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<document id="E17A02C19A5560B3F9163F2DBBE7D53D" ID-CLB-Dataset="30396" ID-DOI="10.11646/zootaxa.4402.2.2" ID-GBIF-Dataset="7a890fa5-7993-4637-8454-7dcbd6839278" ID-ISSN="1175-5326" ID-Zenodo-Dep="1208876" ID-ZooBank="2E03B764-519D-4689-9875-0AEF5C34836A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1522215825127" checkinUser="plazi" docAuthor="Dubey, Anil Kumar &amp; Martin, Jon H." docDate="2018" docId="03CD879FB6025929FF28FB4CDBA1DCE8" docLanguage="en" docName="zootaxa.4402.2.2.pdf" docOrigin="Zootaxa 4402 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tuberaleyrodes glutae (Corbett) Comb., Comb." docType="treatment" docVersion="7" lastPageNumber="265" masterDocId="FFF4FFE7B60E5927FFBFFFC4DB3FDC46" masterDocTitle="A review of the genus Tuberaleyrodes Takahashi (Hemiptera: Aleyrodidae) with description of five new species from Brunei, Hong Kong and Malaysia" masterLastPageNumber="282" masterPageNumber="251" pageNumber="263" updateTime="1698534324385" updateUser="ExternalLinkService">
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<mods:title id="460426355AD6F29DB3B391FC2C228F45">A review of the genus Tuberaleyrodes Takahashi (Hemiptera: Aleyrodidae) with description of five new species from Brunei, Hong Kong and Malaysia</mods:title>
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<mods:namePart id="CA4383B61ADB12CB9A35AE0F76DDAB41">Dubey, Anil Kumar</mods:namePart>
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<mods:namePart id="6E42E4C47E4DCC257F08853FC2434761">Martin, Jon H.</mods:namePart>
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<mods:title id="9F579EADFED0D60C19CFF51AF3FA8889">Zootaxa</mods:title>
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<mods:date id="D2D309447B65510B2600916401390C85">2018</mods:date>
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<treatment id="03CD879FB6025929FF28FB4CDBA1DCE8" ID-DOI="http://doi.org/10.5281/zenodo.6490815" ID-GBIF-Taxon="142657703" ID-Zenodo-Dep="6490815" LSID="urn:lsid:plazi:treatment:03CD879FB6025929FF28FB4CDBA1DCE8" httpUri="http://treatment.plazi.org/id/03CD879FB6025929FF28FB4CDBA1DCE8" lastPageId="14" lastPageNumber="265" pageId="12" pageNumber="263">
<subSubSection id="C37E6502B602592BFF28FB4CDA18D885" pageId="12" pageNumber="263" type="nomenclature">
<paragraph id="8BDB3689B602592BFF28FB4CD98AD8E4" blockId="12.[151,693,1160,1219]" box="[151,693,1160,1186]" pageId="12" pageNumber="263">
<heading id="D09381E5B602592BFF28FB4CD98AD8E4" bold="true" box="[151,693,1160,1186]" fontSize="11" level="1" pageId="12" pageNumber="263" reason="1">
<emphasis id="B910EA9BB602592BFF28FB4CD98AD8E4" bold="true" box="[151,693,1160,1186]" pageId="12" pageNumber="263">
<taxonomicName id="4C644D0AB602592BFF28FB4CD945D8E4" authority="(Corbett) Comb." authorityName="(Corbett) Comb." box="[151,634,1160,1186]" class="Insecta" family="Aleyrodidae" genus="Tuberaleyrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="glutae">
<emphasis id="B910EA9BB602592BFF28FB4CDA9DD8E4" bold="true" box="[151,418,1160,1186]" italics="true" pageId="12" pageNumber="263">Tuberaleyrodes glutae</emphasis>
(Corbett)
<taxonomicNameLabel id="A22357E0B602592BFD99FB4CD945D8E4" box="[550,634,1160,1186]" pageId="12" pageNumber="263">Comb.</taxonomicNameLabel>
</taxonomicName>
nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDB3689B602592BFF28FB6EDA18D885" blockId="12.[151,693,1160,1219]" box="[151,295,1194,1219]" pageId="12" pageNumber="263">
(
<figureCitation id="135F2A0CB602592BFF20FB6EDA20D885" box="[159,287,1194,1219]" captionStart-0="FIGURES 4750" captionStart-1="FIGURES 5160" captionStartId-0="14.[151,264,1927,1949]" captionStartId-1="15.[151,264,1906,1928]" captionTargetBox-0="[162,1436,198,1900]" captionTargetBox-1="[151,1436,186,1885]" captionTargetId-0="figure@14.[151,1436,193,1906]" captionTargetId-1="figure@15.[151,1436,171,1885]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 4750. Tuberaleyrodes glutae (Corbett), camera lucida drawings, puparium, variation, 47, shape, median tubercles, small 1 st abdominal seta,submarginal setae;48, shape, medianand abdominal submedian tubercles;vasiform orifice;49, shape, median tubercles absent, long first abdominal setae, operculum; 50, scattered large tubercles, caudal setae." captionText-1="FIGURES 5160. Tuberaleyrodes glutae (Corbett), microphotographs, puparium, dorsal and ventral views, 51, long first abdominal setae, median tubercles absent; 52, small first abdominal setae, vasiform orifice slightly elongate; 53, cephalothorax pattern of tuberculation; 54, abdomen, tubercles; 55, median tubercles; 56, vasiform orifice and caudal furrow; 57, thoracic tracheal pore; 58, submedian tubercles; 59, reduced number of median tubercles; 60, vasiform orifice." httpUri-0="https://zenodo.org/record/1208894/files/figure.png" httpUri-1="https://zenodo.org/record/1208896/files/figure.png" pageId="12" pageNumber="263">Figs 4760</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37E6502B602592BFF28FB36D962D961" pageId="12" pageNumber="263" type="reference_group">
<paragraph id="8BDB3689B602592BFF28FB36D91AD94E" blockId="12.[151,605,1266,1319]" box="[151,549,1266,1288]" pageId="12" pageNumber="263">
<treatmentCitationGroup id="AB7411A7B602592BFF28FB36D91AD94E" box="[151,549,1266,1288]" pageId="12" pageNumber="263">
<taxonomicName id="4C644D0AB602592BFF28FB36D91FD94E" ID-CoL="358RJ" authority="Corbett, 1935: 759" authorityName="Corbett. Two" authorityPageNumber="759" authorityYear="1935" box="[151,544,1266,1288]" class="Insecta" family="Aleyrodidae" genus="Dialeurodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="glutae">
<emphasis id="B910EA9BB602592BFF28FB36DA67D94E" box="[151,344,1266,1288]" italics="true" pageId="12" pageNumber="263">Dialeurodes glutae</emphasis>
<treatmentCitation id="0AC51098B602592BFEE0FB36D91FD94E" author="" box="[351,544,1266,1288]" page="759" pageId="12" pageNumber="263">
<bibRefCitation id="EFF54B78B602592BFEE0FB36DAD1D94E" box="[351,494,1266,1288]" pageId="12" pageNumber="263" refId="ref12483" refString="Corbett, G. H. (1935) Malayan Aleurodidae. Journal of the Federated Malay States Museum, 17, 722 - 852." type="journal article">Corbett, 1935</bibRefCitation>
: 759
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB3689B602592BFF28FAD5D962D961" blockId="12.[151,605,1266,1319]" box="[151,605,1297,1319]" pageId="12" pageNumber="263">
<taxonomicName id="4C644D0AB602592BFF28FAD5D914D961" authority="(Corbett) Comb." authorityName="(Corbett) Comb." box="[151,555,1297,1319]" class="Insecta" family="Aleyrodidae" genus="Tuberaleyrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="glutae">
<emphasis id="B910EA9BB602592BFF28FAD5DA46D961" box="[151,377,1297,1319]" italics="true" pageId="12" pageNumber="263">Tuberaleyrodes glutae</emphasis>
(Corbett)
<emphasis id="B910EA9BB602592BFE5DFAD5D914D961" bold="true" box="[482,555,1297,1319]" pageId="12" pageNumber="263">
<taxonomicNameLabel id="A22357E0B602592BFE5DFAD5D914D961" box="[482,555,1297,1319]" pageId="12" pageNumber="263">Comb.</taxonomicNameLabel>
</emphasis>
</taxonomicName>
<emphasis id="B910EA9BB602592BFD8EFAD5D962D961" bold="true" box="[561,605,1297,1319]" pageId="12" pageNumber="263">nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37E6502B602592AFF28FA90DA5DD94B" lastPageId="13" lastPageNumber="264" pageId="12" pageNumber="263" type="description">
<paragraph id="8BDB3689B602592BFF28FA90DE5FD9D6" blockId="12.[151,1437,1364,2037]" pageId="12" pageNumber="263">A re-description of this species is provided below. The dorsal setae are found to be variable in length considering basal elevation. Also, the median tubercles are absent, variably present or prominent on abdominal segments.</paragraph>
<paragraph id="8BDB3689B602592BFF78FA58D8F2D9BB" blockId="12.[151,1437,1364,2037]" pageId="12" pageNumber="263">
<emphasis id="B910EA9BB602592BFF78FA58DA77D9F3" bold="true" box="[199,328,1436,1461]" pageId="12" pageNumber="263">Puparium.</emphasis>
In life, yellowish; cryptic; scattered on leaf; 12 puparia per leaf; scale-like; without visible wax secretion; ovate (
<figureCitation id="135F2A0CB602592BFEE9FA7BDAE5D99E" box="[342,474,1471,1496]" captionStart-0="FIGURES 4750" captionStart-1="FIGURES 5160" captionStartId-0="14.[151,264,1927,1949]" captionStartId-1="15.[151,264,1906,1928]" captionTargetBox-0="[162,1436,198,1900]" captionTargetBox-1="[151,1436,186,1885]" captionTargetId-0="figure@14.[151,1436,193,1906]" captionTargetId-1="figure@15.[151,1436,171,1885]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 4750. Tuberaleyrodes glutae (Corbett), camera lucida drawings, puparium, variation, 47, shape, median tubercles, small 1 st abdominal seta,submarginal setae;48, shape, medianand abdominal submedian tubercles;vasiform orifice;49, shape, median tubercles absent, long first abdominal setae, operculum; 50, scattered large tubercles, caudal setae." captionText-1="FIGURES 5160. Tuberaleyrodes glutae (Corbett), microphotographs, puparium, dorsal and ventral views, 51, long first abdominal setae, median tubercles absent; 52, small first abdominal setae, vasiform orifice slightly elongate; 53, cephalothorax pattern of tuberculation; 54, abdomen, tubercles; 55, median tubercles; 56, vasiform orifice and caudal furrow; 57, thoracic tracheal pore; 58, submedian tubercles; 59, reduced number of median tubercles; 60, vasiform orifice." httpUri-0="https://zenodo.org/record/1208894/files/figure.png" httpUri-1="https://zenodo.org/record/1208896/files/figure.png" pageId="12" pageNumber="263">Figs 4752</figureCitation>
); posteriorly narrow; found on lower surface of leaves; dimorphic, female
<date id="FFDA1049B602592BFAA1FA7BDEA3D99E" box="[1310,1436,1471,1496]" pageId="12" pageNumber="263" value="1375" valueMax="1462">13751462</date>
µm long,
<date id="FFDA1049B602592BFEBBFA20DABDD9BB" box="[260,386,1508,1533]" pageId="12" pageNumber="263" value="1012" valueMax="1075">10121075</date>
wide; male
<date id="FFDA1049B602592BFDB4FA20D9B6D9BB" box="[523,649,1508,1533]" pageId="12" pageNumber="263" value="1050" valueMax="1287">10501287</date>
µm long; 755900 µm long.
</paragraph>
<paragraph id="8BDB3689B602592BFF78F9C3DA9DDACA" blockId="12.[151,1437,1364,2037]" pageId="12" pageNumber="263">
<emphasis id="B910EA9BB602592BFF78F9C3DA17DA66" bold="true" box="[199,296,1543,1568]" pageId="12" pageNumber="263">Margin.</emphasis>
Irregularly crenulate, 1423 crenulations in 0.1 mm; submarginal lines forming little projections on margin; margin modified occasionally into deep C-shaped invagination at thoracic tracheal pore area (
<figureCitation id="135F2A0CB602592BFA87F9E8DEB0DA03" box="[1336,1423,1580,1605]" captionStart="FIGURES 5160" captionStartId="15.[151,264,1906,1928]" captionTargetBox="[151,1436,186,1885]" captionTargetId="figure@15.[151,1436,171,1885]" captionTargetPageId="15" captionText="FIGURES 5160. Tuberaleyrodes glutae (Corbett), microphotographs, puparium, dorsal and ventral views, 51, long first abdominal setae, median tubercles absent; 52, small first abdominal setae, vasiform orifice slightly elongate; 53, cephalothorax pattern of tuberculation; 54, abdomen, tubercles; 55, median tubercles; 56, vasiform orifice and caudal furrow; 57, thoracic tracheal pore; 58, submedian tubercles; 59, reduced number of median tubercles; 60, vasiform orifice." httpUri="https://zenodo.org/record/1208896/files/figure.png" pageId="12" pageNumber="263">Fig. 57</figureCitation>
), sometimes and with shallow emargination, and with slight invagination at caudal tracheal pore area, but not forming C-shaped pore.
</paragraph>
<paragraph id="8BDB3689B602592AFF78F953DE3FDD5B" blockId="12.[151,1437,1364,2037]" lastBlockId="13.[151,1437,151,1977]" lastPageId="13" lastPageNumber="264" pageId="12" pageNumber="263">
<emphasis id="B910EA9BB602592BFF78F953DA10DAF6" bold="true" box="[199,303,1687,1712]" pageId="12" pageNumber="263">Dorsum.</emphasis>
Submargin not differentiated from the dorsal disc. Submarginal lines evident. Submargin with a row of 10 pairs of minute capitate setae, more clearly visible on posterior abdominal area, at least 3 pairs from segment VIVIII. Submedian area with a longitudinal row or clusters of minute tubercles near termination of segment sutures (
<figureCitation id="135F2A0CB602592BFF45F8C0DA6CDB5B" box="[250,339,1796,1821]" captionStart="FIGURES 5160" captionStartId="15.[151,264,1906,1928]" captionTargetBox="[151,1436,186,1885]" captionTargetId="figure@15.[151,1436,171,1885]" captionTargetPageId="15" captionText="FIGURES 5160. Tuberaleyrodes glutae (Corbett), microphotographs, puparium, dorsal and ventral views, 51, long first abdominal setae, median tubercles absent; 52, small first abdominal setae, vasiform orifice slightly elongate; 53, cephalothorax pattern of tuberculation; 54, abdomen, tubercles; 55, median tubercles; 56, vasiform orifice and caudal furrow; 57, thoracic tracheal pore; 58, submedian tubercles; 59, reduced number of median tubercles; 60, vasiform orifice." httpUri="https://zenodo.org/record/1208896/files/figure.png" pageId="12" pageNumber="263">Fig. 54</figureCitation>
), many with a geminate pore, often appearing as a single pore. Median tubercles present on abdominal segments IVII (
<figureCitation id="135F2A0CB602592BFE64F8E3D90FDB06" box="[475,560,1831,1856]" captionStart="FIGURES 5160" captionStartId="15.[151,264,1906,1928]" captionTargetBox="[151,1436,186,1885]" captionTargetId="figure@15.[151,1436,171,1885]" captionTargetPageId="15" captionText="FIGURES 5160. Tuberaleyrodes glutae (Corbett), microphotographs, puparium, dorsal and ventral views, 51, long first abdominal setae, median tubercles absent; 52, small first abdominal setae, vasiform orifice slightly elongate; 53, cephalothorax pattern of tuberculation; 54, abdomen, tubercles; 55, median tubercles; 56, vasiform orifice and caudal furrow; 57, thoracic tracheal pore; 58, submedian tubercles; 59, reduced number of median tubercles; 60, vasiform orifice." httpUri="https://zenodo.org/record/1208896/files/figure.png" pageId="12" pageNumber="263">Fig. 55</figureCitation>
) or entirely absent. The distance between large pore and small porette 23 times diameter of the large pore itself, 23 pairs on the first abdominal segment between setae and one pair each on abdominal segments IIVIII. Tubercle clusters present on submedian area of pro- and metathorax with a few tubercles on mesothorax (
<figureCitation id="135F2A0CB602592BFE06F850D932DBEB" box="[441,525,1940,1965]" captionStart="FIGURES 5160" captionStartId="15.[151,264,1906,1928]" captionTargetBox="[151,1436,186,1885]" captionTargetId="figure@15.[151,1436,171,1885]" captionTargetPageId="15" captionText="FIGURES 5160. Tuberaleyrodes glutae (Corbett), microphotographs, puparium, dorsal and ventral views, 51, long first abdominal setae, median tubercles absent; 52, small first abdominal setae, vasiform orifice slightly elongate; 53, cephalothorax pattern of tuberculation; 54, abdomen, tubercles; 55, median tubercles; 56, vasiform orifice and caudal furrow; 57, thoracic tracheal pore; 58, submedian tubercles; 59, reduced number of median tubercles; 60, vasiform orifice." httpUri="https://zenodo.org/record/1208896/files/figure.png" pageId="12" pageNumber="263">Fig. 53</figureCitation>
). Cephalothoracic and abdominal segment sutures reaching submedian area (
<figureCitation id="135F2A0CB602592BFAD0F850DB88DB97" captionStart="FIGURES 5160" captionStartId="15.[151,264,1906,1928]" captionTargetBox="[151,1436,186,1885]" captionTargetId="figure@15.[151,1436,171,1885]" captionTargetPageId="15" captionText="FIGURES 5160. Tuberaleyrodes glutae (Corbett), microphotographs, puparium, dorsal and ventral views, 51, long first abdominal setae, median tubercles absent; 52, small first abdominal setae, vasiform orifice slightly elongate; 53, cephalothorax pattern of tuberculation; 54, abdomen, tubercles; 55, median tubercles; 56, vasiform orifice and caudal furrow; 57, thoracic tracheal pore; 58, submedian tubercles; 59, reduced number of median tubercles; 60, vasiform orifice." httpUri="https://zenodo.org/record/1208896/files/figure.png" pageId="12" pageNumber="263">Fig. 54</figureCitation>
). Meso- and metathorax subequal in mid-length. The longest median length of abdominal segment was of segment I and the smallest of segment VII. The median length of abdominal segments IVIII measured in: female 7783, 5567, 6063, 60, 58, 53, 53 and 4042 µm long, respectively; male
<date id="FFDA1049B603592AFC46FF53DF7EDCF6" box="[1017,1089,151,176]" pageId="13" pageNumber="264" value="1962-02-05">5 262</date>
, 55, 5058, 5060, 4055, 40- 50, 4548 and 3237 µm long, respectively. Longitudinal moulting suture reaching margin and transverse moulting suture reaching submedian to inner subdorsal area. Caudal furrow length in female 190218 µm long, male 145 185 µm long. Thoracic tracheal furrow absent; caudal furrow smooth or faintly tuberculate (
<figureCitation id="135F2A0CB603592AFB20FEC0DFCCDD5B" box="[1183,1267,260,285]" captionStart="FIGURES 5160" captionStartId="15.[151,264,1906,1928]" captionTargetBox="[151,1436,186,1885]" captionTargetId="figure@15.[151,1436,171,1885]" captionTargetPageId="15" captionText="FIGURES 5160. Tuberaleyrodes glutae (Corbett), microphotographs, puparium, dorsal and ventral views, 51, long first abdominal setae, median tubercles absent; 52, small first abdominal setae, vasiform orifice slightly elongate; 53, cephalothorax pattern of tuberculation; 54, abdomen, tubercles; 55, median tubercles; 56, vasiform orifice and caudal furrow; 57, thoracic tracheal pore; 58, submedian tubercles; 59, reduced number of median tubercles; 60, vasiform orifice." httpUri="https://zenodo.org/record/1208896/files/figure.png" pageId="13" pageNumber="264">Fig. 56</figureCitation>
).
</paragraph>
<paragraph id="8BDB3689B603592AFF78FEE3D883DDEB" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="264">
<emphasis id="B910EA9BB603592AFF78FEE3DAAFDD06" bold="true" box="[199,400,295,320]" pageId="13" pageNumber="264">Vasiform orifice.</emphasis>
Subcordate to subcircular, inner lateral and posterior margins with crenulations, posteriorly slightly notched, female 5255 µm long, 5073 µm wide; male 4752 µm long, 4245 wide; operculum elongate cordate, sometimes with lateral emargination, almost covering the orifice, female 35 µm long, 3033 µm wide; male 3037 µm long, 2735 µm wide; lingula concealed by operculum.
</paragraph>
<paragraph id="8BDB3689B603592AFF78FE73DEA2DE7B" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="264">
<emphasis id="B910EA9BB603592AFF78FE73DA26DD96" bold="true" box="[199,281,439,464]" pageId="13" pageNumber="264">Venter.</emphasis>
A pair of ventral abdominal setae female 2325 µm long, 4248 µm apart; male 1725 µm long; 40 42 µm apart. Antennae inside prothoracic legs, female 100 µm (including keel 10 µm) long; male 7790 µm (including keel 3 µm) long. Microsetae present at base of meso- and metathoracic legs, 510 µm long. Stipples present as dots in the thoracic tracheal folds but very faint and in the form of small lines in the caudal tracheal fold.
</paragraph>
<paragraph id="8BDB3689B603592AFF78FD83D9C6DF39" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="264">
<emphasis id="B910EA9BB603592AFF78FD83DA6ADE26" bold="true" box="[199,341,583,608]" pageId="13" pageNumber="264">Chaetotaxy.</emphasis>
Anterior marginal setae 2025 µm long and posterior marginal setae 2532 µm long. Cephalic and first abdominal setae much longer, apically acute, placed on much elevated dorsal cuticle of the puparium; cephalic setae in female 448467 µm (basal elevation 310322 µm + apical setae 137145 µm) long; male 261 340 µm (basal elevation 192230 µm + apical setae 68105 µm) long; first abdominal setae in female 322442 µm (basal elevation 225312 µm + apical setae 98130 µm) long, male 262310 µm (basal elevation 215218 µm + apical setae 7493 µm) long; eighth abdominal setae anterolateral to vasiform orifice, slightly above the base of operculum, pointed or blunt ended, usually 1013 µm long (68 µm long in
<specimenCount id="9D62FD00B603592AFC4DFCE4DF01DF7E" box="[1010,1086,799,824]" pageId="13" pageNumber="264" type="male">2 male</specimenCount>
puparia); caudal setae pointed, located at ends of caudal tracheal pore, not on long elevation as cephalic setae, sometimes on small elevation, 67 72 µm long. Submarginal row of minute setae present.
</paragraph>
<paragraph id="8BDB3689B603592AFF78FC48DA5DD94B" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="264">
<emphasis id="B910EA9BB603592AFF78FC48DA61DFE3" bold="true" box="[199,350,908,933]" pageId="13" pageNumber="264">Third instar.</emphasis>
Oval; 800862 µm long, 575625 µm wide. Anterior and posterior marginal setae 3755 µm and 2033 µm long, respectively. Margin crenulate 2024 crenulations in 0.1 mm. Thoracic tracheal pore area modified into comb and caudal tracheal pore superficially emarginated. Cephalic setae capitate 1215 µm long, first, eighth abdominal setae, anterolateral to vasiform orifice and caudal setae 1520, 79 and 6075 µm long, respectively. Submargin with 12 pairs of minute setae, 35 µm long. Subdorsum tuberculate. Median tubercles present on abdominal segments. Median length of cephalothorax and abdomen almost equal. The median length of abdominal segments IVIII measured as: 3538, 37, 37, 3240, 2735, 2730, 3033, 2735 µm long, respectively. Vasiform orifice elongate cordate 3545 µm long, 3538 µm wide; operculum subcordate, lateral margin emarginated, 2732 µm long, 2526 µm wide. Caudal furrow 95103 µm long, 540 µm wide. Paired ventral setae 1520 µm long, 27 31 µm apart. Antennae inside prothoracic legs, 2730 µm long. Microsetae at base of pro-, meso- and metathoracic legs 35 µm long.
</paragraph>
</subSubSection>
<subSubSection id="C37E6502B603592AFF78FAD3DE4FD986" pageId="13" pageNumber="264" type="materials_examined">
<paragraph id="8BDB3689B603592AFF78FAD3DE4FD986" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="264">
<emphasis id="B910EA9BB603592AFF78FAD3DA96D976" bold="true" box="[199,425,1303,1328]" pageId="13" pageNumber="264">Material examined</emphasis>
.
<taxonomicName id="4C644D0AB603592AFE08FAD3D81ED976" authority="(Corbett)" authorityName="Corbett" box="[439,801,1303,1328]" class="Insecta" family="Aleyrodidae" genus="Tuberaleyrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="264" phylum="Arthropoda" rank="species" species="glutae">
<emphasis id="B910EA9BB603592AFE08FAD3D98ED976" box="[439,689,1303,1328]" italics="true" pageId="13" pageNumber="264">Tuberaleyrodes glutae</emphasis>
(Corbett)
</taxonomicName>
, Indonesia, Sulawesi Utara: Dumoga-Bone N.P., approx.
<quantity id="4C9C9B6CB603592AFF28FAF8DBDCD912" box="[151,227,1340,1365]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="13" pageNumber="264" unit="m" value="215.0">215 m</quantity>
. alt., Toraut forest, near base camp, 3 puparia (2♀, 1♂), 8 third instars on 2 slides, on
<taxonomicName id="4C644D0AB603592AFB75FAF8DE6BD912" box="[1226,1364,1340,1364]" family="Annonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="family">Annonaceae</taxonomicName>
(Kew Gardens opinion),
<taxonomicName id="4C644D0AB603592AFED7FAA5DA88D93E" authorityName="Loureiro" authorityYear="1790" box="[360,439,1377,1400]" class="Magnoliopsida" family="Myristicaceae" genus="Knema" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Tracheophyta" rank="genus">
<emphasis id="B910EA9BB603592AFED7FAA5DA88D93E" box="[360,439,1377,1400]" italics="true" pageId="13" pageNumber="264">Knema</emphasis>
</taxonomicName>
?
<date id="FFDA1049B603592AFE74FA9BD90AD93E" box="[459,565,1375,1400]" pageId="13" pageNumber="264" value="1985-03-04">4.iii.1985</date>
(4574); one puparium, on
<taxonomicName id="4C644D0AB603592AFCDDFAA4DFACD93E" authority=", Kew Gardens" authorityName="Kew Gardens" box="[866,1171,1375,1400]" family="Annonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="family">Annonaceae, Kew Gardens</taxonomicName>
opinion,
<date id="FFDA1049B603592AFB41FA9BDE59D93E" box="[1278,1382,1375,1400]" pageId="13" pageNumber="264" value="1985-04-04">4.iv.1985</date>
(J H Martin 4784); 2 ♀ puparia on one slide, on
<taxonomicName id="4C644D0AB603592AFD21FA41D851D9DB" authorityName="Warb" authorityYear="1897" box="[670,878,1412,1437]" class="Magnoliopsida" family="Myristicaceae" genus="Knema" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Tracheophyta" rank="species" species="tomentella">
<emphasis id="B910EA9BB603592AFD21FA41D851D9DB" box="[670,878,1412,1437]" italics="true" pageId="13" pageNumber="264">Knema tomentella</emphasis>
</taxonomicName>
,
<date id="FFDA1049B603592AFC3FFA40D8C5D9DB" box="[896,1018,1412,1437]" pageId="13" pageNumber="264" value="1985-03-26">26.iii.1985</date>
(J H Martin 4754); West Malaysia: Pahang Province, Lake Chini Basin, one male puparium on
<taxonomicName id="4C644D0AB603592AFC89FA6CD8ECD9F9" authorityName="Schaeffer" authorityYear="1760" box="[822,979,1448,1471]" class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" higherTaxonomySource="CoL" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="genus">
<emphasis id="B910EA9BB603592AFC89FA6CD8ECD9F9" box="[822,979,1448,1471]" italics="true" pageId="13" pageNumber="264">Cinnamomum</emphasis>
</taxonomicName>
sp.,
<date id="FFDA1049B603592AFBB7FA63DFA8D986" box="[1032,1175,1447,1472]" pageId="13" pageNumber="264" value="2013-01-27" valueMax="2013-01-31" valueMin="2013-01-27">27-31.i.2013</date>
(J H Martin 8667).
</paragraph>
</subSubSection>
<subSubSection id="C37E6502B603592AFF78FA08D952DA4E" pageId="13" pageNumber="264" type="biology_ecology">
<paragraph id="8BDB3689B603592AFF78FA08D952DA4E" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="264">
<emphasis id="B910EA9BB603592AFF78FA08DA68D9A3" bold="true" box="[199,343,1484,1509]" pageId="13" pageNumber="264">Host plants.</emphasis>
<taxonomicName id="4C644D0AB603592AFEDFFA08DA9FD9A3" authorityName="Linnaeus" authorityYear="1771" box="[352,416,1484,1509]" class="Magnoliopsida" family="Anacardiaceae" genus="Gluta" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="264" phylum="Tracheophyta" rank="genus">
<emphasis id="B910EA9BB603592AFEDFFA08DA9FD9A3" box="[352,416,1484,1509]" italics="true" pageId="13" pageNumber="264">Gluta</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C644D0AB603592AFE67FA08D9BED9A3" box="[472,641,1484,1509]" family="Anacardiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="family">Anacardiaceae</taxonomicName>
) (
<bibRefCitation id="EFF54B78B603592AFD27FA08D806D9A3" box="[664,825,1484,1509]" pageId="13" pageNumber="264" refId="ref12483" refString="Corbett, G. H. (1935) Malayan Aleurodidae. Journal of the Federated Malay States Museum, 17, 722 - 852." type="journal article">Corbett, 1935</bibRefCitation>
);
<taxonomicName id="4C644D0AB603592AFCF1FA08D8E3D9A2" box="[846,988,1484,1508]" family="Annonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="family">Annonaceae</taxonomicName>
;
<taxonomicName id="4C644D0AB603592AFC56FA09DFB9D9A2" authorityName="Schaeffer" authorityYear="1760" box="[1001,1158,1485,1508]" class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" higherTaxonomySource="CoL" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="genus">
<emphasis id="B910EA9BB603592AFC56FA09DFB9D9A2" box="[1001,1158,1485,1508]" italics="true" pageId="13" pageNumber="264">Cinnamomum</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C644D0AB603592AFB01FA08DE07D9A2" authorityName="" authorityYear="1789" box="[1214,1336,1484,1508]" family="Lauraceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="family">Lauraceae</taxonomicName>
);
<taxonomicName id="4C644D0AB603592AFAF3FA09DAFCDA4E" authorityName="Warb" authorityYear="1897" class="Magnoliopsida" family="Myristicaceae" genus="Knema" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Tracheophyta" rank="species" species="tomentella">
<emphasis id="B910EA9BB603592AFAF3FA09DA32DA4E" italics="true" pageId="13" pageNumber="264">Knema tomentella</emphasis>
(Myristicaceae)
</taxonomicName>
(new records).
</paragraph>
</subSubSection>
<subSubSection id="C37E6502B603592AFF78F9D0D8EDDA6B" box="[199,978,1556,1581]" pageId="13" pageNumber="264" type="distribution">
<paragraph id="8BDB3689B603592AFF78F9D0D8EDDA6B" blockId="13.[151,1437,151,1977]" box="[199,978,1556,1581]" pageId="13" pageNumber="264">
<emphasis id="B910EA9BB603592AFF78F9D0DA5FDA6B" bold="true" box="[199,352,1556,1581]" pageId="13" pageNumber="264">Distribution.</emphasis>
West
<collectingCountry id="F3737619B603592AFE1AF9D0D934DA6B" box="[421,523,1556,1581]" name="Malaysia" pageId="13" pageNumber="264">Malaysia</collectingCountry>
(
<bibRefCitation id="EFF54B78B603592AFDA4F9D0D985DA6B" box="[539,698,1556,1581]" pageId="13" pageNumber="264" refId="ref12483" refString="Corbett, G. H. (1935) Malayan Aleurodidae. Journal of the Federated Malay States Museum, 17, 722 - 852." type="journal article">Corbett, 1935</bibRefCitation>
);
<collectingRegion id="49A0F86BB603592AFD71F9D0D80CDA6B" box="[718,819,1556,1581]" country="Indonesia" name="Sulawesi Utara" pageId="13" pageNumber="264">Sulawesi</collectingRegion>
(new record).
</paragraph>
</subSubSection>
<subSubSection id="C37E6502B6035929FF78F9F3DBA1DCE8" lastPageId="14" lastPageNumber="265" pageId="13" pageNumber="264" type="discussion">
<paragraph id="8BDB3689B603592AFF78F9F3DEA2DBFF" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="264">
<emphasis id="B910EA9BB603592AFF78F9F3DA04DA16" bold="true" box="[199,315,1591,1616]" pageId="13" pageNumber="264">Remarks.</emphasis>
The puparia of this species bear tuberculate setae and agree with the generic description of
<taxonomicName id="4C644D0AB603592AFF28F998DA7BDA33" authorityName="Takahashi" authorityYear="1932" box="[151,324,1628,1653]" class="Insecta" family="Aleyrodidae" genus="Tuberaleyrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA9BB603592AFF28F998DA7BDA33" box="[151,324,1628,1653]" italics="true" pageId="13" pageNumber="264">Tuberaleyrodes</emphasis>
</taxonomicName>
, therefore, this species is transferred from
<taxonomicName id="4C644D0AB603592AFC98F998D88EDA33" authorityName="Cockerell" authorityYear="1902" box="[807,945,1628,1653]" class="Insecta" family="Aleyrodidae" genus="Dialeurodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA9BB603592AFC98F998D88EDA33" box="[807,945,1628,1653]" italics="true" pageId="13" pageNumber="264">Dialeurodes</emphasis>
</taxonomicName>
to
<taxonomicName id="4C644D0AB603592AFC6AF998DFBDDA33" authorityName="Takahashi" authorityYear="1932" box="[981,1154,1628,1653]" class="Insecta" family="Aleyrodidae" genus="Tuberaleyrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B910EA9BB603592AFC6AF998DFBDDA33" box="[981,1154,1628,1653]" italics="true" pageId="13" pageNumber="264">Tuberaleyrodes</emphasis>
</taxonomicName>
. Puparia on
<taxonomicName id="4C644D0AB603592AFAADF998DEA3DA32" box="[1298,1436,1628,1652]" family="Annonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="family">Annonaceae</taxonomicName>
from
<collectingRegion id="49A0F86BB603592AFF6AF9BBDA05DADE" box="[213,314,1663,1688]" country="Indonesia" name="Sulawesi Utara" pageId="13" pageNumber="264">Sulawesi</collectingRegion>
were cryptic and scattered on leaf. Dimorphism was also visible in the length of cephalic and first abdominal setae of male and female puparia in that the female puparia possessed longer setae. In a single puparium, the length of caudal setae varied from left
<emphasis id="B910EA9BB603592AFD4AF90DD833DAA6" box="[757,780,1737,1760]" italics="true" pageId="13" pageNumber="264">vs</emphasis>
right side (left side 35 µm
<emphasis id="B910EA9BB603592AFBF9F90DDF62DAA6" box="[1094,1117,1737,1760]" italics="true" pageId="13" pageNumber="264">vs</emphasis>
right side 57 µm) of caudal pore opening. The length of caudal furrow in female puparia was 3345 µm longer than that of male puparia. No difference could be made in the second and third instars except irregular margins. The dorsal setae are placed on tubercles in third instars, not elevated but in fourth instars (puparia) dorsal setae are in the form of tuberculate setae, hence it is concluded that the elevation of dorsal cuticle that holds apical setae takes place during transformation from third to fourth instar in this species. Minute tubercles on submedian area of abdominal segments are more prominent in puparia collected on
<taxonomicName id="4C644D0AB603592AFD51F865D846DBFF" box="[750,889,1953,1977]" family="Annonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Magnoliopsida" rank="family">Annonaceae</taxonomicName>
but faint and less in puparia from
<taxonomicName id="4C644D0AB603592AFB47F864DEA6DBFF" authorityName="" authorityYear="1810" box="[1272,1433,1952,1977]" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="13" pageNumber="264" phylum="Tracheophyta" rank="family">Myristicaceae</taxonomicName>
.
</paragraph>
<paragraph id="8BDB3689B6005929FF28FF5CDBA1DCE8" blockId="14.[151,158,152,174]" box="[151,158,152,174]" pageId="14" pageNumber="265">\</paragraph>
</subSubSection>
</treatment>
</document>