602 lines
56 KiB
XML
602 lines
56 KiB
XML
<document id="4D73E2B3C77835558E71B8EA94412D6F" ID-CLB-Dataset="29115" ID-DOI="10.11646/zootaxa.4455.1.3" ID-GBIF-Dataset="0d838ca1-52f1-4278-895d-68fc481b7558" ID-ISSN="1175-5326" ID-Zenodo-Dep="1456971" ID-ZooBank="2EA2EE4A-2BA4-438A-A1CB-D9C54F37DC78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539260480029" checkinUser="plazi" docAuthor="Wüster, Wolfgang, Chirio, Laurent, Trape, Jean-François, Ineich, Ivan, Jackson, Kate, Greenbaum, Eli, Barron, Cesar, Kusamba, Chifundera, Nagy, Zoltán T., Storey, Richard, Hall, Cara, Wüster, Catharine E., Barlow, Axel & Broadley, Donald G." docDate="2018" docId="03ADDC33D47DFA69FF5935B8D015FD82" docLanguage="en" docName="zootaxa.4455.1.3.pdf" docOrigin="Zootaxa 4455 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naja (Boulengerina) melanoleuca HAllOwEll 1857" docType="treatment" docVersion="7" masterDocId="FF94A44BD472FA78FFCE3743D41AFF95" masterDocTitle="Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae)" updateTime="1698575068667" updateUser="ExternalLinkService">
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<mods:title id="1FAC2D7F1F30157C024A9C9393EB12A8">Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae)</mods:title>
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<mods:namePart id="5CF06F0E123422A795BEF60AEC2083A2">Wüster, Wolfgang</mods:namePart>
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<mods:namePart id="45DF3CC96F79401D533B86CB554158B0">Chirio, Laurent</mods:namePart>
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<mods:namePart id="1AA3AB2D5A92D0F4EC8903DAA72D2DCF">Trape, Jean-François</mods:namePart>
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<mods:namePart id="5C29B291A72D7E13C0B28BBDE28887C3">Ineich, Ivan</mods:namePart>
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<mods:namePart id="91CC147E8FCA1F6B124E4004591EDE74">Jackson, Kate</mods:namePart>
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<mods:namePart id="2BFD9A19517626222AC8C7C00B343835">Greenbaum, Eli</mods:namePart>
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<mods:namePart id="03EF2B842C8107C9EAB03B12B9EBD61E">Barron, Cesar</mods:namePart>
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<mods:namePart id="F84040394A402A8B46FCF7A12E947159">Kusamba, Chifundera</mods:namePart>
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<mods:namePart id="26A3287607BFD2633359DE2437098978">Nagy, Zoltán T.</mods:namePart>
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<mods:namePart id="428AAF9725808F4A797378E2D70D43C3">Storey, Richard</mods:namePart>
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<mods:namePart id="88D1977D5C31AC6DFD3D8B312EA15B8E">Hall, Cara</mods:namePart>
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<mods:namePart id="D0676CEDD66C205AEE3C89FD955FA388">Wüster, Catharine E.</mods:namePart>
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<mods:namePart id="37F834F40F6A7E3665D37D456AD8980D">Barlow, Axel</mods:namePart>
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<mods:namePart id="F096734896C367EFF28BCAA11BE6827B">Broadley, Donald G.</mods:namePart>
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<mods:typeOfResource id="91E688323A537161C95B537B5517BB35">text</mods:typeOfResource>
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<mods:title id="A56709A93923016B905D68C2DD0DAD81">Zootaxa</mods:title>
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<mods:part id="00958AF84E6C9AADE18F9721DF4AE7BC">
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<mods:date id="B0BBB95EDF7EF377865B2C4D4CACBFF6">2018</mods:date>
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<mods:number id="004970450E07E4BC8DFDFBF499D22A72">2018-08-01</mods:number>
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<mods:number id="40BFF95EF151ADCCAFA5C0E7DAA06FF3">4455</mods:number>
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<mods:number id="CB82C71BEA781752DF225BA19917A798">1</mods:number>
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<mods:classification id="055FC6D3F87A6325D74A478E428F7A96">journal volume</mods:classification>
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<mods:identifier id="797C6C7E2B81B55F73CEEAB68DD0859C" type="DOI">10.11646/zootaxa.4455.1.3</mods:identifier>
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<mods:identifier id="F3A996282DA45560E61012143E0E6C10" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C8E6B395F5AEE1D3020A4912793049AD" type="Zenodo-Dep">1456971</mods:identifier>
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<treatment id="03ADDC33D47DFA69FF5935B8D015FD82" ID-DOI="http://doi.org/10.5281/zenodo.5950784" ID-GBIF-Taxon="148691100" ID-Zenodo-Dep="5950784" LSID="urn:lsid:plazi:treatment:03ADDC33D47DFA69FF5935B8D015FD82" httpUri="http://treatment.plazi.org/id/03ADDC33D47DFA69FF5935B8D015FD82" lastPageId="17" pageId="15">
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<subSubSection id="C31E3EAED47DFA77FF5935B8D6E6FC83" box="[151,764,763,790]" pageId="15" type="nomenclature">
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<paragraph id="8BBB6D25D47DFA77FF5935B8D6E6FC83" blockId="15.[151,764,763,790]" box="[151,764,763,790]" pageId="15">
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<heading id="D0F3DA49D47DFA77FF5935B8D6E6FC83" bold="true" box="[151,764,763,790]" fontSize="11" level="1" pageId="15" reason="1">
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<taxonomicName id="4C0416A6D47DFA77FF5935B8D6E6FC83" authority="Hallowell, 1857" authorityName="HAllOwEll" authorityYear="1857" box="[151,764,763,790]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="species" species="melanoleuca" subGenus="Boulengerina">
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<emphasis id="B970B137D47DFA77FF5935B8D6E6FC83" bold="true" box="[151,764,763,790]" pageId="15">
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<emphasis id="B970B137D47DFA77FF5935B8D4C8FC80" bold="true" box="[151,210,763,789]" italics="true" pageId="15">Naja</emphasis>
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(
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<emphasis id="B970B137D47DFA77FF2D35B8D593FC80" bold="true" box="[227,393,763,789]" italics="true" pageId="15">Boulengerina</emphasis>
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)
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<emphasis id="B970B137D47DFA77FE5435B8D62FFC80" bold="true" box="[410,565,763,789]" italics="true" pageId="15">melanoleuca</emphasis>
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<bibRefCitation id="EF9510D4D47DFA77FDF235BFD6E6FC83" box="[572,764,764,790]" pageId="15" refId="ref13793" refString="Hallowell, E. (1857) Notes of a collection of reptiles from the Gaboon country, West Africa, recently presented to the Academy of Natural Sciences of Philadelphia, by Dr. Henry A. Ford. Proceedings of Academy of Natural Sciences, Philadelphia, 9, 48 - 72." type="journal article">Hallowell, 1857</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E3EAED47DFA77FF59347ED773FC59" pageId="15" type="reference_group">
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<paragraph id="8BBB6D25D47DFA77FF59347ED596FCE5" blockId="15.[151,1436,828,1187]" pageId="15">
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<materialsCitation id="3B6C6778D47DFA77FF59347ED596FCE5" ID-GBIF-Occurrence="1917075145" collectionCode="ANSP" country="Gabon" location="Gabon" pageId="15" specimenCode="ANSP 6875-76, 6878-79" specimenCount="2" typeStatus="syntype">
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<taxonomicName id="4C0416A6D47DFA77FF59347ED67FFCC7" authority="Hallowell, 1857" authorityName="Hallowell" authorityYear="1857" box="[151,613,828,850]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="variety" species="haie" variety="melanoleuca">
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<emphasis id="B970B137D47DFA77FF59347ED4E7FCC7" box="[151,253,828,850]" italics="true" pageId="15">Naia haie</emphasis>
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var.
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<emphasis id="B970B137D47DFA77FEFD347FD5AEFCC7" box="[307,436,828,850]" italics="true" pageId="15">melanoleuca</emphasis>
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<bibRefCitation id="EF9510D4D47DFA77FE73347FD67FFCC7" box="[445,613,828,850]" pageId="15" refId="ref13793" refString="Hallowell, E. (1857) Notes of a collection of reptiles from the Gaboon country, West Africa, recently presented to the Academy of Natural Sciences of Philadelphia, by Dr. Henry A. Ford. Proceedings of Academy of Natural Sciences, Philadelphia, 9, 48 - 72." type="journal article">Hallowell, 1857</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="B970B137D47DFA77FDBC347ED7C3FCC7" box="[626,985,828,850]" italics="true" pageId="15">Proc. Acad. Nat. Sci. Philadelphia</emphasis>
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: 61.
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<typeStatus id="54BFD387D47DFA77FBDE347ED058FCC7" box="[1040,1090,829,850]" pageId="15">Type</typeStatus>
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locality:
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<collectingCountry id="F3132DB5D47DFA77FB69347FD0F7FCC7" box="[1191,1261,828,850]" name="Gabon" pageId="15">Gabon</collectingCountry>
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,
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<typeStatus id="54BFD387D47DFA77FB34347ED149FCC7" box="[1274,1363,829,850]" pageId="15" type="syntype">syntypes</typeStatus>
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<specimenCode id="DBA2C55ED47DFA77FA95347FD538FCE5" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="15">ANSP 6875–76</specimenCode>
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,
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<specimenCode id="DBA2C55ED47DFA77FEE23419D592FCE5" box="[300,392,858,880]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="15">6878–79</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BBB6D25D47DFA77FF593439D6EAFC3B" blockId="15.[151,1436,828,1187]" pageId="15">
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<materialsCitation id="3B6C6778D47DFA77FF593439D6F6FC3B" ID-GBIF-Occurrence="1917075149" collectionCode="ZMH" country="Cameroon" location="Hamburg" pageId="15" specimenCode="ZMH 4280, 7048, 7299-7302" specimenCount="2" stateProvince="Hamburg" typeStatus="syntype">
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<taxonomicName id="4C0416A6D47DFA77FF593439D631FC1A" authority="Fischer, 1885" authorityName="FischEr" authorityYear="1885" box="[151,555,889,911]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="variety" species="haje" variety="leucosticta">
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<emphasis id="B970B137D47DFA77FF593439D583FC1A" box="[151,409,889,911]" italics="true" pageId="15">Naja haje var. leucosticta</emphasis>
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<bibRefCitation id="EF9510D4D47DFA77FE6E343AD631FC1A" box="[416,555,889,911]" pageId="15" refId="ref13603" refString="Fischer, J. G. (1885) Ichthyologische und herpetologische Bemerkungen. V. Herpetologische Bemerkungen. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 2, 82 - 121." type="journal article">Fischer, 1885</bibRefCitation>
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</taxonomicName>
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,
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||
<emphasis id="B970B137D47DFA77FDFB343AD753FC1A" box="[565,841,889,911]" italics="true" pageId="15">
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||
Jahr.
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<collectingRegion id="49C0A3C7D47DFA77FDA3343AD6CAFC1A" box="[621,720,889,911]" country="Germany" name="Hamburg" pageId="15">Hamburg</collectingRegion>
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||
. Wiss. Anst.
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||
</emphasis>
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||
<emphasis id="B970B137D47DFA77FC80343AD740FC1A" bold="true" box="[846,858,889,911]" pageId="15">2</emphasis>
|
||
: 115, pl. v, fig. 11.
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<typeStatus id="54BFD387D47DFA77FBD13439D04BFC1A" box="[1055,1105,890,911]" pageId="15">Type</typeStatus>
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locality:
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<collectingCountry id="F3132DB5D47DFA77FB7F343AD100FC1A" box="[1201,1306,889,911]" name="Cameroon" pageId="15">Cameroon</collectingCountry>
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and Ogooué
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<collectingRegion id="49C0A3C7D47DFA77FF0934DBD51BFC3B" box="[199,257,920,942]" country="Nigeria" name="Rivers" pageId="15">River</collectingRegion>
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,
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<collectingCountry id="F3132DB5D47DFA77FEC534DBD54BFC3B" box="[267,337,920,942]" name="Gabon" pageId="15">Gabon</collectingCountry>
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||
,
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||
<typeStatus id="54BFD387D47DFA77FE9534DAD5AEFC3B" box="[347,436,921,942]" pageId="15" type="syntype">syntypes</typeStatus>
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<specimenCode id="DBA2C55ED47DFA77FE7434DAD637FC3B" box="[442,557,920,942]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="15">ZMH 4280</specimenCode>
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,
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||
<specimenCode id="DBA2C55ED47DFA77FDF634DBD677FC3B" box="[568,621,920,942]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="15">7048</specimenCode>
|
||
,
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||
<specimenCode id="DBA2C55ED47DFA77FDB934DBD6F6FC3B" box="[631,748,920,942]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="15">7299–7302</specimenCode>
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||
</materialsCitation>
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.
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||
</paragraph>
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<paragraph id="8BBB6D25D47DFA77FF5934FBD773FC59" blockId="15.[151,1436,828,1187]" box="[151,873,950,973]" pageId="15">
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<treatmentCitationGroup id="AB144A0BD47DFA77FF5934FBD773FC59" box="[151,873,950,973]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF5934FBD542FC59" box="[151,344,950,973]" italics="true" pageId="15">
|
||
<taxonomicName id="4C0416A6D47DFA77FF5934FBD54BFC59" authorityName="Hallowell" authorityYear="1857" box="[151,337,950,973]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="species" species="melanoleuca">Naia melanoleuca</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
<treatmentCitation id="0AA54B34D47DFA77FE9034F5D620FC59" author="" box="[350,570,950,972]" page="376" pageId="15">
|
||
<bibRefCitation id="EF9510D4D47DFA77FE9034F5D613FC59" box="[350,521,950,972]" pageId="15" refId="ref12241" refString="Boulenger, G. A. (1896) Catalogue of the Snakes in the British Museum (Natural History) III. British Museum (Natural History), London, 727 pp. https: // doi. org / 10.5962 / bhl. title. 54273" type="book">Boulenger, 1896</bibRefCitation>
|
||
: 376
|
||
</treatmentCitation>
|
||
(part, vars.
|
||
<collectingRegion id="49C0A3C7D47DFA77FD7D34F4D6DFFC59" box="[691,709,951,972]" country="Nigeria" name="Rivers" pageId="15">A</collectingRegion>
|
||
[a,b] & B [a,b])
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47DFA77FF593495D6A6FBBD" pageId="15" type="description">
|
||
<paragraph id="8BBB6D25D47DFA77FF593495D750FC7E" blockId="15.[151,1436,828,1187]" box="[151,842,981,1003]" pageId="15">
|
||
<treatmentCitationGroup id="AB144A0BD47DFA77FF593495D750FC7E" box="[151,842,981,1003]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF593495D5C5FC7E" box="[151,479,981,1003]" italics="true" pageId="15">
|
||
<taxonomicName id="4C0416A6D47DFA77FF593495D5C3FC7E" authority="melanoleuca" box="[151,473,981,1003]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="subSpecies" species="melanoleuca" subSpecies="melanoleuca">Naja melanoleuca melanoleuca</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
<treatmentCitation id="0AA54B34D47DFA77FE283495D6B0FC7E" author="" box="[486,682,981,1003]" page="290" pageId="15">
|
||
<bibRefCitation id="EF9510D4D47DFA77FE283495D662FC7E" box="[486,632,981,1003]" pageId="15" refId="ref14506" refString="Laurent, R. F. (1956) Contribution a l'herpetologie de la region des Grands Lacs de l'Afrique centrale. I. Generalites. II. Cheloniens. III. Ophidiens. Annales du Musee royal du Congo Belge, 48, 1 - 390." type="journal article">Laurent, 1956</bibRefCitation>
|
||
: 290
|
||
</treatmentCitation>
|
||
, pl. xxvi, fig. 2.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47DFA77FF5934B6D6CBFB9F" blockId="15.[151,1436,828,1187]" box="[151,721,1012,1034]" pageId="15">
|
||
<taxonomicName id="4C0416A6D47DFA77FF5934B6D5E8FB9F" authorityName="HAllOwEll" authorityYear="1857" box="[151,498,1012,1034]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="species" species="melanoleuca" subGenus="Boulengerina">
|
||
<emphasis id="B970B137D47DFA77FF5934B6D4D3FB9F" box="[151,201,1013,1034]" italics="true" pageId="15">Naja</emphasis>
|
||
(
|
||
<emphasis id="B970B137D47DFA77FF1934B7D57BFB9F" box="[215,353,1012,1034]" italics="true" pageId="15">Boulengerina</emphasis>
|
||
)
|
||
<emphasis id="B970B137D47DFA77FEA134B7D5E8FB9F" box="[367,498,1012,1034]" italics="true" pageId="15">melanoleuca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B970B137D47DFA77FE3D34B7D5E3FB9F" box="[499,505,1012,1034]" italics="true" pageId="15">;</emphasis>
|
||
<bibRefCitation id="EF9510D4D47DFA77FE3134B7D6D7FB9F" box="[511,717,1012,1034]" pageId="15" refId="ref16705" refString="Wallach, V., Wuster, W. & Broadley, D. G. (2009) In praise of subgenera: taxonomic status of cobras of the genus Naja Laurenti (Serpentes: Elapidae). Zootaxa, 2236, 26 - 36." type="journal article">
|
||
Wallach
|
||
<emphasis id="B970B137D47DFA77FD9634B6D691FB9F" box="[600,651,1012,1034]" italics="true" pageId="15">et al.</emphasis>
|
||
, 2009
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47DFA77FF593351D6A6FBBD" blockId="15.[151,1436,828,1187]" box="[151,700,1042,1064]" pageId="15">
|
||
<treatmentCitationGroup id="AB144A0BD47DFA77FF593351D6A6FBBD" box="[151,700,1042,1064]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF593351D5AAFBBD" box="[151,432,1042,1064]" italics="true" pageId="15">
|
||
<taxonomicName id="4C0416A6D47DFA77FF593351D5B0FBBD" box="[151,426,1042,1064]" class="Reptilia" family="Elapidae" genus="Boulengerina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="species" species="melanoleuca">Boulengerina melanoleuca</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
<treatmentCitation id="0AA54B34D47DFA77FE793351D6A2FBBD" author="" box="[439,696,1042,1064]" page="122" pageId="15">
|
||
<bibRefCitation id="EF9510D4D47DFA77FE793351D69FFBBD" box="[439,645,1042,1064]" pageId="15" refId="ref16653" refString="Wallach, V., Williams, K. L. & Boundy, J. (2014) Snakes of the World. A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, 1227 pp. https: // doi. org / 10.1201 / b 16901" type="book">
|
||
Wallach
|
||
<emphasis id="B970B137D47DFA77FDDE3350D659FBBD" box="[528,579,1042,1064]" italics="true" pageId="15">et al.</emphasis>
|
||
, 2014
|
||
</bibRefCitation>
|
||
: 122
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47DFA77FF593372D766FB36" pageId="15" type="reference_group">
|
||
<paragraph id="8BBB6D25D47DFA77FF593372D766FB36" blockId="15.[151,1436,828,1187]" pageId="15">
|
||
<treatmentCitationGroup id="AB144A0BD47DFA77FF593372D766FB36" infered="true" pageId="15">
|
||
<taxonomicName id="4C0416A6D47DFA77FF593372D636FBD2" authority="Sauvage, 1884: 204" authorityName="Sauvage" authorityPageNumber="204" authorityYear="1884" box="[151,556,1073,1095]" class="Reptilia" family="Elapidae" genus="Aspidelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="species" species="bocagei">
|
||
<emphasis id="B970B137D47DFA77FF593372D547FBD2" box="[151,349,1073,1095]" italics="true" pageId="15">Aspidelaps bocagei</emphasis>
|
||
<treatmentCitation id="0AA54B34D47DFA77FEAD3372D636FBD2" author="" box="[355,556,1073,1095]" page="204" pageId="15">
|
||
<bibRefCitation id="EF9510D4D47DFA77FEAD3372D5E6FBD2" box="[355,508,1073,1095]" pageId="15" refId="ref15687" refString="Sauvage, H. E. (1884) Notice sur une collection de reptiles et de poissons recueillie a Majumba, Congo. Bulletin de la Societe Zoologique de France, 9, 199 - 204." type="journal article">Sauvage, 1884</bibRefCitation>
|
||
: 204
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(type locality: Gabon and Majumba; holotype MNHN 1884.0015) has been listed as a synonym
|
||
<taxonomicName id="4C0416A6D47DFA77FEE53313D5FDFBF3" box="[299,487,1104,1126]" class="Reptilia" family="Elapidae" genus="Boulengerina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="subSpecies" species="melanoleuca" subSpecies="melanoleuca">
|
||
of
|
||
<emphasis id="B970B137D47DFA77FE893312D5FDFBF3" box="[327,487,1104,1126]" italics="true" pageId="15">N. melanoleuca</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="EF9510D4D47DFA77FDE93313D6DCFBF3" box="[551,710,1104,1126]" pageId="15" refId="ref12364" refString="Broadley, D. G. (1983) FitzSimons' Snakes of Southern Africa. Delta Books, Johannesburg, 376 pp." type="book">Broadley, 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9510D4D47DFA77FD1F3313D7BBFBF3" box="[721,929,1104,1126]" pageId="15" refId="ref16653" refString="Wallach, V., Williams, K. L. & Boundy, J. (2014) Snakes of the World. A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, 1227 pp. https: // doi. org / 10.1201 / b 16901" type="book">
|
||
Wallach
|
||
<emphasis id="B970B137D47DFA77FCE43312D744FBF3" box="[810,862,1104,1126]" italics="true" pageId="15">et al.</emphasis>
|
||
, 2014
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9510D4D47DFA77FC653313D062FBF3" box="[939,1144,1104,1126]" pageId="15" refId="ref12835" refString="Ceriaco, L. M. P., Marques, M. P., Schmitz, A. & Bauer, A. M. (2017) The " Cobra-preta " of Sao Tome Island, Gulf of Guinea, is a new species of Naja Laurenti, 1768 (Squamata: Elapidae). Zootaxa, 4324 (1), 121 - 141. https: // doi. org / 10.11646 / zootaxa. 4324.1" type="journal article">
|
||
Ceríaco
|
||
<emphasis id="B970B137D47DFA77FBCF3312D02FFBF3" box="[1025,1077,1104,1126]" italics="true" pageId="15">et al.</emphasis>
|
||
, 2017
|
||
</bibRefCitation>
|
||
), but is in fact a synonym of
|
||
<taxonomicName id="4C0416A6D47DFA77FF093333D542FB11" box="[199,344,1134,1157]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="species" species="annulata">
|
||
<emphasis id="B970B137D47DFA77FF093333D542FB11" box="[199,344,1134,1157]" italics="true" pageId="15">Naja annulata</emphasis>
|
||
</taxonomicName>
|
||
, as is evident from Sauvage’s description, which notes approximately 20 dark double bands along the entire body length, and 21 mid-dorsal scale rows (see also
|
||
<bibRefCitation id="EF9510D4D47DFA77FD1733CED76AFB36" box="[729,880,1165,1187]" pageId="15" refId="ref15725" refString="Schmidt, K. P. (1923) Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909 - 1915. Part II. Snakes. Bulletin of the American Museum of Natural History, 49, 1 - 146." type="journal article">Schmidt, 1923</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47DFA77FF59338AD076FA93" pageId="15" type="diagnosis">
|
||
<paragraph id="8BBB6D25D47DFA77FF59338AD076FA93" blockId="15.[151,1436,1225,1430]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF59338AD509FB77" bold="true" box="[151,275,1225,1250]" pageId="15">Diagnosis:</emphasis>
|
||
DOrsum blAck, OftEN with 1–3 sEmidividEd yEllOw crOssbANds ON thE NEck, thE first mAy bE AN OcEllus; vENtEr yEllOw with 4
|
||
<collectingRegion id="49C0A3C7D47DFA77FE4B33ADD581FA93" box="[389,411,1262,1286]" country="South Sudan" name="Lakes" pageId="15">tO</collectingRegion>
|
||
6 blAck bANds iN thE first 100 vENtrAls, thErEAftEr uNifOrm blAck.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47DFA69FF093252D75CFEF0" lastPageId="17" pageId="15" type="description">
|
||
<paragraph id="8BBB6D25D47DFA77FF093252D6A6FA00" blockId="15.[151,1436,1225,1430]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF093252D55BFABF" bold="true" box="[199,321,1297,1322]" pageId="15">Variation:</emphasis>
|
||
DOrsAl scAlE rOws ON NEck 19–27, At midbOdy 19 (vEry rArEly 17 Or 21); vENtrAls 209–230; subcAudAls 59–74 (
|
||
<tableCitation id="C686589ED47DFA77FEA13276D5D2FADB" box="[367,456,1333,1358]" captionStart="TABLE 8" captionStartId="16.[147,224,712,735]" captionText="TABLE 8. Variation in meristic characters in the species of the N. melanoleuca complex." pageId="15">TAblE 8</tableCitation>
|
||
). SuprAlAbiAls 7, thE third ANd fOurth ENtEriNg thE Orbit; iNfrAlAbiAls 8, thE first fOur iN cONtAct with thE ANtEriOr subliNguAls,
|
||
<collectingRegion id="49C0A3C7D47DFA77FD8E3219D646FAE7" box="[576,604,1370,1394]" country="Cameroon" name="North" pageId="15">NO</collectingRegion>
|
||
cuNEAtE; prEOculAr 1; pOstOculArs 3 (vEry rArEly 2 Or 4); tEmpOrAls 1+2 Or 1+3; NuchAls bOrdEriNg tEmpOrAls 5–9, usuAlly 7.
|
||
</paragraph>
|
||
<caption id="DF7B3DADD47DFA77FF5930E0D7CCF863" httpUri="https://zenodo.org/record/1456987/files/figure.png" pageId="15" startId="15.[151,250,1955,1977]" targetBox="[151,1436,1485,1930]" targetPageId="15">
|
||
<paragraph id="8BBB6D25D47DFA77FF5930E0D7CCF863" blockId="15.[151,1436,1955,2038]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF5930E0D50BF82C" bold="true" box="[151,273,1955,1977]" pageId="15">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="4C0416A6D47DFA77FEEC30E7D693F82C" authorityName="HAllOwEll" authorityYear="1857" box="[290,649,1955,1977]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="species" species="melanoleuca" subGenus="Boulengerina">
|
||
<emphasis id="B970B137D47DFA77FEEC30E7D549F82C" box="[290,339,1956,1977]" italics="true" pageId="15">Naja</emphasis>
|
||
(
|
||
<emphasis id="B970B137D47DFA77FEA930E0D5EBF82C" box="[359,497,1955,1977]" italics="true" pageId="15">Boulengerina</emphasis>
|
||
)
|
||
<emphasis id="B970B137D47DFA77FDCA30E0D693F82C" box="[516,649,1955,1977]" italics="true" pageId="15">melanoleuca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B970B137D47DFA77FD4730E0D697F82C" box="[649,653,1955,1977]" italics="true" pageId="15">.</emphasis>
|
||
Adult specimens from Yaoundé, Cameroon (left—photo J.-F. Trape) and Tsibilé, Gabon (right—photo L. Chirio). Note the diffuse but distinct hood mark that is often present in this species, and the combination of broad main bands and narrow accessory bands on the ventral side.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7B3DADD462FA68FF5D358BD7DCFD4A" box="[147,966,711,735]" pageId="16">
|
||
<paragraph id="8BBB6D25D462FA68FF5D358BD7DCFD4A" blockId="16.[147,966,711,735]" box="[147,966,711,735]" pageId="16">
|
||
<emphasis id="B970B137D462FA68FF5D358BD4E2FD4A" bold="true" box="[147,248,712,735]" pageId="16">TABLE 8.</emphasis>
|
||
Variation in meristic characters in the species of the
|
||
<taxonomicName id="4C0416A6D462FA68FD13358BD776FD4A" authorityName="HAllOwEll" authorityYear="1857" box="[733,876,711,735]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" phylum="Chordata" rank="species" species="melanoleuca">
|
||
<emphasis id="B970B137D462FA68FD13358BD776FD4A" box="[733,876,711,735]" italics="true" pageId="16">N. melanoleuca</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBB6D25D462FA68FE2E35BED359FA31" pageId="16">
|
||
<table id="F9049F85D4620587FF5D35BFD341FA57" box="[147,1883,764,1474]" gridcols="11" gridrows="12" pageId="16">
|
||
<tr id="35346F67D4620587FF5D35BFD341FCC5" box="[147,1883,764,848]" gridrow="0" pageId="16" rowspan-0="1" rowspan-2="1">
|
||
<th id="76E5061BD4620587FE2E35BFD668FCC5" box="[480,626,764,848]" gridcol="1" gridrow="0" pageId="16">
|
||
<emphasis id="B970B137D462FA68FE2E35BED5F5FCA7" italics="true" pageId="16">
|
||
<taxonomicName id="4C0416A6D462FA68FE2E35BED668FC81" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[480,626,765,788]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" phylum="Chordata" rank="species" species="guineensis">Naja guineensis</taxonomicName>
|
||
N
|
||
</emphasis>
|
||
=71
|
||
</th>
|
||
<th id="76E5061BD4620587FCD635BFD7E2FCC5" box="[792,1016,764,848]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="16">
|
||
<emphasis id="B970B137D462FA68FCD635BED73DFCA7" italics="true" pageId="16">
|
||
<taxonomicName id="4C0416A6D462FA68FCD635BED7A5FC81" authorityName="HAllOwEll" authorityYear="1857" box="[792,959,764,788]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" phylum="Chordata" rank="species" species="melanoleuca">Naja melanoleuca</taxonomicName>
|
||
N
|
||
</emphasis>
|
||
=307
|
||
</th>
|
||
<th id="76E5061BD4620587FBD435BFD110FCC5" box="[1050,1290,764,848]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="16">
|
||
<emphasis id="B970B137D462FA68FBD435BED033FCA7" italics="true" pageId="16">
|
||
<taxonomicName id="4C0416A6D462FA68FBD435BED0B0FC81" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[1050,1194,764,788]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" phylum="Chordata" rank="species" species="savannula">Naja savannula</taxonomicName>
|
||
N
|
||
</emphasis>
|
||
=24
|
||
</th>
|
||
<th id="76E5061BD4620587FAF135BFD224FCC5" box="[1343,1598,764,848]" colspan="2" colspanRight="1" gridcol="7" gridrow="0" pageId="16">
|
||
<taxonomicName id="4C0416A6D462FA68FAF135BED1A4FC81" authorityName="LAurENt" authorityYear="1955" box="[1343,1470,764,788]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" phylum="Chordata" rank="species" species="subfulva">
|
||
<emphasis id="B970B137D462FA68FAF135BED1A4FC81" box="[1343,1470,764,788]" italics="true" pageId="16">Naja subfulva</emphasis>
|
||
</taxonomicName>
|
||
(excl. eastern and southern African pops.)
|
||
<emphasis id="B970B137D462FA68FAF1347AD154FCC5" box="[1343,1358,825,848]" italics="true" pageId="16">N</emphasis>
|
||
=202
|
||
</th>
|
||
<th id="76E5061BD4620587F9A935BFD341FCC5" box="[1639,1883,764,848]" colspan="2" colspanRight="1" gridcol="9" gridrow="0" pageId="16">
|
||
<emphasis id="B970B137D462FA68F9A935BED26CFCA7" italics="true" pageId="16">
|
||
<taxonomicName id="4C0416A6D462FA68F9A935BED30BFC81" box="[1639,1809,764,788]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" phylum="Chordata" rank="species" species="peroescobari">Naja peroescobari</taxonomicName>
|
||
N
|
||
</emphasis>
|
||
= 8
|
||
</th>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D3435D341FC18" box="[147,1883,886,909]" gridrow="1" pageId="16" rowspan-0="1">
|
||
<td id="76E5061BD4620587FE2E3435D668FC18" box="[480,626,886,909]" gridcol="1" gridrow="1" pageId="16">Mean ± 1SD</td>
|
||
<td id="76E5061BD4620587FD473435D6C0FC18" box="[649,730,886,909]" gridcol="2" gridrow="1" pageId="16">Range</td>
|
||
<td id="76E5061BD4620587FCD63435D797FC18" box="[792,909,886,909]" gridcol="3" gridrow="1" pageId="16">Mean ± 1SD</td>
|
||
<td id="76E5061BD4620587FC663435D7E2FC18" box="[936,1016,886,909]" gridcol="4" gridrow="1" pageId="16">Range</td>
|
||
<td id="76E5061BD4620587FBD43435D095FC18" box="[1050,1167,886,909]" gridcol="5" gridrow="1" pageId="16">Mean ± 1SD</td>
|
||
<td id="76E5061BD4620587FB773435D110FC18" box="[1209,1290,886,909]" gridcol="6" gridrow="1" pageId="16">Range</td>
|
||
<td id="76E5061BD4620587FAF13435D1AEFC18" box="[1343,1460,886,909]" gridcol="7" gridrow="1" pageId="16">Mean ± 1SD</td>
|
||
<td id="76E5061BD4620587FA193435D224FC18" box="[1495,1598,886,909]" gridcol="8" gridrow="1" pageId="16">Range</td>
|
||
<td id="76E5061BD4620587F9A93435D2C6FC18" box="[1639,1756,886,909]" gridcol="9" gridrow="1" pageId="16">Mean ± 1SD</td>
|
||
<td id="76E5061BD4620587F8C43435D341FC18" box="[1802,1883,886,909]" gridcol="10" gridrow="1" pageId="16">Range</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D34EBD341FC2A" box="[147,1883,936,959]" gridrow="2" pageId="16">
|
||
<th id="76E5061BD4620587FF5D34EBD5A3FC2A" box="[147,441,936,959]" gridcol="0" gridrow="2" pageId="16">Midbody dorsal scale rows</th>
|
||
<td id="76E5061BD4620587FE2E34EBD668FC2A" box="[480,626,936,959]" gridcol="1" gridrow="2" pageId="16">17.0 ± 1.0</td>
|
||
<td id="76E5061BD4620587FD4734EBD6C0FC2A" box="[649,730,936,959]" gridcol="2" gridrow="2" pageId="16">17–19</td>
|
||
<td id="76E5061BD4620587FCD634EBD797FC2A" box="[792,909,936,959]" gridcol="3" gridrow="2" pageId="16">19.0 ± 0.3</td>
|
||
<td id="76E5061BD4620587FC6634EBD7E2FC2A" box="[936,1016,936,959]" gridcol="4" gridrow="2" pageId="16">17–21</td>
|
||
<td id="76E5061BD4620587FBD434EBD095FC2A" box="[1050,1167,936,959]" gridcol="5" gridrow="2" pageId="16">19.0 ± 0</td>
|
||
<td id="76E5061BD4620587FB7734EBD110FC2A" box="[1209,1290,936,959]" gridcol="6" gridrow="2" pageId="16">19</td>
|
||
<td id="76E5061BD4620587FAF134EBD1AEFC2A" box="[1343,1460,936,959]" gridcol="7" gridrow="2" pageId="16">19.0 ± 0.2</td>
|
||
<td id="76E5061BD4620587FA1934EBD224FC2A" box="[1495,1598,936,959]" gridcol="8" gridrow="2" pageId="16">18–21</td>
|
||
<td id="76E5061BD4620587F9A934EBD2C6FC2A" box="[1639,1756,936,959]" gridcol="9" gridrow="2" pageId="16">19.0 ± 0.0</td>
|
||
<td id="76E5061BD4620587F8C434EBD341FC2A" box="[1802,1883,936,959]" gridcol="10" gridrow="2" pageId="16">19</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D3499D341FC64" box="[147,1883,986,1009]" gridrow="3" pageId="16">
|
||
<th id="76E5061BD4620587FF5D3499D5A3FC64" box="[147,441,986,1009]" gridcol="0" gridrow="3" pageId="16">Ventrals</th>
|
||
<td id="76E5061BD4620587FE2E3499D668FC64" box="[480,626,986,1009]" gridcol="1" gridrow="3" pageId="16">211.8 ± 4.2</td>
|
||
<td id="76E5061BD4620587FD473499D6C0FC64" box="[649,730,986,1009]" gridcol="2" gridrow="3" pageId="16">203–221</td>
|
||
<td id="76E5061BD4620587FCD63499D797FC64" box="[792,909,986,1009]" gridcol="3" gridrow="3" pageId="16">217.8 ± 4.4</td>
|
||
<td id="76E5061BD4620587FC663499D7E2FC64" box="[936,1016,986,1009]" gridcol="4" gridrow="3" pageId="16">206–232</td>
|
||
<td id="76E5061BD4620587FBD43499D095FC64" box="[1050,1167,986,1009]" gridcol="5" gridrow="3" pageId="16">216.9 ± 4.2</td>
|
||
<td id="76E5061BD4620587FB773499D110FC64" box="[1209,1290,986,1009]" gridcol="6" gridrow="3" pageId="16">211–226</td>
|
||
<td id="76E5061BD4620587FAF13499D1AEFC64" box="[1343,1460,986,1009]" gridcol="7" gridrow="3" pageId="16">210.0 ± 4.8</td>
|
||
<td id="76E5061BD4620587FA193499D224FC64" box="[1495,1598,986,1009]" gridcol="8" gridrow="3" pageId="16">197–228</td>
|
||
<td id="76E5061BD4620587F9A93499D2C6FC64" box="[1639,1756,986,1009]" gridcol="9" gridrow="3" pageId="16">209.4 ± 3.4</td>
|
||
<td id="76E5061BD4620587F8C43499D341FC64" box="[1802,1883,986,1009]" gridcol="10" gridrow="3" pageId="16">204–215</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D3348D341FBB7" box="[147,1883,1035,1058]" gridrow="4" pageId="16">
|
||
<th id="76E5061BD4620587FF5D3348D5A3FBB7" box="[147,441,1035,1058]" gridcol="0" gridrow="4" pageId="16">Subcaudals</th>
|
||
<td id="76E5061BD4620587FE2E3348D668FBB7" box="[480,626,1035,1058]" gridcol="1" gridrow="4" pageId="16">65.1 ± 2.3</td>
|
||
<td id="76E5061BD4620587FD473348D6C0FBB7" box="[649,730,1035,1058]" gridcol="2" gridrow="4" pageId="16">60–70</td>
|
||
<td id="76E5061BD4620587FCD63348D797FBB7" box="[792,909,1035,1058]" gridcol="3" gridrow="4" pageId="16">65.5 ± 2.7</td>
|
||
<td id="76E5061BD4620587FC663348D7E2FBB7" box="[936,1016,1035,1058]" gridcol="4" gridrow="4" pageId="16">57–74</td>
|
||
<td id="76E5061BD4620587FBD43348D095FBB7" box="[1050,1167,1035,1058]" gridcol="5" gridrow="4" pageId="16">70.0 ± 1.6</td>
|
||
<td id="76E5061BD4620587FB773348D110FBB7" box="[1209,1290,1035,1058]" gridcol="6" gridrow="4" pageId="16">67–73</td>
|
||
<td id="76E5061BD4620587FAF13348D1AEFBB7" box="[1343,1460,1035,1058]" gridcol="7" gridrow="4" pageId="16">62.8 ± 2.8</td>
|
||
<td id="76E5061BD4620587FA193348D224FBB7" box="[1495,1598,1035,1058]" gridcol="8" gridrow="4" pageId="16">57–72</td>
|
||
<td id="76E5061BD4620587F9A93348D2C6FBB7" box="[1639,1756,1035,1058]" gridcol="9" gridrow="4" pageId="16">66.2 ± 4.6</td>
|
||
<td id="76E5061BD4620587F8C43348D341FBB7" box="[1802,1883,1035,1058]" gridcol="10" gridrow="4" pageId="16">57–70</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D337FD341FBE4" box="[147,1883,1084,1137]" gridrow="5" pageId="16">
|
||
<th id="76E5061BD4620587FF5D337FD5A3FBE4" box="[147,441,1084,1137]" gridcol="0" gridrow="5" pageId="16">First dark ventral of first main band</th>
|
||
<td id="76E5061BD4620587FE2E337FD668FBE4" box="[480,626,1084,1137]" gridcol="1" gridrow="5" pageId="16">18.0 ± 2.3</td>
|
||
<td id="76E5061BD4620587FD47337FD6C0FBE4" box="[649,730,1084,1137]" gridcol="2" gridrow="5" pageId="16">10–23</td>
|
||
<td id="76E5061BD4620587FCD6337FD797FBE4" box="[792,909,1084,1137]" gridcol="3" gridrow="5" pageId="16">16.3 ± 1.7</td>
|
||
<td id="76E5061BD4620587FC66337FD7E2FBE4" box="[936,1016,1084,1137]" gridcol="4" gridrow="5" pageId="16">11–21</td>
|
||
<td id="76E5061BD4620587FBD4337FD095FBE4" box="[1050,1167,1084,1137]" gridcol="5" gridrow="5" pageId="16">16.4 ± 2.2</td>
|
||
<td id="76E5061BD4620587FB77337FD110FBE4" box="[1209,1290,1084,1137]" gridcol="6" gridrow="5" pageId="16">11–21</td>
|
||
<td id="76E5061BD4620587FAF1337FD1AEFBE4" box="[1343,1460,1084,1137]" gridcol="7" gridrow="5" pageId="16">14.2 ± 1.7</td>
|
||
<td id="76E5061BD4620587FA19337FD224FBE4" box="[1495,1598,1084,1137]" gridcol="8" gridrow="5" pageId="16">10–22</td>
|
||
<td id="76E5061BD4620587F9A9337FD2C6FBE4" box="[1639,1756,1084,1137]" gridcol="9" gridrow="5" pageId="16">18.2 ± 3.4</td>
|
||
<td id="76E5061BD4620587F8C4337FD341FBE4" box="[1802,1883,1084,1137]" gridcol="10" gridrow="5" pageId="16">14–22</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D333BD341FB39" box="[147,1883,1144,1196]" gridrow="6" pageId="16">
|
||
<th id="76E5061BD4620587FF5D333BD5A3FB39" box="[147,441,1144,1196]" gridcol="0" gridrow="6" pageId="16">Last dark ventral of first main band</th>
|
||
<td id="76E5061BD4620587FE2E333BD668FB39" box="[480,626,1144,1196]" gridcol="1" gridrow="6" pageId="16">22.2 ± 2.3</td>
|
||
<td id="76E5061BD4620587FD47333BD6C0FB39" box="[649,730,1144,1196]" gridcol="2" gridrow="6" pageId="16">16.26</td>
|
||
<td id="76E5061BD4620587FCD6333BD797FB39" box="[792,909,1144,1196]" gridcol="3" gridrow="6" pageId="16">19.9 ± 1.9</td>
|
||
<td id="76E5061BD4620587FC66333BD7E2FB39" box="[936,1016,1144,1196]" gridcol="4" gridrow="6" pageId="16">15–25</td>
|
||
<td id="76E5061BD4620587FBD4333BD095FB39" box="[1050,1167,1144,1196]" gridcol="5" gridrow="6" pageId="16">19.4 ± 2.4</td>
|
||
<td id="76E5061BD4620587FB77333BD110FB39" box="[1209,1290,1144,1196]" gridcol="6" gridrow="6" pageId="16">14–25</td>
|
||
<td id="76E5061BD4620587FAF1333BD1AEFB39" box="[1343,1460,1144,1196]" gridcol="7" gridrow="6" pageId="16">18.2 ± 2.1</td>
|
||
<td id="76E5061BD4620587FA19333BD224FB39" box="[1495,1598,1144,1196]" gridcol="8" gridrow="6" pageId="16">13–26</td>
|
||
<td id="76E5061BD4620587F9A9333BD2C6FB39" box="[1639,1756,1144,1196]" gridcol="9" gridrow="6" pageId="16">22.63 ± 6.4</td>
|
||
<td id="76E5061BD4620587F8C4333BD341FB39" box="[1802,1883,1144,1196]" gridcol="10" gridrow="6" pageId="16">15–29</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D33F0D341FB7D" box="[147,1883,1203,1256]" gridrow="7" pageId="16">
|
||
<th id="76E5061BD4620587FF5D33F0D5A3FB7D" box="[147,441,1203,1256]" gridcol="0" gridrow="7" pageId="16">Width of first main band (in ventral scales)</th>
|
||
<td id="76E5061BD4620587FE2E33F0D668FB7D" box="[480,626,1203,1256]" gridcol="1" gridrow="7" pageId="16">5.2 ± 1.4</td>
|
||
<td id="76E5061BD4620587FD4733F0D6C0FB7D" box="[649,730,1203,1256]" gridcol="2" gridrow="7" pageId="16">2–9</td>
|
||
<td id="76E5061BD4620587FCD633F0D797FB7D" box="[792,909,1203,1256]" gridcol="3" gridrow="7" pageId="16">4.5 ± 1.0</td>
|
||
<td id="76E5061BD4620587FC6633F0D7E2FB7D" box="[936,1016,1203,1256]" gridcol="4" gridrow="7" pageId="16">2–9</td>
|
||
<td id="76E5061BD4620587FBD433F0D095FB7D" box="[1050,1167,1203,1256]" gridcol="5" gridrow="7" pageId="16">4.0 ± 0.8</td>
|
||
<td id="76E5061BD4620587FB7733F0D110FB7D" box="[1209,1290,1203,1256]" gridcol="6" gridrow="7" pageId="16">3–5</td>
|
||
<td id="76E5061BD4620587FAF133F0D1AEFB7D" box="[1343,1460,1203,1256]" gridcol="7" gridrow="7" pageId="16">5.0 ± 1.0</td>
|
||
<td id="76E5061BD4620587FA1933F0D224FB7D" box="[1495,1598,1203,1256]" gridcol="8" gridrow="7" pageId="16">2–10</td>
|
||
<td id="76E5061BD4620587F9A933F0D2C6FB7D" box="[1639,1756,1203,1256]" gridcol="9" gridrow="7" pageId="16">5.4 ± 3.1</td>
|
||
<td id="76E5061BD4620587F8C433F0D341FB7D" box="[1802,1883,1203,1256]" gridcol="10" gridrow="7" pageId="16">2–9</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D33ACD341FA93" box="[147,1883,1263,1286]" gridrow="8" pageId="16">
|
||
<th id="76E5061BD4620587FF5D33ACD5A3FA93" box="[147,441,1263,1286]" gridcol="0" gridrow="8" pageId="16">Number of main bands</th>
|
||
<td id="76E5061BD4620587FE2E33ACD668FA93" box="[480,626,1263,1286]" gridcol="1" gridrow="8" pageId="16">1.5 ± 0.7</td>
|
||
<td id="76E5061BD4620587FD4733ACD6C0FA93" box="[649,730,1263,1286]" gridcol="2" gridrow="8" pageId="16">1–4</td>
|
||
<td id="76E5061BD4620587FCD633ACD797FA93" box="[792,909,1263,1286]" gridcol="3" gridrow="8" pageId="16">3.7 ± 0.9</td>
|
||
<td id="76E5061BD4620587FC6633ACD7E2FA93" box="[936,1016,1263,1286]" gridcol="4" gridrow="8" pageId="16">1–7</td>
|
||
<td id="76E5061BD4620587FBD433ACD095FA93" box="[1050,1167,1263,1286]" gridcol="5" gridrow="8" pageId="16">3.6 ± 1.3</td>
|
||
<td id="76E5061BD4620587FB7733ACD110FA93" box="[1209,1290,1263,1286]" gridcol="6" gridrow="8" pageId="16">1–7</td>
|
||
<td id="76E5061BD4620587FAF133ACD1AEFA93" box="[1343,1460,1263,1286]" gridcol="7" gridrow="8" pageId="16">2.5 ± 0.8</td>
|
||
<td id="76E5061BD4620587FA1933ACD224FA93" box="[1495,1598,1263,1286]" gridcol="8" gridrow="8" pageId="16">1–5</td>
|
||
<td id="76E5061BD4620587F9A933ACD2C6FA93" box="[1639,1756,1263,1286]" gridcol="9" gridrow="8" pageId="16">1.7 ± 0.7</td>
|
||
<td id="76E5061BD4620587F8C433ACD341FA93" box="[1802,1883,1263,1286]" gridcol="10" gridrow="8" pageId="16">1–3</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D3263D341FAA2" box="[147,1883,1312,1335]" gridrow="9" pageId="16">
|
||
<th id="76E5061BD4620587FF5D3263D5A3FAA2" box="[147,441,1312,1335]" gridcol="0" gridrow="9" pageId="16">Number of accessory bands</th>
|
||
<td id="76E5061BD4620587FE2E3263D668FAA2" box="[480,626,1312,1335]" gridcol="1" gridrow="9" pageId="16">0.8 ± 1.2</td>
|
||
<td id="76E5061BD4620587FD473263D6C0FAA2" box="[649,730,1312,1335]" gridcol="2" gridrow="9" pageId="16">0–5</td>
|
||
<td id="76E5061BD4620587FCD63263D797FAA2" box="[792,909,1312,1335]" gridcol="3" gridrow="9" pageId="16">1.4 ± 1.7</td>
|
||
<td id="76E5061BD4620587FC663263D7E2FAA2" box="[936,1016,1312,1335]" gridcol="4" gridrow="9" pageId="16">0–9</td>
|
||
<td id="76E5061BD4620587FBD43263D095FAA2" box="[1050,1167,1312,1335]" gridcol="5" gridrow="9" pageId="16">1.4 ± 1.9</td>
|
||
<td id="76E5061BD4620587FB773263D110FAA2" box="[1209,1290,1312,1335]" gridcol="6" gridrow="9" pageId="16">0–6</td>
|
||
<td id="76E5061BD4620587FAF13263D1AEFAA2" box="[1343,1460,1312,1335]" gridcol="7" gridrow="9" pageId="16">0.3 ± 0.79</td>
|
||
<td id="76E5061BD4620587FA193263D224FAA2" box="[1495,1598,1312,1335]" gridcol="8" gridrow="9" pageId="16">0–4</td>
|
||
<td id="76E5061BD4620587F9A93263D2C6FAA2" box="[1639,1756,1312,1335]" gridcol="9" gridrow="9" pageId="16">0.7 ± 0.9</td>
|
||
<td id="76E5061BD4620587F8C43263D341FAA2" box="[1802,1883,1312,1335]" gridcol="10" gridrow="9" pageId="16">0–2</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D3212D341FA13" box="[147,1883,1361,1414]" gridrow="10" pageId="16">
|
||
<th id="76E5061BD4620587FF5D3212D5A3FA13" box="[147,441,1361,1414]" gridcol="0" gridrow="10" pageId="16">Total number of discrete dark bands</th>
|
||
<td id="76E5061BD4620587FE2E3212D668FA13" box="[480,626,1361,1414]" gridcol="1" gridrow="10" pageId="16">2.3 ± 1.4</td>
|
||
<td id="76E5061BD4620587FD473212D6C0FA13" box="[649,730,1361,1414]" gridcol="2" gridrow="10" pageId="16">1–6</td>
|
||
<td id="76E5061BD4620587FCD63212D797FA13" box="[792,909,1361,1414]" gridcol="3" gridrow="10" pageId="16">5.1 ± 1.7</td>
|
||
<td id="76E5061BD4620587FC663212D7E2FA13" box="[936,1016,1361,1414]" gridcol="4" gridrow="10" pageId="16">2–12</td>
|
||
<td id="76E5061BD4620587FBD43212D095FA13" box="[1050,1167,1361,1414]" gridcol="5" gridrow="10" pageId="16">5.0 ± 2.5</td>
|
||
<td id="76E5061BD4620587FB773212D110FA13" box="[1209,1290,1361,1414]" gridcol="6" gridrow="10" pageId="16">2–12</td>
|
||
<td id="76E5061BD4620587FAF13212D1AEFA13" box="[1343,1460,1361,1414]" gridcol="7" gridrow="10" pageId="16">2.8 ± 1.1</td>
|
||
<td id="76E5061BD4620587FA193212D224FA13" box="[1495,1598,1361,1414]" gridcol="8" gridrow="10" pageId="16">1–8</td>
|
||
<td id="76E5061BD4620587F9A93212D2C6FA13" box="[1639,1756,1361,1414]" gridcol="9" gridrow="10" pageId="16">2.4 ± 1.2</td>
|
||
<td id="76E5061BD4620587F8C43212D341FA13" box="[1802,1883,1361,1414]" gridcol="10" gridrow="10" pageId="16">1–5</td>
|
||
</tr>
|
||
<tr id="35346F67D4620587FF5D32CED341FA57" box="[147,1883,1421,1474]" gridrow="11" pageId="16">
|
||
<th id="76E5061BD4620587FF5D32CED5A3FA57" box="[147,441,1421,1474]" gridcol="0" gridrow="11" pageId="16">Last ventral of last discrete dark band</th>
|
||
<td id="76E5061BD4620587FE2E32CED668FA57" box="[480,626,1421,1474]" gridcol="1" gridrow="11" pageId="16">28.5 ± 9.2</td>
|
||
<td id="76E5061BD4620587FD4732CED6C0FA57" box="[649,730,1421,1474]" gridcol="2" gridrow="11" pageId="16">17–52</td>
|
||
<td id="76E5061BD4620587FCD632CED797FA57" box="[792,909,1421,1474]" gridcol="3" gridrow="11" pageId="16">68.6 ± 15.4</td>
|
||
<td id="76E5061BD4620587FC6632CED7E2FA57" box="[936,1016,1421,1474]" gridcol="4" gridrow="11" pageId="16">24–114</td>
|
||
<td id="76E5061BD4620587FBD432CED095FA57" box="[1050,1167,1421,1474]" gridcol="5" gridrow="11" pageId="16">59.4 ± 19.1</td>
|
||
<td id="76E5061BD4620587FB7732CED110FA57" box="[1209,1290,1421,1474]" gridcol="6" gridrow="11" pageId="16">19–103</td>
|
||
<td id="76E5061BD4620587FAF132CED1AEFA57" box="[1343,1460,1421,1474]" gridcol="7" gridrow="11" pageId="16">36.8 ± 10.0</td>
|
||
<td id="76E5061BD4620587FA1932CED224FA57" box="[1495,1598,1421,1474]" gridcol="8" gridrow="11" pageId="16">13–82</td>
|
||
<td id="76E5061BD4620587F9A932CED2C6FA57" box="[1639,1756,1421,1474]" gridcol="9" gridrow="11" pageId="16">27.6 ± 15.4</td>
|
||
<td id="76E5061BD4620587F8C432CED341FA57" box="[1802,1883,1421,1474]" gridcol="10" gridrow="11" pageId="16">15–55</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D463FA69FF0937D4D513FE88" blockId="17.[151,1437,151,536]" pageId="17">
|
||
<emphasis id="B970B137D463FA69FF0937D4D57BFF25" bold="true" box="[199,353,151,176]" pageId="17">Colouration:</emphasis>
|
||
HEAd brOwN, thE suprAlAbiAls bArrEd blAck ANd yEllOw,
|
||
<collectingRegion id="49C0A3C7D463FA69FC1137D4D015FF25" box="[991,1039,151,176]" country="Myanmar" name="Chin" pageId="17">chiN</collectingRegion>
|
||
yEllOw. BlAck AbOvE, sOmEtimEs
|
||
<collectingRegion id="49C0A3C7D463FA69FA5E37DBD186FF25" box="[1424,1436,152,176]" country="Nigeria" name="Rivers" pageId="17">A</collectingRegion>
|
||
yEllOw mONOcEllAtE mArkiNg ON thE hOOd, Or 1–3 smAll yEllOw blOtchEs, thE dOrsAl scAlEs mAy bE tippEd with whitE iN juvENilEs (
|
||
<figureCitation id="133F71A0D463FA69FEE6379CD571FF6D" box="[296,363,223,248]" captionStart="FIGURE 8" captionStartId="15.[151,250,1955,1977]" captionTargetBox="[151,1436,1485,1930]" captionTargetId="figure@15.[151,1436,1485,1930]" captionTargetPageId="15" captionText="FIGURE 8. Naja (Boulengerina) melanoleuca. Adult specimens from Yaoundé, Cameroon (left—photo J.-F. Trape) and Tsibilé, Gabon (right—photo L. Chirio). Note the diffuse but distinct hood mark that is often present in this species, and the combination of broad main bands and narrow accessory bands on the ventral side." httpUri="https://zenodo.org/record/1456987/files/figure.png" pageId="17">Fig. 8</figureCitation>
|
||
). YEllOw Or whitE bElOw with 4
|
||
<collectingRegion id="49C0A3C7D463FA69FD1D37A3D6F3FF6D" box="[723,745,224,248]" country="South Sudan" name="Lakes" pageId="17">tO</collectingRegion>
|
||
6 blAck bANds ON thE first 100 vENtrAls, usuAlly uNifOrm blAck thErEAftEr.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D463FA69FF093664D0FBFED5" blockId="17.[151,1437,151,536]" box="[199,1249,295,320]" pageId="17">
|
||
<emphasis id="B970B137D463FA69FF093664D584FED5" bold="true" box="[199,414,295,320]" pageId="17">Largest recorded:</emphasis>
|
||
2250+420 =
|
||
<quantity id="4CFCC0C0D463FA69FDFA3664D6BBFEAA" box="[564,673,295,320]" metricMagnitude="0" metricUnit="m" metricValue="2.67" pageId="17" unit="mm" value="2670.0">2670 mm</quantity>
|
||
, frOm MONiyA, IbAdAN,
|
||
<collectingCountry id="F3132DB5D463FA69FC673664D7E7FED5" box="[937,1021,295,320]" name="Nigeria" pageId="17">NigEriA</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF9510D4D463FA69FBC23664D0CEFED5" box="[1036,1236,295,320]" pageId="17" refId="ref12547" refString="Butler, J. A. (1982) Capture au Nigeria d'un Naja noir et blanc de grande taille et notes sur les autres Najas de l'Ouest Africain. Notes Africaines, 176, 110 - 111." type="journal article">ButlEr, 1982: 110</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D463FA69FF09360FD75CFEF0" blockId="17.[151,1437,151,536]" box="[199,838,332,357]" pageId="17">
|
||
<materialsCitation id="3B6C6778D463FA69FF09360FD75CFEF0" ID-GBIF-Occurrence="1917075143" box="[199,838,332,357]" location="CENtrAl" pageId="17" specimenCount="1" stateProvince="CENtrAl">
|
||
<emphasis id="B970B137D463FA69FF09360FD5EEFEF0" bold="true" box="[199,500,332,357]" pageId="17">Suggested common name</emphasis>
|
||
:
|
||
<collectingRegion id="49C0A3C7D463FA69FDCD360FD64CFEF0" box="[515,598,332,357]" country="Ghana" name="Central" pageId="17">CENtrAl</collectingRegion>
|
||
AfricAN fOrEst cObrA.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED463FA69FF09362CD015FD82" pageId="17" type="distribution">
|
||
<paragraph id="8BBB6D25D463FA69FF09362CD015FD82" blockId="17.[151,1437,151,536]" pageId="17">
|
||
<emphasis id="B970B137D463FA69FF09362CD578FE1D" bold="true" box="[199,354,367,392]" pageId="17">Distribution:</emphasis>
|
||
CENtErEd ON thE
|
||
<collectingRegion id="49C0A3C7D463FA69FDED3633D674FE1D" box="[547,622,368,392]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="17">
|
||
<collectingCountry id="F3132DB5D463FA69FDED3633D674FE1D" box="[547,622,368,392]" name="Democratic Republic of the Congo" pageId="17">CONgO</collectingCountry>
|
||
</collectingRegion>
|
||
BAsiN,
|
||
<collectingRegion id="49C0A3C7D463FA69FD0D3633D6EDFE12" box="[707,759,368,391]" country="Cameroon" name="West" pageId="17">wEst</collectingRegion>
|
||
<collectingRegion id="49C0A3C7D463FA69FD303633D70EFE1D" box="[766,788,368,392]" country="South Sudan" name="Lakes" pageId="17">tO</collectingRegion>
|
||
sOuthwEstErN
|
||
<collectingCountry id="F3132DB5D463FA69FC79362CD011FE1D" box="[951,1035,367,392]" name="Nigeria" pageId="17">NigEriA</collectingCountry>
|
||
ANd pOssibly
|
||
<collectingRegion id="49C0A3C7D463FA69FB68362CD11DFE1D" box="[1190,1287,367,392]" country="Zambia" name="Southern" pageId="17">sOuthErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D463FA69FAC0362CD14EFE1D" box="[1294,1364,367,392]" name="Benin" pageId="17">BENiN</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C0A3C7D463FA69FAAE362CD186FE1D" box="[1376,1436,367,392]" country="Cameroon" name="South" pageId="17">sOuth</collectingRegion>
|
||
<collectingRegion id="49C0A3C7D463FA69FF5936D7D4B7FE39" box="[151,173,404,428]" country="South Sudan" name="Lakes" pageId="17">tO</collectingRegion>
|
||
<collectingRegion id="49C0A3C7D463FA69FF7D36D7D508FE38" box="[179,274,404,429]" country="Ghana" name="Northern" pageId="17">NOrthErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D463FA69FED636D7D574FE38" box="[280,366,404,429]" name="Angola" pageId="17">ANgOlA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C0A3C7D463FA69FEB636D7D586FE39" box="[376,412,404,428]" country="Cameroon" name="North" pageId="17">NOt</collectingRegion>
|
||
ExtENdiNg
|
||
<collectingRegion id="49C0A3C7D463FA69FDD936D7D659FE39" box="[535,579,404,428]" country="Cameroon" name="East" pageId="17">EAst</collectingRegion>
|
||
Of thE AlbErtiNE Rift VAllEy, whErE it is rEplAcEd by
|
||
<emphasis id="B970B137D463FA69FB4836D6D117FE39" box="[1158,1293,404,428]" italics="true" pageId="17">
|
||
<taxonomicName id="4C0416A6D463FA69FB4836D6D113FE39" authorityName="LAurENt" authorityYear="1955" box="[1158,1289,404,428]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" phylum="Chordata" rank="species" species="subfulva">N. subfulva</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
ThE
|
||
<collectingRegion id="49C0A3C7D463FA69FA8B36D6D186FE39" box="[1349,1436,405,428]" country="Ghana" name="Western" pageId="17">wEstErN</collectingRegion>
|
||
rANgE limits ArE pOOrly uNdErstOOd. POpulAtiONs frOm sOuthEAstErN
|
||
<collectingCountry id="F3132DB5D463FA69FC5A36F4D7F2FE45" box="[916,1000,439,464]" name="Nigeria" pageId="17">NigEriA</collectingCountry>
|
||
ArE clEArly AssigNAblE
|
||
<collectingRegion id="49C0A3C7D463FA69FB3E36FBD11CFE45" box="[1264,1286,440,464]" country="South Sudan" name="Lakes" pageId="17">tO</collectingRegion>
|
||
this fOrm.
|
||
<collectingRegion id="49C0A3C7D463FA69FA4636FBD186FE45" box="[1416,1436,440,464]" country="Nigeria" name="Rivers" pageId="17">A</collectingRegion>
|
||
fEw spEcimENs frOm LAmtA,
|
||
<collectingRegion id="49C0A3C7D463FA69FE19369FD62DFE60" box="[471,567,476,501]" country="Zambia" name="Southern" pageId="17">sOuthErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D463FA69FDF1369FD698FE60" box="[575,642,476,501]" name="Benin" pageId="17">BENiN</collectingCountry>
|
||
(IRD 12.B, IRD 54.B, IRD 60.B), ANd
|
||
<collectingCountry id="F3132DB5D463FA69FBF3369FD09CFE60" box="[1085,1158,476,501]" name="Ghana" pageId="17">GhANA</collectingCountry>
|
||
(MNHN 1983.0663–64;
|
||
<collectingRegion id="49C0A3C7D463FA69FF593543D4A9FD8D" box="[151,179,512,536]" country="Cameroon" name="North" pageId="17">NO</collectingRegion>
|
||
furthEr lOcAlity iNfOrmAtiON) AlsO AppEAr
|
||
<collectingRegion id="49C0A3C7D463FA69FD4C3543D682FD8D" box="[642,664,512,536]" country="South Sudan" name="Lakes" pageId="17">tO</collectingRegion>
|
||
bE AssigNAblE
|
||
<collectingRegion id="49C0A3C7D463FA69FCF23543D748FD8D" box="[828,850,512,536]" country="South Sudan" name="Lakes" pageId="17">tO</collectingRegion>
|
||
<taxonomicName id="4C0416A6D463FA69FC973542D013FD8D" authorityName="HAllOwEll" authorityYear="1857" box="[857,1033,512,536]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" phylum="Chordata" rank="species" species="melanoleuca">
|
||
<emphasis id="B970B137D463FA69FC973542D013FD8D" box="[857,1033,512,536]" italics="true" pageId="17">N. melanoleuca</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |