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<document id="612FAAD15C81D43632B5EC875E36B61C" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFE5FFD1BF14FF08085D039B" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus niger Cameron 1918" docType="treatment" docVersion="2" lastPageNumber="60" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="60" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A9F5CBA5BEFB6F9B71FBEF5036C829A4">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:namePart id="D089D7CD34D8FC46D60F03C5F818AA3E">Frisch, Johannes</mods:namePart>
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<mods:namePart id="2BCA89D1A96F19C6E6F2A8912D9A8074">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="35907CA396D30DC2598E99C98A11E9D0">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:date id="C20957DCD49C1A8BE4079249813113EC">2022</mods:date>
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<treatment id="03B7C77AFFE5FFD1BF14FF08085D039B" ID-DOI="http://doi.org/10.5281/zenodo.10724329" ID-Zenodo-Dep="10724329" LSID="urn:lsid:plazi:treatment:03B7C77AFFE5FFD1BF14FF08085D039B" httpUri="http://treatment.plazi.org/id/03B7C77AFFE5FFD1BF14FF08085D039B" lastPageNumber="60" pageId="37" pageNumber="60">
<subSubSection id="C30425E7FFE5FFD1BF14FF0809D10504" box="[847,1228,239,266]" pageId="37" pageNumber="60" type="nomenclature">
<paragraph id="8BA1766CFFE5FFD1BF14FF0809D10504" blockId="37.[847,1228,239,299]" box="[847,1228,239,266]" pageId="37" pageNumber="60">
<heading id="D0E9C100FFE5FFD1BF14FF0809D10504" box="[847,1228,239,266]" fontSize="11" level="2" pageId="37" pageNumber="60" reason="5">
<taxonomicName id="4C1E0DEFFFE5FFD1BF14FF0809D10504" ID-CoL="79YYG" authority="Cameron, 1918" authorityName="Cameron" authorityYear="1918" box="[847,1228,239,266]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96AAA7EFFE5FFD1BF14FF0809D10504" bold="true" box="[847,1228,239,266]" pageId="37" pageNumber="60">
<emphasis id="B96AAA7EFFE5FFD1BF14FF0809140507" bold="true" box="[847,1033,240,266]" italics="true" pageId="37" pageNumber="60">Scopaeus niger</emphasis>
<bibRefCitation id="EF8F0B9DFFE5FFD1B84BFF1709D10504" author="Cameron, M." box="[1040,1228,239,265]" pageId="37" pageNumber="60" pagination="58 - 90" refId="ref30971" refString="Cameron, M. (1918): New species of Staphylinidae from Singapore. Part 1. - Transactions of the Entomological Society of London 1918: 58 - 90." type="journal article" year="1918">Cameron, 1918</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE5FFD1BF14FEE80EF30526" box="[847,1006,272,299]" pageId="37" pageNumber="60" type="description">
<paragraph id="8BA1766CFFE5FFD1BF14FEE80EF30526" blockId="37.[847,1228,239,299]" box="[847,1006,272,299]" pageId="37" pageNumber="60">
(
<figureCitation id="13256AE9FFE5FFD1BF0CFEE80EAD0526" box="[855,944,272,299]" captionStart="Figures 2326" captionStartId="14.[151,224,1930,1952]" captionTargetBox="[263,1352,435,1899]" captionTargetId="figure-131@14.[151,1398,422,1911]" captionTargetPageId="14" captionText="Figures 2326. Habitus of Scopaeus cuspidatus, paratype, Jawa Barat: Mt Salak (23); S. sulawesianus, paratype, Sulawesi Tengah: Tambarana (24); S. niger, paralectotype, Singapore (25); S. borneensis, holotype, Kalimantan Barat: Pontianak (26)." figureDoi="http://doi.org/10.5281/zenodo.10724252" httpUri="https://zenodo.org/record/10724252/files/figure.png" pageId="37" pageNumber="60">Figs 25</figureCitation>
,
<figureCitation id="13256AE9FFE5FFD1BFE7FEE90EF80526" box="[956,997,273,299]" captionStart="Figures 179181" captionStartId="44.[151,224,925,947]" captionTargetBox="[230,1323,252,854]" captionTargetId="figure-384@44.[157,1398,225,897]" captionTargetPageId="44" captionText="Figures 179181. Sperm pump (179, 181) and bursal duct (180) of ♀ of S. niger, lectotype, Singapore (179); S. borneensis, holotype, Kalimantan Barat: Pontianak (180, with lateral gonocoxal plates; 181). Long scale bar: 179, 181; short scale bar: 180." figureDoi="http://doi.org/10.5281/zenodo.10724317" httpUri="https://zenodo.org/record/10724317/files/figure.png" pageId="37" pageNumber="60">179</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE5FFD1BF14FEAD09EC0561" box="[847,1265,340,364]" pageId="37" pageNumber="60" type="reference_group">
<paragraph id="8BA1766CFFE5FFD1BF14FEAD09EC0561" blockId="37.[847,1265,340,364]" box="[847,1265,340,364]" pageId="37" pageNumber="60">
<treatmentCitationGroup id="AB0E5142FFE5FFD1BF14FEAD09EC0561" box="[847,1265,340,364]" pageId="37" pageNumber="60">
<treatmentCitation id="0ABF507DFFE5FFD1BF14FEAD09D80561" author="Cameron, M." box="[847,1221,340,364]" page="77" pageId="37" pageNumber="60" year="1918">
<taxonomicName id="4C1E0DEFFFE5FFD1BF14FEAD09D80561" ID-CoL="79YYG" authority="Cameron, 1918: 77" authorityName="Cameron" authorityPageNumber="77" authorityYear="1918" box="[847,1221,340,364]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96AAA7EFFE5FFD1BF14FEAD0EE90561" box="[847,1012,341,364]" italics="true" pageId="37" pageNumber="60">Scopaeus niger</emphasis>
<bibRefCitation id="EF8F0B9DFFE5FFD1BFA0FEAC09D80561" author="Cameron, M." box="[1019,1221,340,364]" pageId="37" pageNumber="60" pagination="58 - 90" refId="ref30971" refString="Cameron, M. (1918): New species of Staphylinidae from Singapore. Part 1. - Transactions of the Entomological Society of London 1918: 58 - 90." type="journal article" year="1918">Cameron, 1918: 77</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 78.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE5FFD1BF14FE6E095F07F3" pageId="37" pageNumber="60" type="materials_examined">
<paragraph id="8BA1766CFFE5FFD1BF14FE6E092B0734" blockId="37.[824,1443,406,1942]" pageId="37" pageNumber="60">
<emphasis id="B96AAA7EFFE5FFD1BF14FE6E099B05A3" bold="true" box="[847,1158,406,430]" pageId="37" pageNumber="60">Type specimens examined:</emphasis>
<typeStatus id="54A5C8CEFFE5FFD1B8D4FE6E09E005A3" box="[1167,1277,406,430]" pageId="37" pageNumber="60" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D18BDE5FFE5FFD1B95DFE6E080405A0" box="[1286,1305,406,429]" count="1" pageId="37" pageNumber="60" type="female"></specimenCount>
,
<collectingCountry id="F30936FCFFE5FFD1B972FE6E088405A3" box="[1321,1433,406,430]" name="Singapore" pageId="37" pageNumber="60">Singapore</collectingCountry>
, Mandai, by present designation; labeled Type (round, printed label with red margin), Type (red, handwritten), Mandai, /
<collectingCountry id="F30936FCFFE5FFD1BFF6FE000900061D" box="[941,1053,504,528]" name="Singapore" pageId="37" pageNumber="60">Singapore</collectingCountry>
. / Dr. Cameron. (printed), Bank of Stream (printed), M. Cameron. / Bequest. / B.M.1955- 147. (printed),
<typeStatus id="54A5C8CEFFE5FFD1BFAAFDC20952065F" box="[1009,1103,570,594]" pageId="37" pageNumber="60" type="syntype">Syntype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFE5FFD1B82EFDC20806065F" authority="Cam." authorityName="Cameron" authorityYear="1918" box="[1141,1307,570,594]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">S. niger Cam.</taxonomicName>
/ (original det. label / missing? / M. T. O. Brendell.
<date id="FFA050ACFFE5FFD1B962FDA30885067E" box="[1337,1432,603,627]" pageId="37" pageNumber="60" value="2000-02">ii. 2000</date>
(handwritten);
<typeStatus id="54A5C8CEFFE5FFD1BFB9FD8409490699" box="[994,1108,636,660]" pageId="37" pageNumber="60" type="lectotype">Lectotype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFE5FFD1B82AFD8408010699" authorityName="Cameron" authorityYear="1918" box="[1137,1308,636,660]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96AAA7EFFE5FFD1B82AFD8408010699" box="[1137,1308,636,660]" italics="true" pageId="37" pageNumber="60">Scopaeus niger</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF8F0B9DFFE5FFD1B962FD840E7406B8" author="Cameron, M." pageId="37" pageNumber="60" pagination="58 - 90" refId="ref30971" refString="Cameron, M. (1918): New species of Staphylinidae from Singapore. Part 1. - Transactions of the Entomological Society of London 1918: 58 - 90." type="journal article" year="1918">Cameron 1918</bibRefCitation>
/ label by J.
<bibRefCitation id="EF8F0B9DFFE5FFD1BFA4FD65099106B8" author="Frisch, J. &amp; T. Mainda" box="[1023,1164,669,693]" pageId="37" pageNumber="60" pagination="139 - 147" refId="ref31869" refString="Frisch, J. &amp; T. Mainda (2022): The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species. - Soil Organisms 94 (3): 139 - 147." type="journal article" year="2022">Frisch, 2022</bibRefCitation>
(red, printed) (
<collectionCode id="ED0FEEA9FFE5FFD1B967FD65088A06B8" box="[1340,1431,669,693]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="37" pageNumber="60">NHML</collectionCode>
).
<typeStatus id="54A5C8CEFFE5FFD1BF63FD460EC606DB" box="[824,987,702,726]" pageId="37" pageNumber="60" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="9D18BDE5FFE5FFD1BFB7FD46090706D8" box="[1004,1050,702,726]" count="3" pageId="37" pageNumber="60" type="female">3 ♀</specimenCount>
, same labels as
<typeStatus id="54A5C8CEFFE5FFD1B8BEFD46085406DB" box="[1253,1353,702,726]" pageId="37" pageNumber="60" type="lectotype">lectotype</typeStatus>
except for type label; subsequent
<typeStatus id="54A5C8CEFFE5FFD1B82BFD2709D206FA" box="[1136,1231,735,759]" pageId="37" pageNumber="60" type="paratype">paratype</typeStatus>
label
<typeStatus id="54A5C8CEFFE5FFD1B978FD27089A06FA" box="[1315,1415,735,759]" pageId="37" pageNumber="60" type="paratype">Paratype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFE5FFD1BF63FCF80EC0071A" authorityName="Cameron" authorityYear="1918" box="[824,989,768,791]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96AAA7EFFE5FFD1BF63FCF80EC0071A" box="[824,989,768,791]" italics="true" pageId="37" pageNumber="60">Scopaeus niger</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF8F0B9DFFE5FFD1BFAAFCF809960715" author="Cameron, M." box="[1009,1163,768,792]" pageId="37" pageNumber="60" pagination="58 - 90" refId="ref30971" refString="Cameron, M. (1918): New species of Staphylinidae from Singapore. Part 1. - Transactions of the Entomological Society of London 1918: 58 - 90." type="journal article" year="1918">Cameron 1918</bibRefCitation>
/ label by J.
<bibRefCitation id="EF8F0B9DFFE5FFD1B954FCF808850715" author="Frisch, J. &amp; T. Mainda" box="[1295,1432,768,792]" pageId="37" pageNumber="60" pagination="139 - 147" refId="ref31869" refString="Frisch, J. &amp; T. Mainda (2022): The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species. - Soil Organisms 94 (3): 139 - 147." type="journal article" year="2022">Frisch, 2022</bibRefCitation>
(red, printed) (
<collectionCode id="ED0FEEA9FFE5FFD1BF8EFCD909370734" box="[981,1066,801,825]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="37" pageNumber="60">NHML</collectionCode>
).
</paragraph>
<paragraph id="8BA1766CFFE5FFD1BF14FCB9095F07F3" blockId="37.[824,1443,406,1942]" pageId="37" pageNumber="60">
<bibRefCitation id="EF8F0B9DFFE5FFD1BF14FCB909310754" author="Cameron, M." box="[847,1068,833,857]" pageId="37" pageNumber="60" pagination="58 - 90" refId="ref30971" refString="Cameron, M. (1918): New species of Staphylinidae from Singapore. Part 1. - Transactions of the Entomological Society of London 1918: 58 - 90." type="journal article" year="1918">Cameron (1918: 77</bibRefCitation>
, 78) described
<taxonomicName id="4C1E0DEFFFE5FFD1B8B7FCBA08810754" authorityName="Cameron" authorityYear="1918" box="[1260,1436,834,857]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96AAA7EFFE5FFD1B8B7FCBA08810754" box="[1260,1436,834,857]" italics="true" pageId="37" pageNumber="60">Scopaeus niger</emphasis>
</taxonomicName>
based on
<specimenCount id="9D18BDE5FFE5FFD1BFFEFC9A093B0777" box="[933,1062,866,890]" count="4" pageId="37" pageNumber="60" type="female">four female</specimenCount>
specimens without designating a
<typeStatus id="54A5C8CEFFE5FFD1BF63FC7B0E840796" box="[824,921,899,923]" pageId="37" pageNumber="60" type="holotype">holotype</typeStatus>
, which thus are
<typeStatus id="54A5C8CEFFE5FFD1B81FFC7C09B50791" box="[1092,1192,900,924]" pageId="37" pageNumber="60" type="syntype">syntypes</typeStatus>
. As no male
<typeStatus id="54A5C8CEFFE5FFD1B974FC7C08980791" box="[1327,1413,900,924]" pageId="37" pageNumber="60" type="syntype">syntype</typeStatus>
is available, I designate a female specimen as
<typeStatus id="54A5C8CEFFE5FFD1B941FC5C086307B1" box="[1306,1406,932,956]" pageId="37" pageNumber="60" type="lectotype">lectotype</typeStatus>
to stabilize the name
<taxonomicName id="4C1E0DEFFFE5FFD1B852FC3D081607D0" authority="Cameron, 1918" authorityName="Cameron" authorityYear="1918" box="[1033,1291,965,989]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96AAA7EFFE5FFD1B852FC3D094307D0" box="[1033,1118,965,989]" italics="true" pageId="37" pageNumber="60">S. niger</emphasis>
<bibRefCitation id="EF8F0B9DFFE5FFD1B83CFC3D081607D0" author="Cameron, M." box="[1127,1291,965,989]" pageId="37" pageNumber="60" pagination="58 - 90" refId="ref30971" refString="Cameron, M. (1918): New species of Staphylinidae from Singapore. Part 1. - Transactions of the Entomological Society of London 1918: 58 - 90." type="journal article" year="1918">Cameron, 1918</bibRefCitation>
</taxonomicName>
according to ICZN 1999, Article 74.1.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE5FFD1BF14FBFE0EA80339" pageId="37" pageNumber="60" type="description">
<paragraph id="8BA1766CFFE5FFD1BF14FBFE096B0206" blockId="37.[824,1443,406,1942]" pageId="37" pageNumber="60">
<emphasis id="B96AAA7EFFE5FFD1BF14FBFE0EEB0013" bold="true" box="[847,1014,1030,1054]" pageId="37" pageNumber="60">Redescription:</emphasis>
Habitus as in
<figureCitation id="13256AE9FFE5FFD1B8D0FBFF09C40013" box="[1163,1241,1030,1055]" captionStart="Figures 2326" captionStartId="14.[151,224,1930,1952]" captionTargetBox="[263,1352,435,1899]" captionTargetId="figure-131@14.[151,1398,422,1911]" captionTargetPageId="14" captionText="Figures 2326. Habitus of Scopaeus cuspidatus, paratype, Jawa Barat: Mt Salak (23); S. sulawesianus, paratype, Sulawesi Tengah: Tambarana (24); S. niger, paralectotype, Singapore (25); S. borneensis, holotype, Kalimantan Barat: Pontianak (26)." figureDoi="http://doi.org/10.5281/zenodo.10724252" httpUri="https://zenodo.org/record/10724252/files/figure.png" pageId="37" pageNumber="60">Fig. 25</figureCitation>
. Head subcircular. Penultimate antennal segment elongate. Mesotibia slender. Body surface subnitid with fine, dense, setose punctation; microreticulation absent. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body color of
<typeStatus id="54A5C8CEFFE5FFD1BFC3FB540EDB00C9" box="[920,966,1196,1220]" pageId="37" pageNumber="60">type</typeStatus>
specimens reddish medium brown to dark brown; maxillary palpi medium brown; antennae medium brown proximally, gradually lighter towards yellow-brown terminal segments; legs yellow-brown with light brown femora.
<typeStatus id="54A5C8CEFFE5FFD1BF80FAD7090D014A" box="[987,1040,1327,1351]" pageId="37" pageNumber="60">Type</typeStatus>
specimens probably bleached during last century, because
<bibRefCitation id="EF8F0B9DFFE5FFD1B862FAA8083B0165" author="Cameron, M." box="[1081,1318,1360,1384]" pageId="37" pageNumber="60" pagination="58 - 90" refId="ref30971" refString="Cameron, M. (1918): New species of Staphylinidae from Singapore. Part 1. - Transactions of the Entomological Society of London 1918: 58 - 90." type="journal article" year="1918">Cameron (1918: 77)</bibRefCitation>
described the coloration as follows: Black, moderately shining; antennae with first six segments pitchy-testaceous, the others clear testaceous; legs testaceous, the femora more or less infuscate. Total body length 3.4 4.0 mm; forebody length 2.0
<quantity id="4CE6DB89FFE5FFD1B846FA0C096F0206" box="[1053,1138,1524,1548]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="37" pageNumber="60" unit="mm" value="2.1">2.1 mm</quantity>
.
</paragraph>
<paragraph id="8BA1766CFFE5FFD1BF14F9ED091B0220" blockId="37.[824,1443,406,1942]" box="[847,1030,1557,1581]" pageId="37" pageNumber="60">Male: Unknown.</paragraph>
<paragraph id="8BA1766CFFE5FFD1BF14F9CE09E30282" blockId="37.[824,1443,406,1942]" pageId="37" pageNumber="60">
Female: Chamber segment of sperm pump short, circular, not projecting from large apopysis towards bursal duct (
<figureCitation id="13256AE9FFE5FFD1BFE5F98F090A0282" box="[958,1047,1655,1679]" captionStart="Figures 179181" captionStartId="44.[151,224,925,947]" captionTargetBox="[230,1323,252,854]" captionTargetId="figure-384@44.[157,1398,225,897]" captionTargetPageId="44" captionText="Figures 179181. Sperm pump (179, 181) and bursal duct (180) of ♀ of S. niger, lectotype, Singapore (179); S. borneensis, holotype, Kalimantan Barat: Pontianak (180, with lateral gonocoxal plates; 181). Long scale bar: 179, 181; short scale bar: 180." figureDoi="http://doi.org/10.5281/zenodo.10724317" httpUri="https://zenodo.org/record/10724317/files/figure.png" pageId="37" pageNumber="60">Fig. 179</figureCitation>
); bursa membranous.
</paragraph>
<paragraph id="8BA1766CFFE5FFD1BF14F9600EA80339" blockId="37.[824,1443,406,1942]" pageId="37" pageNumber="60">
<emphasis id="B96AAA7EFFE5FFD1BF14F9600ECD02BD" bold="true" box="[847,976,1688,1712]" pageId="37" pageNumber="60">Phylogeny:</emphasis>
Judging from the body shape with long, slender antennae and legs,
<taxonomicName id="4C1E0DEFFFE5FFD1B831F941080802DC" authorityName="Cameron" authorityYear="1918" box="[1130,1301,1721,1745]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96AAA7EFFE5FFD1B831F941080802DC" box="[1130,1301,1721,1745]" italics="true" pageId="37" pageNumber="60">Scopaeus niger</emphasis>
</taxonomicName>
is expected to belong to the
<taxonomicName id="4C1E0DEFFFE5FFD1BFAAF922097E02FF" authorityName="Oke" authorityYear="1933" box="[1009,1123,1754,1778]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96AAA7EFFE5FFD1BFAAF922097E02FF" box="[1009,1123,1754,1778]" italics="true" pageId="37" pageNumber="60">S. gracilis</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF8F0B9DFFE5FFD1B94EF9220E81031E" author="Frisch, J. &amp; D. Burckhardt &amp; V. Wolters" pageId="37" pageNumber="60" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt &amp; V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant &amp; Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity &amp; Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
). Male specimens are needed to confirm this hypothesis.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE5FFD1BF14F8C4085D039B" pageId="37" pageNumber="60" type="distribution">
<paragraph id="8BA1766CFFE5FFD1BF14F8C4085D039B" blockId="37.[824,1443,406,1942]" pageId="37" pageNumber="60">
<emphasis id="B96AAA7EFFE5FFD1BF14F8C40EF50359" bold="true" box="[847,1000,1852,1876]" pageId="37" pageNumber="60">Distribution:</emphasis>
<taxonomicName id="4C1E0DEFFFE5FFD1BFB5F8C5098A0359" authorityName="Cameron" authorityYear="1918" box="[1006,1175,1853,1876]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96AAA7EFFE5FFD1BFB5F8C5098A0359" box="[1006,1175,1853,1876]" italics="true" pageId="37" pageNumber="60">Scopaeus niger</emphasis>
</taxonomicName>
is hitherto known from
<collectingCountry id="F30936FCFFE5FFD1BF63F8A50EBA0378" box="[824,935,1885,1909]" name="Singapore" pageId="37" pageNumber="60">Singapore</collectingCountry>
only, but might be found in Sumatra as well, which is why it is included in this contribution.
</paragraph>
</subSubSection>
</treatment>
</document>