Birula" authorityYear="1908" box="[785,908,1505,1530]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis box="[785,908,1505,1530]" italics="true" pageId="7" pageNumber="44">Hottentotta</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Levy, Amitai & Shulov" authorityYear="1973" box="[973,1100,1504,1530]" class="Arachnida" family="Buthidae" genus="Vachoniolus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis box="[973,1100,1504,1530]" italics="true" pageId="7" pageNumber="44">Vachoniolus</emphasis>
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</taxonomicName>
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, which are also elements of the North African and Palaearctic faunas. The moderately marked carapace carinae which form only an incomplete ‘lyra’ configuration associates the new genus to
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<taxonomicName authorityName="Birula" authorityYear="1908" box="[345,468,1611,1636]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis box="[345,468,1611,1636]" italics="true" pageId="7" pageNumber="44">Hottentotta</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Vachon" authorityYear="1950" box="[528,654,1610,1636]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis box="[528,654,1610,1636]" italics="true" pageId="7" pageNumber="44">Mesobuthus</emphasis>
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</taxonomicName>
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, but metasomal segments II to IV with strong spinoid granules on the ventral carinae, are more similar to those of the genus
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</paragraph>
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<paragraph blockId="8.[140,1108,167,510]" pageId="8" pageNumber="45">
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<taxonomicName authorityName="Vachon" authorityYear="1950" box="[140,293,167,193]" class="Arachnida" family="Buthidae" genus="Odontobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis box="[140,293,167,193]" italics="true" pageId="8" pageNumber="45">Odontobuthus</emphasis>
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</taxonomicName>
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. Th e very central position of the median ocular tubercle on the carapace and the shape of the telson with a strongly globular vesicle and a very short aculeus associates
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<taxonomicName authority="Lourenco & Duhem, 2009" authorityName="Lourenco & Duhem" authorityYear="2009" box="[254,387,237,263]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[254,387,237,263]" italics="true" pageId="8" pageNumber="45">Pantobuthus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[396,474,238,264]" pageId="8" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
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with
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<taxonomicName authorityName="Leach" authorityYear="1815" box="[544,619,237,263]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis box="[544,619,237,263]" italics="true" pageId="8" pageNumber="45">Buthus</emphasis>
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</taxonomicName>
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. In this last genus, however, both carapace and mesosomal carinae are strongly developed and the ‘lyra’ configuration is totally
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</paragraph>
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<paragraph blockId="8.[140,1108,167,510]" pageId="8" pageNumber="45">
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complete. Th e very short and bulky pedipalps again associate
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<taxonomicName authority="Lourenco & Duhem, 2009" authorityName="Lourenco & Duhem" authorityYear="2009" box="[831,964,307,333]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[831,964,307,333]" italics="true" pageId="8" pageNumber="45">Pantobuthus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[972,1049,308,334]" pageId="8" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
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with
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<taxonomicName authorityName="Leach" authorityYear="1815" box="[140,215,343,369]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis box="[140,215,343,369]" italics="true" pageId="8" pageNumber="45">Buthus</emphasis>
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</taxonomicName>
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. However, the very strongly globular chela recall those of the male
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<taxonomicName authorityName="Levy, Amitai & Shulov" authorityYear="1973" box="[973,1100,343,369]" class="Arachnida" family="Buthidae" genus="Vachoniolus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis box="[973,1100,343,369]" italics="true" pageId="8" pageNumber="45">Vachoniolus</emphasis>
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</taxonomicName>
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. The male of
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<taxonomicName authority="Lourenco & Duhem, 2009" authorityName="Lourenco & Duhem" authorityYear="2009" box="[277,539,378,404]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
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<emphasis box="[277,539,378,404]" italics="true" pageId="8" pageNumber="45">Pantobuthus complicatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[546,605,378,404]" pageId="8" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
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remains unknown. Finally, the total displacement of chela trichobothrium
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<emphasis bold="true" box="[490,521,413,439]" pageId="8" pageNumber="45">db</emphasis>
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to the internal surface of fixed finger, the retreated position of lateral eyes and the tarsi with two series of short spine-like setae, but with external series reduced or totally absent appear to be particular characteristics of this genus.
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</paragraph>
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</subSubSection>
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</subSection>
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</treatment>
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</document> |