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<document id="4E647BB99BB7C20478DD4AFE12DD918B" ID-CLB-Dataset="46598" ID-DOI="10.5281/zenodo.277961" ID-GBIF-Dataset="ebeb34ec-8338-4e31-bcf7-757e5b4ce497" ID-ISSN="1175-5326" ID-Zenodo-Dep="277961" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460373518696" checkinUser="plazi" docAuthor="Arnold, Rachel J. &amp; Pietsch, Theodore W." docDate="2011" docId="03CD878CFF90FFAEFF6C3484FA924449" docLanguage="en" docName="zt02925p068.pdf" docOrigin="Zootaxa 2925" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Histiophryne maggiewalker Arnold &amp; Pietsch, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="67" masterDocId="FFF4FFF4FF91FFABFFFB3002FFD9452D" masterDocTitle="A new species of frogfish of the genus Histiophryne (Teleostei: Lophiiformes: Antennariidae) from Queensland, Australia" masterLastPageNumber="68" masterPageNumber="63" pageNumber="64" updateTime="1698296048463" updateUser="ExternalLinkService">
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<mods:title id="EA13753C973C87FDCF5408BB7E370B48">A new species of frogfish of the genus Histiophryne (Teleostei: Lophiiformes: Antennariidae) from Queensland, Australia</mods:title>
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<mods:namePart id="C77EBBDBFFA72F1E9020760DA9C9B27A">Arnold, Rachel J.</mods:namePart>
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<mods:namePart id="971E6C71401C6FF3D69AA9C0B84012FC">Pietsch, Theodore W.</mods:namePart>
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<mods:date id="F82BC6300115D615BF291F5134772699">2011</mods:date>
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<mods:identifier id="F62747E1582937B4347EF9E8C0294CC4" type="CLB-Dataset">46598</mods:identifier>
<mods:identifier id="5E91807E323FAAA1ABA28AF833BF50FE" type="DOI">10.5281/zenodo.277961</mods:identifier>
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<treatment id="03CD878CFF90FFAEFF6C3484FA924449" ID-DOI="http://doi.org/10.5281/zenodo.6188976" ID-GBIF-Taxon="119412642" ID-Zenodo-Dep="6188976" LSID="urn:lsid:zoobank.org:act:3FB75687-57D5-4162-B9D9-04D4CBC061E1" httpUri="http://treatment.plazi.org/id/03CD878CFF90FFAEFF6C3484FA924449" lastPageId="5" lastPageNumber="67" pageId="1" pageNumber="64">
<subSubSection id="C37E6511FF90FFAAFF6C3484FD93418D" box="[151,586,1158,1184]" pageId="1" pageNumber="64" type="nomenclature">
<paragraph id="8BDB369AFF90FFAAFF6C3484FD93418D" blockId="1.[151,586,1158,1184]" box="[151,586,1158,1184]" pageId="1" pageNumber="64">
<heading id="D09381F6FF90FFAAFF6C3484FD93418D" bold="true" box="[151,586,1158,1184]" fontSize="11" level="1" pageId="1" pageNumber="64" reason="1">
<emphasis id="B910EA88FF90FFAAFF6C3484FD93418D" bold="true" box="[151,586,1158,1184]" pageId="1" pageNumber="64">
<taxonomicName id="4C644D19FF90FFAAFF6C3484FE3C418D" ID-CoL="6LWH7" box="[151,485,1158,1184]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="64" phylum="Chordata" rank="species" species="maggiewalker" status="sp. nov.">
<emphasis id="B910EA88FF90FFAAFF6C3484FE3C418D" bold="true" box="[151,485,1158,1184]" italics="true" pageId="1" pageNumber="64">Histiophryne maggiewalker</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357F3FF90FFAAFE173484FD93418D" box="[492,586,1158,1184]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E6511FF90FFAAFF6C34CFFCE14058" pageId="1" pageNumber="64" type="materials_examined">
<paragraph id="8BDB369AFF90FFAAFF6C34CFFC9E4024" blockId="1.[151,1436,1229,2010]" pageId="1" pageNumber="64">
<emphasis id="B910EA88FF90FFAAFF6C34CFFE9F41CB" bold="true" box="[151,326,1229,1254]" pageId="1" pageNumber="64">
<typeStatus id="54DF8838FF90FFAAFF6C34CFFF0B41C8" box="[151,210,1229,1253]" pageId="1" pageNumber="64">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54DF8838FF90FFAAFEB534CFFE6341C8" box="[334,442,1229,1253]" pageId="1" pageNumber="64" type="holotype">Holotype</typeStatus>
: QM I.38176, 55 mm, found washed up on the beach of Alexandria Bay, Noosa Heads, southeast Queensland, K. Long,
<date id="FFDA105AFF90FFAAFDF834F3FD1A4024" box="[515,707,1265,1289]" pageId="1" pageNumber="64" value="2007-10-20">20 October 2007</date>
.
<figureCitation id="135F2A1FFF90FFAAFD3634F3FC9B4024" box="[717,834,1265,1289]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1913,1936]" captionTargetBox="[151,1436,481,1873]" captionTargetId="figure@2.[151,1436,481,1874]" captionTargetPageId="2" captionText="FIGURE 1. A) Histiophryne maggiewalker, new species, holotype, QM I. 38176, 55 mm SL, showing some dehydration, and B) Histiophryne maggiewalker, paratype, QM I. 37621, 60.1 mm SL." httpUri="https://zenodo.org/record/277962/files/figure.png" pageId="1" pageNumber="64">Figure 1A</figureCitation>
.
</paragraph>
<paragraph id="8BDB369AFF90FFAAFF3E3514FCE14058" blockId="1.[151,1436,1229,2010]" pageId="1" pageNumber="64">
<typeStatus id="54DF8838FF90FFAAFF3E3514FEEE4003" box="[197,311,1302,1326]" pageId="1" pageNumber="64" type="paratype">Paratypes</typeStatus>
: QM I.29079,
<quantity id="4C9C9B7FFF90FFAAFE173517FD804000" box="[492,601,1301,1325]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="1" pageNumber="64" unit="mm" value="16.6">16.6 mm</quantity>
, north side of Mujimba Island, Mujimba, Queensland,
<collectingCountry id="F373760AFF90FFAAFB083517FA844000" box="[1267,1373,1301,1325]" name="Australia" pageId="1" pageNumber="64">Australia</collectingCountry>
; QM I.37621,
<specimenCount id="9D62FD13FF90FFAAFF01353BFE9C407C" box="[250,325,1337,1361]" pageId="1" pageNumber="64" type="male">1 male</specimenCount>
(54.0 mm),
<specimenCount id="9D62FD13FF90FFAAFE34353BFDE9407C" box="[463,560,1337,1361]" pageId="1" pageNumber="64" type="female">1 female</specimenCount>
(
<quantity id="4C9C9B7FFF90FFAAFDC5353BFD7E407C" box="[574,679,1337,1361]" metricMagnitude="-2" metricUnit="m" metricValue="6.01" pageId="1" pageNumber="64" unit="mm" value="60.1">60.1 mm</quantity>
), and two juveniles (
<quantity id="4C9C9B7FFF90FFAAFC76353BFBEC407C" box="[909,1077,1337,1361]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" metricValueMax="1.41" metricValueMin="1.25" pageId="1" pageNumber="64" unit="mm" value="13.3" valueMax="14.1" valueMin="12.5">12.5–14.1 mm</quantity>
), Shag Rock, east side off North Stradbroke Island, Queensland,
<collectingCountry id="F373760AFF90FFAAFE06355FFDBE4058" box="[509,615,1373,1397]" name="Australia" pageId="1" pageNumber="64">Australia</collectingCountry>
.
<figureCitation id="135F2A1FFF90FFAAFD8A355FFD0E4058" box="[625,727,1373,1397]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1913,1936]" captionTargetBox="[151,1436,481,1873]" captionTargetId="figure@2.[151,1436,481,1874]" captionTargetPageId="2" captionText="FIGURE 1. A) Histiophryne maggiewalker, new species, holotype, QM I. 38176, 55 mm SL, showing some dehydration, and B) Histiophryne maggiewalker, paratype, QM I. 37621, 60.1 mm SL." httpUri="https://zenodo.org/record/277962/files/figure.png" pageId="1" pageNumber="64">Figures 1</figureCitation>
B and 2.
</paragraph>
</subSubSection>
<subSubSection id="C37E6511FF90FFAAFF3E3583FEAA432B" pageId="1" pageNumber="64" type="diagnosis">
<paragraph id="8BDB369AFF90FFAAFF3E3583FEAA432B" blockId="1.[151,1436,1229,2010]" pageId="1" pageNumber="64">
<emphasis id="B910EA88FF90FFAAFF3E3583FEE640B7" bold="true" box="[197,319,1409,1434]" pageId="1" pageNumber="64">Diagnosis.</emphasis>
A frogfish of the lophiiform family
<taxonomicName id="4C644D19FF90FFAAFD113583FC5540B4" box="[746,908,1409,1433]" class="Actinopterygii" family="Antennariidae" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="64" phylum="Chordata" rank="family">Antennariidae</taxonomicName>
, genus
<taxonomicName id="4C644D19FF90FFAAFC1F3583FBAF40B4" box="[996,1142,1409,1433]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="B910EA88FF90FFAAFC1F3583FBAF40B4" box="[996,1142,1409,1433]" italics="true" pageId="1" pageNumber="64">Histiophryne</emphasis>
</taxonomicName>
, unique in having a relatively long illicium and esca, the latter expanded distally, its shape more or less rectangular with rounded edges (
<figureCitation id="135F2A1FFF90FFAAFF6535CBFEDB40CC" box="[158,258,1481,1505]" captionStart="FIGURE 3" captionStartId="3.[151,250,1967,1990]" captionTargetBox="[520,1077,876,1954]" captionTargetId="figure@3.[520,1077,876,1954]" captionTargetPageId="3" captionText="FIGURE 3. The illicium of Histiophryne maggiewalker, QM I. 38176, 55 mm SL, and H. bougainvilli, AMS I. 17304 - 001, 58.9 mm SL." httpUri="https://zenodo.org/record/277964/files/figure.png" pageId="1" pageNumber="64">Figure 3</figureCitation>
), and genetic divergence in the nuclear recombination activation gene-2 (RAG2) and cytochrome oxidase-I (COI) genes.
</paragraph>
</subSubSection>
<subSubSection id="C37E6511FF90FFAFFF3E3613FE0C4449" lastPageId="4" lastPageNumber="67" pageId="1" pageNumber="64" type="description">
<paragraph id="8BDB369AFF90FFA9FF3E3613FAF245F9" blockId="1.[151,1436,1229,2010]" lastBlockId="2.[151,1437,151,428]" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64">
<emphasis id="B910EA88FF90FFAAFF3E3613FE8F4307" bold="true" box="[197,342,1553,1578]" pageId="1" pageNumber="64">Description.</emphasis>
Body short and deep, somewhat compressed laterally, vertebral column distinctly sigmoid in shape (as determined by radiographs); head large, its length approximately 29−39% standard length (SL); mouth small (relative to other antennariid taxa), opening strongly oblique to nearly vertical. Jaws bearing 2
<emphasis id="B910EA88FF90FFAAFAEE365AFAFD435F" box="[1301,1316,1624,1650]" italics="true" pageId="1" pageNumber="64">–</emphasis>
4 irregular rows of small villiform teeth; similar teeth present on vomer and palatine. Eye diameter 4.5
<emphasis id="B910EA88FF90FFAAFB573679FB6243B8" box="[1196,1211,1659,1685]" italics="true" pageId="1" pageNumber="64">–</emphasis>
4.9% and 8.8
<emphasis id="B910EA88FF90FFAAFAA93679FAB843B8" box="[1362,1377,1659,1685]" italics="true" pageId="1" pageNumber="64">–</emphasis>
9.1% SL for adults (four specimens) and juveniles (two specimens), respectively. Spinous dorsal fin consisting of three widely spaced spines, anteriormost spine (illicium) relatively long (2.5
<emphasis id="B910EA88FF90FFAAFC2B36C1FC0643F0" box="[976,991,1731,1757]" italics="true" pageId="1" pageNumber="64">–</emphasis>
4.7% SL); second and third dorsal-fin spines laid back and bound down to surface of head by thick skin; esca relatively long (4.1
<emphasis id="B910EA88FF90FFAAFB7036EAFB43422F" box="[1163,1178,1768,1794]" italics="true" pageId="1" pageNumber="64">–</emphasis>
6.4% SL), more or less rectangular with rounded edges. Pectoral-fin lobe elongate, leg-like, attached to side of body by loose skin for most of its length. Opercular opening restricted to a small, elongate, tube-like opening situated immediately posteroventral to base of pectoral fin. Caudal peduncle absent, thick fleshy posteriormost margin of soft-dorsal and anal fins extending posteriorly well beyond base of caudal fin, connecting to outermost caudal-fin rays. Skin covering head and body everywhere thick, fleshy, and naked, lacking any visible trace of dermal spinules or cutaneous filaments or appendages. Rays of dorsal, anal, pectoral, and pelvic fins all simple; those of caudal fin all bifurcate. Dorsal-fin rays 15; anal-fin rays 8; pectoral-fin rays 8; pelvic fin with 1 spine and 5 rays; caudal-fin rays 9; vertebrae 21–22, including the anteriormost and posteriormost centra fused to the cranium and hypural plate, respectively.
</paragraph>
<paragraph id="8BDB369AFF93FFA9FF3E30DDFDE74481" blockId="2.[151,1437,151,428]" pageId="2" pageNumber="65">
<emphasis id="B910EA88FF93FFA9FF3E30DDFE0B45D5" bold="true" box="[197,466,223,248]" pageId="2" pageNumber="65">Color in preservation.</emphasis>
The three largest specimens completely black, a reticulate pattern easily discernible with aid of a dissecting scope except in the
<typeStatus id="54DF8838FF93FFA9FD7A3106FD3F4431" box="[641,742,260,284]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
, for which the pattern can only be found with more difficulty, perhaps because the skin may have been damaged while the specimen lay washed up on the beach; the two juveniles and smaller adult creamy white with small brown spots on the head and surrounding the eyes, the spots continuing on the body in the small adult, but differ on the bodies of the two smaller juveniles in that the spots are relatively larger and fewer in number.
</paragraph>
<caption id="DF1B6612FF93FFA9FF6C377BFC974280" httpUri="https://zenodo.org/record/277962/files/figure.png" pageId="2" pageNumber="65" targetBox="[151,1436,481,1873]" targetPageId="2">
<paragraph id="8BDB369AFF93FFA9FF6C377BFC974280" blockId="2.[151,1436,1913,1966]" pageId="2" pageNumber="65">
<emphasis id="B910EA88FF93FFA9FF6C377BFECD42A3" bold="true" box="[151,276,1913,1936]" pageId="2" pageNumber="65">FIGURE 1.</emphasis>
A)
<taxonomicName id="4C644D19FF93FFA9FEC7377BFD8F42A2" box="[316,598,1913,1935]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="maggiewalker" status="sp. nov.">
<emphasis id="B910EA88FF93FFA9FEC7377BFD8F42A2" box="[316,598,1913,1935]" italics="true" pageId="2" pageNumber="65">Histiophryne maggiewalker</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22357F3FF93FFA9FD9F3778FD3842A2" box="[612,737,1913,1935]" pageId="2" pageNumber="65" rank="species">new species</taxonomicNameLabel>
, holotype, QM I.38176, 55 mm SL, showing some dehydration, and B)
<taxonomicName id="4C644D19FF93FFA9FF4D379AFE164283" box="[182,463,1944,1966]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="maggiewalker">
<emphasis id="B910EA88FF93FFA9FF4D379AFE164283" box="[182,463,1944,1966]" italics="true" pageId="2" pageNumber="65">Histiophryne maggiewalker</emphasis>
</taxonomicName>
, paratype, QM I.37621, 60.1 mm SL.
</paragraph>
</caption>
<caption id="DF1B6612FF92FFA8FF6C3328FC68466C" box="[151,945,810,833]" httpUri="https://zenodo.org/record/277963/files/figure.png" pageId="3" pageNumber="66" targetBox="[178,1417,176,799]" targetPageId="3">
<paragraph id="8BDB369AFF92FFA8FF6C3328FC68466C" blockId="3.[151,945,810,833]" box="[151,945,810,833]" pageId="3" pageNumber="66">
<emphasis id="B910EA88FF92FFA8FF6C3328FECA466D" bold="true" box="[151,275,810,833]" pageId="3" pageNumber="66">FIGURE 2.</emphasis>
<taxonomicName id="4C644D19FF92FFA8FEE23329FDE8466C" box="[281,561,811,833]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="66" phylum="Chordata" rank="species" species="maggiewalker">
<emphasis id="B910EA88FF92FFA8FEE23329FDE8466C" box="[281,561,811,833]" italics="true" pageId="3" pageNumber="66">Histiophryne maggiewalker</emphasis>
</taxonomicName>
, paratype, QM I.37621, 14.1 mm SL.
</paragraph>
</caption>
<caption id="DF1B6612FF92FFA8FF6C37ADFF3042C9" httpUri="https://zenodo.org/record/277964/files/figure.png" pageId="3" pageNumber="66" targetBox="[520,1077,876,1954]" targetPageId="3">
<paragraph id="8BDB369AFF92FFA8FF6C37ADFF3042C9" blockId="3.[151,1436,1967,2020]" pageId="3" pageNumber="66">
<emphasis id="B910EA88FF92FFA8FF6C37ADFECB42E8" bold="true" box="[151,274,1967,1990]" pageId="3" pageNumber="66">FIGURE 3.</emphasis>
The illicium of
<taxonomicName id="4C644D19FF92FFA8FE4E37ADFD1642E8" box="[437,719,1967,1989]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="66" phylum="Chordata" rank="species" species="maggiewalker">
<emphasis id="B910EA88FF92FFA8FE4E37ADFD1642E8" box="[437,719,1967,1989]" italics="true" pageId="3" pageNumber="66">Histiophryne maggiewalker</emphasis>
</taxonomicName>
, QM I.38176, 55 mm SL, and
<taxonomicName id="4C644D19FF92FFA8FBFE37B3FB4442E8" box="[1029,1181,1967,1990]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="66" phylum="Chordata" rank="species" species="bougainvilli">
<emphasis id="B910EA88FF92FFA8FBFE37B3FB4442E8" box="[1029,1181,1967,1990]" italics="true" pageId="3" pageNumber="66">H. bougainvilli</emphasis>
</taxonomicName>
, AMS I.17304-001, 58.9 mm SL.
</paragraph>
</caption>
<paragraph id="8BDB369AFF95FFAFFF3E309AFE0C4449" blockId="4.[151,1436,151,356]" pageId="4" pageNumber="67">
<emphasis id="B910EA88FF95FFAFFF3E309AFE47459D" bold="true" box="[197,414,152,176]" pageId="4" pageNumber="67">Molecular results.</emphasis>
Results are summarized in
<figureCitation id="135F2A1FFF95FFAFFD203095FCE74582" box="[731,830,151,175]" captionStart="FIGURE 4. 50" captionStartId="4.[151,250,1474,1497]" captionTargetBox="[237,1334,413,1430]" captionTargetId="figure@4.[237,1339,409,1435]" captionTargetPageId="4" captionText="FIGURE 4. 50 % majority rule phylogeny for all described species of Histiophryne, from trees sampled in the posterior, generated from Bayesian analyses on a partitioned dataset that included the nuclear recombination activating gene 2 (RAG 2) and mitochondrial cytochrome oxidase-I (COI) gene." httpUri="https://zenodo.org/record/277965/files/figure.png" pageId="4" pageNumber="67">Figure 4</figureCitation>
. Analysis of RAG2 and COI sequences included 441 and 570 bps, respectively.
<taxonomicName id="4C644D19FF95FFAFFE3730BEFCDE45F9" box="[460,775,188,212]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="maggiewalker">
<emphasis id="B910EA88FF95FFAFFE3730BEFCDE45F9" box="[460,775,188,212]" italics="true" pageId="4" pageNumber="67">Histiophryne maggiewalker</emphasis>
</taxonomicName>
was recovered as sister to
<taxonomicName id="4C644D19FF95FFAFFBBD30BFFB2B45F9" box="[1094,1266,188,212]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="bougainvilli">
<emphasis id="B910EA88FF95FFAFFBBD30BFFB2B45F9" box="[1094,1266,188,212]" italics="true" pageId="4" pageNumber="67">H. bougainvilli</emphasis>
</taxonomicName>
with posterior probability of 1.0.
<taxonomicName id="4C644D19FF95FFAFFE8830E3FDE145D5" box="[371,568,224,248]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="cryptacanthus">
<emphasis id="B910EA88FF95FFAFFE8830E3FDE145D5" box="[371,568,224,248]" italics="true" pageId="4" pageNumber="67">H. cryptacanthus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C644D19FF95FFAFFD8E30E3FCF545D5" box="[629,812,224,248]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="psychedelica">
<emphasis id="B910EA88FF95FFAFFD8E30E3FCF545D5" box="[629,812,224,248]" italics="true" pageId="4" pageNumber="67">H. psychedelica</emphasis>
</taxonomicName>
, the two tropical species included in this study, were recovered as sister to
<taxonomicName id="4C644D19FF95FFAFFE713107FDEA4431" box="[394,563,260,284]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="bougainvilli">
<emphasis id="B910EA88FF95FFAFFE713107FDEA4431" box="[394,563,260,284]" italics="true" pageId="4" pageNumber="67">H. bougainvilli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C644D19FF95FFAFFD913107FCF54431" box="[618,812,260,284]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="maggiewalker">
<emphasis id="B910EA88FF95FFAFFD913107FCF54431" box="[618,812,260,284]" italics="true" pageId="4" pageNumber="67">H. maggiewalker</emphasis>
</taxonomicName>
with posterior probability of 1.0. The intraspecific COI DNA sequence divergence between
<taxonomicName id="4C644D19FF95FFAFFDD4312BFD28446D" box="[559,753,296,320]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="cryptacanthus">
<emphasis id="B910EA88FF95FFAFFDD4312BFD28446D" box="[559,753,296,320]" italics="true" pageId="4" pageNumber="67">H. cryptacanthus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C644D19FF95FFAFFCDC312BFC02446D" box="[807,987,296,320]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="psychedelica">
<emphasis id="B910EA88FF95FFAFFCDC312BFC02446D" box="[807,987,296,320]" italics="true" pageId="4" pageNumber="67">H. psychedelica</emphasis>
</taxonomicName>
was 6.1%, and between
<taxonomicName id="4C644D19FF95FFAFFB09312BFA45446D" box="[1266,1436,296,320]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="bougainvilli">
<emphasis id="B910EA88FF95FFAFFB09312BFA45446D" box="[1266,1436,296,320]" italics="true" pageId="4" pageNumber="67">H. bougainvilli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C644D19FF95FFAFFF3D314FFE5C4449" box="[198,389,332,356]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="maggiewalker">
<emphasis id="B910EA88FF95FFAFFF3D314FFE5C4449" box="[198,389,332,356]" italics="true" pageId="4" pageNumber="67">H. maggiewalker</emphasis>
</taxonomicName>
, 7.1%.
</paragraph>
</subSubSection>
<caption id="DF1B6612FF95FFAFFF6C35C0FD5E433B" httpUri="https://zenodo.org/record/277965/files/figure.png" pageId="4" pageNumber="67" targetBox="[237,1334,413,1430]" targetPageId="4">
<paragraph id="8BDB369AFF95FFAFFF6C35C0FD5E433B" blockId="4.[151,1436,1474,1559]" pageId="4" pageNumber="67">
<emphasis id="B910EA88FF95FFAFFF6C35C0FECB40F5" bold="true" box="[151,274,1474,1497]" pageId="4" pageNumber="67">FIGURE 4.</emphasis>
50% majority rule phylogeny for all described species of
<taxonomicName id="4C644D19FF95FFAFFCA535C1FC3B40F4" box="[862,994,1475,1497]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="genus">
<emphasis id="B910EA88FF95FFAFFCA535C1FC3B40F4" box="[862,994,1475,1497]" italics="true" pageId="4" pageNumber="67">Histiophryne</emphasis>
</taxonomicName>
, from trees sampled in the posterior, generated from Bayesian analyses on a partitioned dataset that included the nuclear recombination activating gene 2 (RAG2) and mitochondrial cytochrome oxidase-I (COI) gene.
</paragraph>
</caption>
<subSubSection id="C37E6511FF95FFAFFF3E3641FB344352" pageId="4" pageNumber="67" type="etymology">
<paragraph id="8BDB369AFF95FFAFFF3E3641FB344352" blockId="4.[151,1436,1603,2024]" pageId="4" pageNumber="67">
<emphasis id="B910EA88FF95FFAFFF3E3641FE924376" bold="true" box="[197,331,1603,1627]" pageId="4" pageNumber="67">Etymology.</emphasis>
This species is named in honor of Maggie Walker for her service and dedication to science and education at the University of Washington and The Burke Museum of Natural History and Culture.
</paragraph>
</subSubSection>
<subSubSection id="C37E6511FF95FFAFFF3E3688FCF3427A" pageId="4" pageNumber="67" type="distribution">
<paragraph id="8BDB369AFF95FFAFFF3E3688FCF3427A" blockId="4.[151,1436,1603,2024]" pageId="4" pageNumber="67">
<emphasis id="B910EA88FF95FFAFFF3E3688FDD8438E" bold="true" box="[197,513,1674,1699]" pageId="4" pageNumber="67">Distribution and behavior.</emphasis>
Known only from shallow inshore waters of Queensland,
<collectingCountry id="F373760AFF95FFAFFB743689FB21438E" box="[1167,1272,1675,1699]" name="Australia" pageId="4" pageNumber="67">Australia</collectingCountry>
, all specimens collected within 130 kilometers, in
<quantity id="4C9C9B7FFF95FFAFFDC836ADFD5243EA" box="[563,651,1711,1735]" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="4" pageNumber="67" unit="m" value="9.5" valueMax="12.0" valueMin="7.0">7–12 m</quantity>
of water; the
<typeStatus id="54DF8838FF95FFAFFCC836ADFC4C43EA" box="[819,917,1711,1735]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
washed up on the beach of Alexandria Bay, Noosa Heads, southeast Queensland. Four specimens, a non-gravid female, male, and two juveniles, were collected together. While some frogfishes are notorious for their insatiable appetite, including tendencies to cannibalize one another, it is possible these four were living together peacefully. We might even be tempted to speculate that the two juveniles are the offspring of one or both of the adults.
</paragraph>
</subSubSection>
<subSubSection id="C37E6511FF95FFAEFF3C3760FA924449" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67" type="materials_examined">
<paragraph id="8BDB369AFF95FFAEFF3C3760FA924449" blockId="4.[151,1436,1603,2024]" lastBlockId="5.[151,1436,151,356]" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
<emphasis id="B910EA88FF95FFAFFF3C3760FE6D4256" bold="true" box="[199,436,1890,1915]" pageId="4" pageNumber="67">Material examined.</emphasis>
<taxonomicName id="4C644D19FF95FFAFFE3B3761FCC44256" authority="Abrolmus" authorityName="Abrolmus" box="[448,797,1891,1915]" class="Actinopterygii" family="Antennariidae" genus="Tathicarpus" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="butleri">
<emphasis id="B910EA88FF95FFAFFE3B3761FD434256" box="[448,666,1891,1915]" italics="true" pageId="4" pageNumber="67">Tathicarpus butleri</emphasis>
, Abrolmus
</taxonomicName>
Island, Western
<collectingCountry id="F373760AFF95FFAFFC1E3761FB964256" box="[997,1103,1891,1915]" name="Australia" pageId="4" pageNumber="67">Australia</collectingCountry>
,
<collectionCode id="ED75AE5FFF95FFAFFBA53761FB7B4256" LSID="urn:lsid:biocol.org:col:34981" box="[1118,1186,1891,1915]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="4" pageNumber="67">WAM</collectionCode>
32904.001, Genbank accession nos.
<accessionNumber id="9437AB79FF95FFAFFEB93785FE1842B2" box="[322,449,1927,1951]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GU188502" pageId="4" pageNumber="67">GU188502</accessionNumber>
,
<accessionNumber id="9437AB79FF95FFAFFE343785FD9642B2" box="[463,591,1927,1951]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GU188551" pageId="4" pageNumber="67">GU188551</accessionNumber>
;
<taxonomicName id="4C644D19FF95FFAFFDA73785FC2542B2" authority="Aquarium" authorityName="Aquarium" box="[604,1020,1927,1951]" class="Actinopterygii" family="Antennariidae" genus="Lophiocharon" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="trisignatus">
<emphasis id="B910EA88FF95FFAFFDA73785FCA242B2" box="[604,891,1927,1951]" italics="true" pageId="4" pageNumber="67">Lophiocharon trisignatus</emphasis>
, Aquarium
</taxonomicName>
trade, UW 115748, Genbank accession nos.
<accessionNumber id="9437AB79FF95FFAFFEF937A9FEAA42EE" box="[258,371,1963,1987]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ219610" pageId="4" pageNumber="67">FJ219610</accessionNumber>
,
<accessionNumber id="9437AB79FF95FFAFFE8537A9FE2842EE" box="[382,497,1963,1987]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ219617" pageId="4" pageNumber="67">FJ219617</accessionNumber>
;
<taxonomicName id="4C644D19FF95FFAFFE0737AEFCF642EE" authority="Aquarium" authorityName="Aquarium" box="[508,815,1963,1987]" class="Actinopterygii" family="Antennariidae" genus="Lophiocharon" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="lithinostomus">
<emphasis id="B910EA88FF95FFAFFE0737AEFD6942EE" box="[508,688,1963,1987]" italics="true" pageId="4" pageNumber="67">L. lithinostomus</emphasis>
, Aquarium
</taxonomicName>
trade, UW 115749, Genbank accession nos.
<accessionNumber id="9437AB79FF95FFAFFADC37A9FA4142EE" box="[1319,1432,1963,1987]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ219611" pageId="4" pageNumber="67">FJ219611</accessionNumber>
,
<accessionNumber id="9437AB79FF95FFAFFF6C37CDFED042CA" box="[151,265,1999,2023]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ219618" pageId="4" pageNumber="67">FJ219618</accessionNumber>
;
<taxonomicName id="4C644D19FF95FFAFFEED37CDFC6E42CA" authority="Alexandria Bay, Noosa Heads" authorityName="Alexandria Bay, Noosa Heads" box="[278,951,1999,2024]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="maggiewalker">
<emphasis id="B910EA88FF95FFAFFEED37CDFD9742CA" box="[278,590,1999,2023]" italics="true" pageId="4" pageNumber="67">Histiophryne maggiewalker</emphasis>
, Alexandria Bay, Noosa Heads
</taxonomicName>
, southeast Queensland, QM I.38176, Genbank accession nos. FJ755972,
<accessionNumber id="9437AB79FF94FFAEFE0D3095FDB14582" box="[502,616,151,175]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JF755974" pageId="5" pageNumber="68">JF755974</accessionNumber>
;
<taxonomicName id="4C644D19FF94FFAEFD88309BFC74459D" authority="Nelson Bay" authorityName="Nelson Bay" box="[627,941,151,176]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="bougainvilli">
<emphasis id="B910EA88FF94FFAEFD88309BFCC2459D" box="[627,795,152,176]" italics="true" pageId="5" pageNumber="68">H. bougainvilli</emphasis>
, Nelson Bay
</taxonomicName>
,
<collectionCode id="ED75AE5FFF94FFAEFC433095FC2E4582" LSID="urn:lsid:biocol.org:col:15627" box="[952,1015,151,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="5" pageNumber="68">NSW</collectionCode>
,
<collectingCountry id="F373760AFF94FFAEFBFA3095FBB24582" box="[1025,1131,151,175]" name="Australia" pageId="5" pageNumber="68">Australia</collectingCountry>
, UW 118990 (1 of 2), Genbank accession nos.
<accessionNumber id="9437AB79FF94FFAEFE7C30BEFE2145F9" box="[391,504,188,212]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JF755970" pageId="5" pageNumber="68">JF755970</accessionNumber>
,
<accessionNumber id="9437AB79FF94FFAEFDFC30BEFDA045F9" box="[519,633,188,212]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JF755973" pageId="5" pageNumber="68">JF755973</accessionNumber>
;
<taxonomicName id="4C644D19FF94FFAEFD7C30BFFC1745F8" authority="Nelson Bay" authorityName="Nelson Bay" box="[647,974,188,213]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="bougainvilli">
<emphasis id="B910EA88FF94FFAEFD7C30BFFCED45F9" box="[647,820,188,212]" italics="true" pageId="5" pageNumber="68">H. bougainvilli</emphasis>
, Nelson Bay
</taxonomicName>
,
<collectionCode id="ED75AE5FFF94FFAEFC2630BEFBC545F9" LSID="urn:lsid:biocol.org:col:15627" box="[989,1052,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="5" pageNumber="68">NSW</collectionCode>
,
<collectingCountry id="F373760AFF94FFAEFBD130BEFB4D45F9" box="[1066,1172,188,212]" name="Australia" pageId="5" pageNumber="68">Australia</collectingCountry>
, UW 118990 (2 of 2), Genbank accession nos.
<accessionNumber id="9437AB79FF94FFAEFE4A30DDFDFA45DA" box="[433,547,223,247]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JF755973" pageId="5" pageNumber="68">JF755973</accessionNumber>
,
<accessionNumber id="9437AB79FF94FFAEFDCB30DDFD7A45DA" box="[560,675,223,247]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JF755976" pageId="5" pageNumber="68">JF755976</accessionNumber>
;
<taxonomicName id="4C644D19FF94FFAEFD4B30E3FCBC45D5" box="[688,869,224,248]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="psychedelica">
<emphasis id="B910EA88FF94FFAEFD4B30E3FCBC45D5" box="[688,869,224,248]" italics="true" pageId="5" pageNumber="68">H. psychedelica</emphasis>
</taxonomicName>
,
<collectingCountry id="F373760AFF94FFAEFC8E30DDFC1045DA" box="[885,969,223,247]" name="Indonesia" pageId="5" pageNumber="68">Ambon</collectingCountry>
Island,
<collectingCountry id="F373760AFF94FFAEFBD330DDFB4145DA" box="[1064,1176,223,247]" name="Indonesia" pageId="5" pageNumber="68">Indonesia</collectingCountry>
,
<collectionCode id="ED75AE5FFF94FFAEFB5E30DDFB3C45DA" box="[1189,1253,223,247]" httpUri="http://grbio.org/cool/h4qb-ekf6" name="Pusat Penelitian dan Pengembangan Oseanologi" pageId="5" pageNumber="68">NCIP</collectionCode>
6377, Genbank accession nos.
<accessionNumber id="9437AB79FF94FFAEFEBC3106FE614431" box="[327,440,260,284]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ219608" pageId="5" pageNumber="68">FJ219608</accessionNumber>
,
<accessionNumber id="9437AB79FF94FFAEFE333106FDE34431" box="[456,570,260,284]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ219615" pageId="5" pageNumber="68">FJ219615</accessionNumber>
;
<taxonomicName id="4C644D19FF94FFAEFDB23107FC4B4431" authority="Aquarium" authorityName="Aquarium" box="[585,914,260,284]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="cryptacanthus">
<emphasis id="B910EA88FF94FFAEFDB23107FCD74431" box="[585,782,260,284]" italics="true" pageId="5" pageNumber="68">H. cryptacanthus</emphasis>
, Aquarium
</taxonomicName>
trade, UW 117820, Genbank accession nos.
<accessionNumber id="9437AB79FF94FFAEFF6C3125FED14412" box="[151,264,295,319]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ219614" pageId="5" pageNumber="68">FJ219614</accessionNumber>
,
<accessionNumber id="9437AB79FF94FFAEFEE03125FE554412" box="[283,396,295,319]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ219621" pageId="5" pageNumber="68">FJ219621</accessionNumber>
;
<taxonomicName id="4C644D19FF94FFAEFE64312BFD374412" authority="Aquarium" authorityName="Aquarium" box="[415,750,295,320]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="cryptacanthus">
<emphasis id="B910EA88FF94FFAEFE64312BFDB1446D" box="[415,616,296,320]" italics="true" pageId="5" pageNumber="68">H. cryptacanthus</emphasis>
, Aquarium
</taxonomicName>
trade, UW 117821, Genbank accession nos.
<accessionNumber id="9437AB79FF94FFAEFAE33125FA414412" box="[1304,1432,295,319]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GU188512" pageId="5" pageNumber="68">GU188512</accessionNumber>
,
<accessionNumber id="9437AB79FF94FFAEFF6C314EFECE4449" box="[151,279,332,356]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GU188545" pageId="5" pageNumber="68">GU188545</accessionNumber>
;
<taxonomicName id="4C644D19FF94FFAEFED9314FFDBB4449" authority="Aquarium" authorityName="Aquarium" box="[290,610,332,356]" class="Actinopterygii" family="Antennariidae" genus="Histiophryne" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="cryptacanthus">
<emphasis id="B910EA88FF94FFAEFED9314FFE3A4449" box="[290,483,332,356]" italics="true" pageId="5" pageNumber="68">H. cryptacanthus</emphasis>
, Aquarium
</taxonomicName>
trade, UW 117816, Genbank accession nos.
<accessionNumber id="9437AB79FF94FFAEFBA2314EFB124449" box="[1113,1227,332,356]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JF755971" pageId="5" pageNumber="68">JF755971</accessionNumber>
,
<accessionNumber id="9437AB79FF94FFAEFB2E314EFA914449" box="[1237,1352,332,356]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JF755977" pageId="5" pageNumber="68">JF755977</accessionNumber>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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