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<document id="1E1EE0CBB963204A32D70E3B87A2CE02" ID="10.11646/zootaxa.4189.3.6" ID-DOI="10.11646/zootaxa.4189.3.6" ID-GBIF-Dataset="36961809-00bf-492f-a723-ae7baf258057" ID-ISSN="1175-5326" ID-Zenodo-Dep="166209" ID-ZooBank="D3C2ED63-FF05-442F-8425-3960050DDA18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478850269485" checkinUser="plazi" docAuthor="Kozlov, Mikhail V." docDate="2016" docId="03B7E700FF9CFF80FF6DDEE2E33EA3C6" docLanguage="en" docName="zootaxa.4189.3.6.pdf" docOrigin="Zootaxa 4189 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nemophora humerella Kozlov, sp. nov." docType="treatment" docVersion="8" lastPageNumber="566" masterDocId="FF8E9F78FF9AFF87FFFAD81BE225A703" masterDocTitle="A taxonomic revision of the kalshoveni species-group of the genus Nemophora Hoffmannsegg (Lepidoptera, Adelidae), with descriptions of six new species from Indonesia and Papua New Guinea" masterLastPageNumber="570" masterPageNumber="559" pageNumber="565" updateTime="1698393258691" updateUser="plazi">
<mods:mods id="BDBCE1B71B439514303C95D481A7FB42" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EA7F32EED57EF7A7B3D909EE67369268">A taxonomic revision of the kalshoveni species-group of the genus Nemophora Hoffmannsegg (Lepidoptera, Adelidae), with descriptions of six new species from Indonesia and Papua New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="EA048D44325F9965AB1C4CE16A3D4DE8" type="personal">
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<mods:namePart id="F5901E4CBE34C7E52294D18787AB9EEA">Kozlov, Mikhail V.</mods:namePart>
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<mods:title id="1E84AA4FF6A8A842058F22D72385C547">Zootaxa</mods:title>
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<mods:date id="4B244C56CF90C879DC73E14D623537C4">2016</mods:date>
<mods:detail id="8C400D101EF018182C1C531153A0A1CB" type="volume">
<mods:number id="046F98248F1E8D40BE3BD960A392C2F3">4189</mods:number>
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<mods:classification id="B0AF2A0741D177D20CE4BD7D9DCD9D22">journal article</mods:classification>
<mods:identifier id="48AA9593812B5AD6E9ACE4DC91104422" type="DOI">10.11646/zootaxa.4189.3.6</mods:identifier>
<mods:identifier id="3F7E86EFEFC5B2FBB56CDA6AC3604452" type="GBIF-Dataset">36961809-00bf-492f-a723-ae7baf258057</mods:identifier>
<mods:identifier id="91BE1BD616AD52F52FD20381E15362BC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BE8FC01EB093F91F996048A47F32E174" type="Zenodo-Dep">166209</mods:identifier>
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<treatment id="03B7E700FF9CFF80FF6DDEE2E33EA3C6" ID-DOI="http://doi.org/10.5281/zenodo.6088822" ID-GBIF-Taxon="124691848" ID-Zenodo-Dep="6088822" LSID="urn:lsid:plazi:treatment:03B7E700FF9CFF80FF6DDEE2E33EA3C6" httpUri="http://treatment.plazi.org/id/03B7E700FF9CFF80FF6DDEE2E33EA3C6" lastPageId="7" lastPageNumber="566" pageId="6" pageNumber="565">
<subSubSection id="C304059DFF9CFF81FF6DDEE2E356A036" pageId="6" pageNumber="565" type="nomenclature">
<paragraph id="8BA15616FF9CFF81FF6DDEE2E056A017" blockId="6.[151,627,1785,1845]" box="[151,627,1785,1812]" pageId="6" pageNumber="565">
<heading id="D0E9E17AFF9CFF81FF6DDEE2E056A017" bold="true" box="[151,627,1785,1812]" fontSize="11" level="1" pageId="6" pageNumber="565" reason="1">
<emphasis id="B96A8A04FF9CFF81FF6DDEE2E056A017" bold="true" box="[151,627,1785,1812]" pageId="6" pageNumber="565">
<taxonomicName id="4C1E2D95FF9CFF81FF6DDEE2E02DA017" authority="Kozlov" authorityName="Kozlov" box="[151,520,1785,1812]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="humerella" status="sp. nov.">
<emphasis id="B96A8A04FF9CFF81FF6DDEE2E38CA010" bold="true" box="[151,425,1785,1811]" italics="true" pageId="6" pageNumber="565">Nemophora humerella</emphasis>
Kozlov
</taxonomicName>
,
<taxonomicNameLabel id="A259377FFF9CFF81FDEFDEE1E056A017" box="[533,627,1786,1812]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA15616FF9CFF81FF6DDF07E356A036" blockId="6.[151,627,1785,1845]" box="[151,371,1820,1845]" pageId="6" pageNumber="565">
(
<figureCitation id="13254A93FF9CFF81FF65DF07E2CBA036" box="[159,238,1820,1845]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1853,1876]" captionTargetBox="[188,1397,193,1831]" captionTargetId="figure@3.[188,1398,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Adults of Nemophora kalshoveni species-group. 1, N. kalshoveni Diakonoff, 1951, male, from Cisaura; 2, N. humerella sp. nov., male, holotype, from Sangir Island; 3, N. agassizi sp. nov., female, holotype, from Wabag; 4, N. dohertyi sp. nov., male, holotype, from Fakfak; 5, N. bistrigata sp. nov., male, holotype, from Jayapura; 6, N. toxopeusi sp. nov., female, holotype, from Irian Jaya; 7, N. stenopterella sp. nov., female, holotype, from Irian Jaya; 8, labels of the holotype of N. kalshoveni Diakonoff. Scale: 2 mm." httpUri="https://zenodo.org/record/166210/files/figure.png" pageId="6" pageNumber="565">Figs. 2</figureCitation>
,
<figureCitation id="13254A93FF9CFF81FF00DF06E332A036" box="[250,279,1821,1845]" captionStart="FIGURES 10 – 17" captionStartId="4.[151,264,1940,1963]" captionTargetBox="[175,1402,875,1912]" captionTargetId="figure@4.[173,1414,871,1919]" captionTargetPageId="4" captionText="FIGURES 10 – 17. Forewing pattern and venation of Nemophora kalshoveni species-group. 10, N. kalshoveni Diakonoff; 11, N. humerella sp. nov.; 12, N. agassizi sp. nov.; 13, N. dohertyi sp. nov.; 14, 16, N. bistrigata sp. nov.; 15, N. toxopeusi sp. nov.; 17 N. stenopterella sp. nov. Scale: 2 mm." httpUri="https://zenodo.org/record/166212/files/figure.png" pageId="6" pageNumber="565">11</figureCitation>
,
<figureCitation id="13254A93FF9CFF81FED9DF06E34FA036" box="[291,362,1821,1845]" captionStart="FIGURES 18 - 37" captionStartId="5.[151,264,1514,1537]" captionTargetBox="[169,1424,200,1474]" captionTargetId="figure@5.[151,1436,193,1493]" captionTargetPageId="5" captionText="FIGURES 18 - 37. Male genitalia of Nemophora kalshoveni species-group. 18 – 22, N. kalshoveni Diakonoff; 23 – 27, N. humerella sp. nov.; 28 – 32, N. dohertyi sp. nov.; 33 – 37, N. bistrigata sp. nov. 18, 23, 28, 33: genital complex, ventral view (right valva not shown); 19, 24, 29, 34: genital complex, lateral view; 20, 25, 30, 35: juxta; 21, 26, 31, 36: phallus, ventral view; 22, 27, 32, 37: phallus, lateral view. Scale: 0.5 mm." httpUri="https://zenodo.org/record/166213/files/figure.png" pageId="6" pageNumber="565">23–27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C304059DFF9CFF81FF6DDF7CE647A0EF" pageId="6" pageNumber="565" type="materials_examined">
<paragraph id="8BA15616FF9CFF81FF6DDF7CE647A0EF" blockId="6.[151,1436,1892,2028]" pageId="6" pageNumber="565">
<materialsCitation id="3B765C4BFF9CFF81FF6DDF7CE179A07C" ID-GBIF-Occurrence="1324964196" box="[151,860,1892,1920]" country="Indonesia" latitude="3.6666667" location="Sangir Island" longLatPrecision="1307" longitude="125.5" pageId="6" pageNumber="565" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A5E8B4FF9CFF81FF6DDF7CE324A07C" box="[151,257,1895,1919]" pageId="6" pageNumber="565" type="holotype">
<emphasis id="B96A8A04FF9CFF81FF6DDF7CE324A07C" bold="true" box="[151,257,1895,1919]" pageId="6" pageNumber="565">Holotype</emphasis>
</typeStatus>
<specimenCount id="9D189D9FFF9CFF81FEFDDF73E33AA07C" box="[263,287,1896,1919]" pageId="6" pageNumber="565" type="male">♂</specimenCount>
:
<collectingCountry id="F3091686FF9CFF81FED0DF7CE3B2A083" box="[298,407,1895,1920]" name="Indonesia" pageId="6" pageNumber="565">Indonesia</collectingCountry>
,
<location id="8EC100CDFF9CFF81FE5BDF7CE017A083" LSID="urn:lsid:plazi:treatment:03B7E700FF9CFF80FF6DDEE2E33EA3C6:8EC100CDFF9CFF81FE5BDF7CE017A083" box="[417,562,1895,1920]" country="Indonesia" latitude="3.6666667" longLatPrecision="1307" longitude="125.5" name="Sangir Island" pageId="6" pageNumber="565">Sangir Island</location>
(approx.
<geoCoordinate id="EE2A30D1FF9CFF81FD60DF7CE0C1A07C" box="[666,740,1892,1919]" direction="north" orientation="latitude" pageId="6" pageNumber="565" precision="925" value="3.6666667">3o40'N</geoCoordinate>
,
<geoCoordinate id="EE2A30D1FF9CFF81FD0ADF7CE171A07C" box="[752,852,1892,1920]" direction="east" orientation="longitude" pageId="6" pageNumber="565" precision="925" value="125.5">125o30'E</geoCoordinate>
)
</materialsCitation>
; labelled:
<quantity id="4CE6FBF3FF9CFF81FC36DF7CE62EA07C" box="[972,1035,1895,1919]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="565" unit="mm" value="8.0">8 mm</quantity>
circle with red margin, print ‘Holo- | type’; 7 ×
<quantity id="4CE6FBF3FF9CFF81FEEBDF97E341A0A0" box="[273,356,1932,1956]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="565" unit="mm" value="13.0">13 mm</quantity>
, print + black ink ‘SANGUIR. | Doherty, 1892. | No. 41745’; 7 ×
<quantity id="4CE6FBF3FF9CFF81FBA4DF97E694A0A0" box="[1118,1201,1932,1956]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="565" unit="mm" value="10.0">10 mm</quantity>
, print ‘Walsingham | Collection. | 1910-427.’; 11 ×
<quantity id="4CE6FBF3FF9CFF81FE1FDFB4E013A0C4" box="[485,566,1967,1991]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="565" unit="mm" value="23.0">23 mm</quantity>
, black ink ‘
<taxonomicName id="4C1E2D95FF9CFF81FD49DFB4E100A0CB" box="[691,805,1967,1992]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="humerella">humerella</taxonomicName>
| Type ♂’; 12 ×
<quantity id="4CE6FBF3FF9CFF81FC1ADFB4E614A0C4" box="[992,1073,1967,1991]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="565" unit="mm" value="12.0">12 mm</quantity>
, print ‘B. M.
<materialsCitation id="3B765C4BFF9CFF81FB30DFB4E647A0EF" ID-GBIF-Occurrence="1324964202" collectionCode="NHM" country="Indonesia" location="Kozlov'" pageId="6" pageNumber="565" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<specimenCount id="9D189D9FFF9CFF81FB30DFB4E6C4A0C4" box="[1226,1249,1967,1991]" pageId="6" pageNumber="565" type="male">♂</specimenCount>
| Genitalia Slide | No. 27823’; 8 ×
<quantity id="4CE6FBF3FF9CFF81FEB1DFCFE3BEA0E8" box="[331,411,2004,2028]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="565" unit="mm" value="18.0">18 mm</quantity>
, print ‘
<typeStatus id="54A5E8B4FF9CFF81FE19DFCFE05CA0EF" box="[483,633,2004,2028]" pageId="6" pageNumber="565" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D189D9FFF9CFF81FD7ADFCFE0BDA0EF" box="[640,664,2004,2028]" pageId="6" pageNumber="565" type="male">♂</specimenCount>
|
<taxonomicName id="4C1E2D95FF9CFF81FD5BDFC8E107A0E8" box="[673,802,2003,2027]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B96A8A04FF9CFF81FD5BDFC8E107A0E8" box="[673,802,2003,2027]" italics="true" pageId="6" pageNumber="565">Nemophora</emphasis>
</taxonomicName>
|
<taxonomicName id="4C1E2D95FF9CFF81FCCEDFC8E184A0E8" box="[820,929,2003,2027]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="humerella">
<emphasis id="B96A8A04FF9CFF81FCCEDFC8E184A0E8" box="[820,929,2003,2027]" italics="true" pageId="6" pageNumber="565">humerella</emphasis>
</taxonomicName>
<location id="8EC100CDFF9CFF81FC52DFC8E624A0EF" LSID="urn:lsid:plazi:treatment:03B7E700FF9CFF80FF6DDEE2E33EA3C6:8EC100CDFF9CFF81FC52DFC8E624A0EF" box="[936,1025,2003,2028]" country="Indonesia" name="Kozlov'" pageId="6" pageNumber="565">Kozlov’</location>
(
<collectionCode id="ED0FCED3FF9CFF81FBEADFCFE673A0EF" box="[1040,1110,2004,2028]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="6" pageNumber="565">NHM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C304059DFF9DFF80FF3DD88CE7B9A667" pageId="7" pageNumber="566" type="diagnosis">
<paragraph id="8BA15616FF9DFF80FF3DD88CE7B9A667" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="566">
<emphasis id="B96A8A04FF9DFF80FF3DD88CE31FA7B3" bold="true" box="[199,314,151,176]" pageId="7" pageNumber="566">Diagnosis</emphasis>
. Most similar to
<taxonomicName id="4C1E2D95FF9DFF80FE04D882E0B2A7B3" box="[510,663,152,176]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="kalshoveni">
<emphasis id="B96A8A04FF9DFF80FE04D882E0B2A7B3" box="[510,663,152,176]" italics="true" pageId="7" pageNumber="566">N. kalshoveni</emphasis>
</taxonomicName>
, from which it differs by grey background colour of forewing (dark brown in
<taxonomicName id="4C1E2D95FF9DFF80FEFFD8A6E3BAA7D7" box="[261,415,188,212]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="kalshoveni">
<emphasis id="B96A8A04FF9DFF80FEFFD8A6E3BAA7D7" box="[261,415,188,212]" italics="true" pageId="7" pageNumber="566">N. kalshoveni</emphasis>
</taxonomicName>
), white colour of light scales scattered in the basal field of forewing and forming oblique stripe in apical part of it (yellow in
<taxonomicName id="4C1E2D95FF9DFF80FDE4D8FAE093A7FB" box="[542,694,224,248]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="kalshoveni">
<emphasis id="B96A8A04FF9DFF80FDE4D8FAE093A7FB" box="[542,694,224,248]" italics="true" pageId="7" pageNumber="566">N. kalshoveni</emphasis>
</taxonomicName>
), and by narrower, widely rounded valva (distal part of valva in
<taxonomicName id="4C1E2D95FF9DFF80FA78D8FAE32AA61F" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="kalshoveni">
<emphasis id="B96A8A04FF9DFF80FA78D8FAE32AA61F" italics="true" pageId="7" pageNumber="566">N. kalshoveni</emphasis>
</taxonomicName>
is nearly rectangular) extending beyond the apex of the tegumen (in
<taxonomicName id="4C1E2D95FF9DFF80FBE8D91EE68FA61F" box="[1042,1194,260,284]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="kalshoveni">
<emphasis id="B96A8A04FF9DFF80FBE8D91EE68FA61F" box="[1042,1194,260,284]" italics="true" pageId="7" pageNumber="566">N. kalshoveni</emphasis>
</taxonomicName>
, apex of the tegumen extends beyond the apex of valva). Also resembles
<taxonomicName id="4C1E2D95FF9DFF80FD0AD932E14BA63C" box="[752,878,296,320]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="agassizi">
<emphasis id="B96A8A04FF9DFF80FD0AD932E14BA63C" box="[752,878,296,320]" italics="true" pageId="7" pageNumber="566">N. agassizi</emphasis>
</taxonomicName>
, from which it differs by presence of both dark brown and yellow scales near forewing base (basal quarter of forewing in
<taxonomicName id="4C1E2D95FF9DFF80FC29D956E668A667" box="[979,1101,333,356]" class="Insecta" family="Adelidae" genus="Nemophora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="agassizi">
<emphasis id="B96A8A04FF9DFF80FC29D956E668A667" box="[979,1101,333,356]" italics="true" pageId="7" pageNumber="566">N. agassizi</emphasis>
</taxonomicName>
is monotonously silver grey).
</paragraph>
</subSubSection>
<subSubSection id="C304059DFF9DFF80FF3DD96BE16FA68B" box="[199,842,367,392]" pageId="7" pageNumber="566" type="etymology">
<paragraph id="8BA15616FF9DFF80FF3DD96BE16FA68B" blockId="7.[151,1437,151,1221]" box="[199,842,367,392]" pageId="7" pageNumber="566">
<emphasis id="B96A8A04FF9DFF80FF3DD96BE366A68B" bold="true" box="[199,323,368,392]" pageId="7" pageNumber="566">Etymology</emphasis>
. From humerus (Latin)—upper arm, shoulder.
</paragraph>
</subSubSection>
<subSubSection id="C304059DFF9DFF80FF3DD988E649A35B" pageId="7" pageNumber="566" type="description">
<paragraph id="8BA15616FF9DFF80FF3DD988E11BA43B" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="566">
<emphasis id="B96A8A04FF9DFF80FF3DD988E368A6AF" bold="true" box="[199,333,403,428]" pageId="7" pageNumber="566">Description</emphasis>
. Male (
<figureCitation id="13254A93FF9DFF80FE5FD98FE3D6A6AF" box="[421,499,404,429]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1853,1876]" captionTargetBox="[188,1397,193,1831]" captionTargetId="figure@3.[188,1398,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Adults of Nemophora kalshoveni species-group. 1, N. kalshoveni Diakonoff, 1951, male, from Cisaura; 2, N. humerella sp. nov., male, holotype, from Sangir Island; 3, N. agassizi sp. nov., female, holotype, from Wabag; 4, N. dohertyi sp. nov., male, holotype, from Fakfak; 5, N. bistrigata sp. nov., male, holotype, from Jayapura; 6, N. toxopeusi sp. nov., female, holotype, from Irian Jaya; 7, N. stenopterella sp. nov., female, holotype, from Irian Jaya; 8, labels of the holotype of N. kalshoveni Diakonoff. Scale: 2 mm." httpUri="https://zenodo.org/record/166210/files/figure.png" pageId="7" pageNumber="566">Figs. 2</figureCitation>
,
<figureCitation id="13254A93FF9DFF80FDFBD98FE038A6AF" box="[513,541,404,428]" captionStart="FIGURES 10 – 17" captionStartId="4.[151,264,1940,1963]" captionTargetBox="[175,1402,875,1912]" captionTargetId="figure@4.[173,1414,871,1919]" captionTargetPageId="4" captionText="FIGURES 10 – 17. Forewing pattern and venation of Nemophora kalshoveni species-group. 10, N. kalshoveni Diakonoff; 11, N. humerella sp. nov.; 12, N. agassizi sp. nov.; 13, N. dohertyi sp. nov.; 14, 16, N. bistrigata sp. nov.; 15, N. toxopeusi sp. nov.; 17 N. stenopterella sp. nov. Scale: 2 mm." httpUri="https://zenodo.org/record/166212/files/figure.png" pageId="7" pageNumber="566">11</figureCitation>
). FWL
<quantity id="4CE6FBF3FF9DFF80FD8CD98FE0EBA6AF" box="[630,718,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="7" pageNumber="566" unit="mm" value="4.8">4.8 mm</quantity>
; WLR 0.38. Vertex with sparse row of brown (medially) to grey (laterally) piliform scales along the occipital margin; frons silver grey. Eyes enlarged, touching each other occipitally; interocular index 1.2. PLB 0.6 × vertical eye diameter and 1.8 × scape, dark brown. Proboscis dark brown. Antenna 2.5 × FWL. Scape and basal 20–25 segments of flagellum dark brown; distal part of flagellum light brown to silver grey, with dark marks on ventral side of each segment. Tegula and thorax brown. Forewing grey; base of costa with small dark brown spot; basal half of forewing sparsely scattered with both dark brown and white scales (approximately in 1:1 ratio). Oblique stripe in the apical part of forewing straight, bright white, on both sides is bordered by narrow lines of dark brown scales; it reaches about 0.5 × forewing width; wing apex behind this oblique stripe densely suffused with both dark brown and white scales. Wing margin between the veins RS1 and M3 with white spot. Fringe brownish grey. Hindwing light brown; costal area grey; fringe light brown apically to light grey basally. Legs brown, apical parts of all tarsomeres and base of the first tarsomere white. Epiphysis at 0.5, reaching tip of tibia. Abdomen light brown, distal margins of both tergites and sternites white.
</paragraph>
<paragraph id="8BA15616FF9DFF80FF3DDB5FE3B5A45E" blockId="7.[151,1437,151,1221]" box="[199,400,836,861]" pageId="7" pageNumber="566">Female unknown.</paragraph>
<paragraph id="8BA15616FF9DFF80FF3DDB7CE649A35B" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="566">
Male genitalia (
<figureCitation id="13254A93FF9DFF80FE85DB7CE02BA483" box="[383,526,871,896]" captionStart="FIGURES 18 - 37" captionStartId="5.[151,264,1514,1537]" captionTargetBox="[169,1424,200,1474]" captionTargetId="figure@5.[151,1436,193,1493]" captionTargetPageId="5" captionText="FIGURES 18 - 37. Male genitalia of Nemophora kalshoveni species-group. 18 – 22, N. kalshoveni Diakonoff; 23 – 27, N. humerella sp. nov.; 28 – 32, N. dohertyi sp. nov.; 33 – 37, N. bistrigata sp. nov. 18, 23, 28, 33: genital complex, ventral view (right valva not shown); 19, 24, 29, 34: genital complex, lateral view; 20, 25, 30, 35: juxta; 21, 26, 31, 36: phallus, ventral view; 22, 27, 32, 37: phallus, lateral view. Scale: 0.5 mm." httpUri="https://zenodo.org/record/166213/files/figure.png" pageId="7" pageNumber="566">Figs. 23–27</figureCitation>
). Tegumen long and narrow, with W-shaped apex. Socius elongate, 1.6–1.8 × diameter of phallus. Vinculum 1.8 × length of valva, with almost straight lateral margins, widely rounded basally; distal margin of vinculum indistinct, fused to valvae. Apex of valva extends beyond the apex of tegumen. Valvae fused basally up to 0.55 × total length; their internal margins indistinct; dorsal margin of valva almost straight; apex of valva widely rounded. Anellus 0.55 × length of valva. Transtilla with small triangular medial process. Juxta 0.3 × length of phallus; arrow head very narrow (WLR 0.4), with widely rounded tip and without lateral arms. Phallus 1.1–1.2 × length of vinculum, very thin, almost straight; base of phallus anchor-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C304059DFF9DFF80FF3DDC78E0C0A37E" box="[199,741,1123,1149]" pageId="7" pageNumber="566" type="distribution">
<paragraph id="8BA15616FF9DFF80FF3DDC78E0C0A37E" blockId="7.[151,1437,151,1221]" box="[199,741,1123,1149]" pageId="7" pageNumber="566">
<emphasis id="B96A8A04FF9DFF80FF3DDC78E37CA37F" bold="true" box="[199,345,1123,1148]" pageId="7" pageNumber="566">Distribution</emphasis>
(
<figureCitation id="13254A93FF9DFF80FE92DC7FE388A37F" box="[360,429,1124,1149]" captionStart="FIGURE 9" captionStartId="4.[151,250,799,821]" captionTargetBox="[208,1376,208,762]" captionTargetId="figure@4.[199,1384,196,770]" captionTargetPageId="4" captionText="FIGURE 9. Distribution of Nemophora kalshoveni species-group." httpUri="https://zenodo.org/record/261896/files/figure.png" pageId="7" pageNumber="566">Fig. 9</figureCitation>
).
<collectingCountry id="F3091686FF9DFF80FE38DC7FE00AA37E" box="[450,559,1124,1149]" name="Indonesia" pageId="7" pageNumber="566">Indonesia</collectingCountry>
(Sangir
<collectingCountry id="F3091686FF9DFF80FD75DC7FE0FDA37E" box="[655,728,1124,1149]" name="Iceland" pageId="7" pageNumber="566">Island</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C304059DFF9DFF80FF3DDC9CE33EA3C6" pageId="7" pageNumber="566" type="discussion">
<paragraph id="8BA15616FF9DFF80FF3DDC9CE33EA3C6" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="566">
<emphasis id="B96A8A04FF9DFF80FF3DDC9CE369A3A3" bold="true" box="[199,332,1159,1184]" pageId="7" pageNumber="566">Comments.</emphasis>
The species’ name was proposed by Walsingham who labelled the holotype but never published the description.
</paragraph>
</subSubSection>
</treatment>
</document>
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