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<document ID-DOI="http://doi.org/10.5281/zenodo.3792224" ID-GBIF-Dataset="6bed073a-175c-42e6-bb99-645c99d55b4f" ID-GBIF-Taxon="163750549" ID-ISSN="13132970" ID-Zenodo-Dep="3792224" ID-ZooBank="C631A347-306E-4773-84A4-E4712329186B" checkinTime="1587984202003" checkinUser="plazi" docAuthor="Miller, Jeremy, Griswold, Charles &amp; Yin, Chang" docDate="2009" docId="03ACB915FFA47428FF4E41B4FBCAF9DD" docLanguage="en" docName="ZK_article_1993.pdf" docOrigin="ZooKeys 11 (11)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Crassignatha Wunderlich 1995" docType="treatment" docVersion="9" lastPageId="60" lastPageNumber="69" masterDocId="FF95C16DFF9F7414FFC2455DFFF1FFB8" masterDocTitle="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers" masterLastPageNumber="195" masterPageNumber="9" pageId="59" pageNumber="68" updateTime="1635491590422" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Jeremy</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Griswold, Charles</mods:namePart>
<mods:affiliation>California Academy of Sciences ,, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Chang</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University ,, China</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2009</mods:date>
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<mods:number>2009-05-31</mods:number>
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<mods:number>11</mods:number>
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<mods:number>11</mods:number>
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<mods:identifier type="DOI">10.3897/zookeys.11.160</mods:identifier>
<mods:identifier type="GBIF-Dataset">6bed073a-175c-42e6-bb99-645c99d55b4f</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792224" ID-GBIF-Taxon="163750549" ID-Zenodo-Dep="3792224" LSID="urn:lsid:plazi:treatment:03ACB915FFA47428FF4E41B4FBCAF9DD" httpUri="http://treatment.plazi.org/id/03ACB915FFA47428FF4E41B4FBCAF9DD" lastPageId="60" lastPageNumber="69" pageId="59" pageNumber="68">
<subSubSection box="[140,602,1257,1284]" pageId="59" pageNumber="68" type="nomenclature">
<paragraph blockId="59.[140,602,1257,1284]" box="[140,602,1257,1284]" pageId="59" pageNumber="68">
<heading box="[140,602,1257,1284]" fontSize="11" level="2" pageId="59" pageNumber="68" reason="3">
<emphasis bold="true" box="[140,602,1257,1284]" pageId="59" pageNumber="68">
Genus
<taxonomicName authority="Wunderlich, 1995" authorityName="Wunderlich" authorityYear="1995" box="[223,602,1257,1284]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[223,378,1257,1284]" italics="true" pageId="59" pageNumber="68">Crassignatha</emphasis>
<bibRefCitation author="Wunderlich J" box="[385,602,1257,1283]" pageId="59" pageNumber="68" pagination="543 - 551" refId="ref47317" refString="Wunderlich J (1995) Drei bisher unbekannte Arten und Gattungen der Familie Anapidae (s. l.) aus Sud-Afrika, Brasilien und Malaysia (Arachnida: Araneae). Beitrage zur Araneologie 4: 543 - 551 (date on volume 1994; actually published 1995)." type="journal article" year="1995">Wunderlich, 1995</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="68" type="reference_group">
<paragraph blockId="59.[140,1107,1328,1390]" pageId="59" pageNumber="68">
<treatmentCitationGroup pageId="59" pageNumber="68">
<treatmentCitation author="Wunderlich J" box="[140,564,1328,1355]" page="546" pageId="59" pageNumber="68" year="1995">
<taxonomicName authority="Wunderlich, 1995: 546" authorityName="Wunderlich" authorityPageNumber="546" authorityYear="1995" box="[140,564,1328,1355]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis box="[140,278,1328,1354]" italics="true" pageId="59" pageNumber="68">Crassignatha</emphasis>
<bibRefCitation author="Wunderlich J" box="[288,564,1328,1355]" pageId="59" pageNumber="68" pagination="543 - 551" refId="ref47317" refString="Wunderlich J (1995) Drei bisher unbekannte Arten und Gattungen der Familie Anapidae (s. l.) aus Sud-Afrika, Brasilien und Malaysia (Arachnida: Araneae). Beitrage zur Araneologie 4: 543 - 551 (date on volume 1994; actually published 1995)." type="journal article" year="1995">Wunderlich, 1995: 546</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[577,632,1329,1355]" pageId="59" pageNumber="68">Type</typeStatus>
species
<taxonomicName authority="Wunderlich, 1995" authorityName="Wunderlich" authorityYear="1995" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="species" species="haeneli">
<emphasis box="[730,955,1328,1354]" italics="true" pageId="59" pageNumber="68">Crassignatha haeneli</emphasis>
<bibRefCitation author="Wunderlich J" pageId="59" pageNumber="68" pagination="543 - 551" refId="ref47317" refString="Wunderlich J (1995) Drei bisher unbekannte Arten und Gattungen der Familie Anapidae (s. l.) aus Sud-Afrika, Brasilien und Malaysia (Arachnida: Araneae). Beitrage zur Araneologie 4: 543 - 551 (date on volume 1994; actually published 1995)." type="journal article" year="1995">Wunderlich, 1995</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="60" lastPageNumber="69" pageId="59" pageNumber="68" type="discussion">
<paragraph blockId="59.[140,1108,1433,1636]" lastBlockId="60.[140,1108,167,1637]" lastPageId="60" lastPageNumber="69" pageId="59" pageNumber="68">
<emphasis bold="true" box="[140,362,1433,1460]" pageId="59" pageNumber="68">
<emphasis box="[140,222,1433,1459]" inLineHeading="true" pageId="59" pageNumber="68" reason="1">Family</emphasis>
placement.
</emphasis>
Conflicting opinions have led to instability in the family placement of
<taxonomicName authorityName="Wunderlich" authorityYear="1995" box="[241,379,1469,1495]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis box="[241,379,1469,1495]" italics="true" pageId="59" pageNumber="68">Crassignatha</emphasis>
</taxonomicName>
.
<bibRefCitation author="Wunderlich J" box="[396,616,1469,1496]" pageId="59" pageNumber="68" pagination="543 - 551" refId="ref47317" refString="Wunderlich J (1995) Drei bisher unbekannte Arten und Gattungen der Familie Anapidae (s. l.) aus Sud-Afrika, Brasilien und Malaysia (Arachnida: Araneae). Beitrage zur Araneologie 4: 543 - 551 (date on volume 1994; actually published 1995)." type="journal article" year="1995">Wunderlich (1995)</bibRefCitation>
described
<taxonomicName authorityName="Wunderlich" authorityYear="1995" box="[743,881,1469,1495]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis box="[743,881,1469,1495]" italics="true" pageId="59" pageNumber="68">Crassignatha</emphasis>
</taxonomicName>
as part of the family
<taxonomicName baseAuthorityName="Marusik and Lehtinen" baseAuthorityYear="2003" box="[176,316,1504,1530]" class="Arachnida" family="Synaphridae" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="family">Synaphridae</taxonomicName>
(which he considered a subfamily of a broadly circumscribed
<taxonomicName authorityName="Simon" authorityYear="1895" class="Arachnida" family="Anapidae" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="family">Anapidae</taxonomicName>
).
<bibRefCitation author="Marusik YM &amp; Lehtinen PT" box="[202,539,1540,1566]" pageId="59" pageNumber="68" pagination="143 - 152" refId="ref46406" refString="Marusik YM, Lehtinen PT (2003) Synaphridae Wunderlich, 1986 (Aranei: Araneoidea), a new family status, with a description of a new species from Turkmenistan. Arthropoda Selecta 11: 143 - 152 (date on volume 2002; actually published 2003)." type="journal article" year="2003">Marusik and Lehtinen (2003)</bibRefCitation>
suggested it might belong to
<taxonomicName authorityName="Hickman" authorityYear="1931" box="[874,1104,1540,1566]" class="Arachnida" family="Symphytognathidae" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="family">Symphytognathidae</taxonomicName>
;
<bibRefCitation author="Wunderlich J" box="[140,430,1575,1601]" pageId="59" pageNumber="68" pagination="1020 - 1111" refId="ref47368" refString="Wunderlich J (2004) The fossil spiders of the family Anapidae s. l. (Aeaneae) [sic] in Baltic, Dominican and Mexican amber and their extant relatives, with the description of the new subfamily Comarominae. Beitrage zur Araneologie 3: 1020 - 1111." type="journal article" year="2004">Wunderlich (2004: 1081)</bibRefCitation>
suggested
<taxonomicName authorityName="Simon" authorityYear="1895" box="[550,656,1575,1601]" class="Arachnida" family="Anapidae" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="family">Anapidae</taxonomicName>
(
<emphasis box="[672,797,1576,1601]" italics="true" pageId="59" pageNumber="68">sensu stricto</emphasis>
);
<bibRefCitation author="Platnick N" box="[821,997,1575,1601]" pageId="59" pageNumber="68" refId="ref46753" refString="Platnick N (2008) The World Spider Catalog, Version 9.0. American Museum of Natural History. http: // research. amnh. org / entomology / spiders / catalog / INTRO 1. html" type="url" year="2008">Platnick (2008)</bibRefCitation>
has cataloged it under
<taxonomicName authorityName="Petrunkevitch" authorityYear="1928" box="[295,443,1610,1636]" class="Arachnida" family="Mysmenidae" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="68" phylum="Arthropoda" rank="family">Mysmenidae</taxonomicName>
. The systematics and circumscription of symphytognathoid spider families, especially
<taxonomicName authorityName="Simon" authorityYear="1895" box="[440,546,167,193]" class="Arachnida" family="Anapidae" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="family">Anapidae</taxonomicName>
and
<taxonomicName authorityName="Hickman" authorityYear="1931" box="[609,839,167,193]" class="Arachnida" family="Symphytognathidae" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="family">Symphytognathidae</taxonomicName>
, is in need of revision. Nevertheless, we predict that an affinity between
<taxonomicName authorityName="Wunderlich" authorityYear="1995" box="[704,842,202,228]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[704,842,202,228]" italics="true" pageId="60" pageNumber="69">Crassignatha</emphasis>
</taxonomicName>
and at least some
<taxonomicName authorityName="Marples" authorityYear="1951" box="[1057,1108,203,228]" class="Arachnida" family="Symphytognathidae" genus="Patu" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[1057,1108,203,228]" italics="true" pageId="60" pageNumber="69">Patu</emphasis>
</taxonomicName>
among the
<taxonomicName authorityName="Hickman" authorityYear="1931" box="[271,497,237,263]" class="Arachnida" family="Symphytognathidae" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="family">Symphytognathidae</taxonomicName>
will be born out, supported in part by similarities in cheliceral armature and the male tibia II clasping spur.
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="69" type="diagnosis">
<paragraph blockId="60.[140,1108,167,1637]" pageId="60" pageNumber="69">
<emphasis bold="true" box="[185,317,307,333]" pageId="60" pageNumber="69">
<emphasis box="[185,311,307,333]" inLineHeading="true" pageId="60" pageNumber="69" reason="1">Diagnosis</emphasis>
.
</emphasis>
Distinguished from other symphytognathid genera except
<taxonomicName authorityName="Marples" authorityYear="1951" box="[1005,1058,308,333]" class="Arachnida" family="Symphytognathidae" genus="Patu" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[1005,1058,308,333]" italics="true" pageId="60" pageNumber="69">Patu</emphasis>
</taxonomicName>
and
<taxonomicName authority="Forster &amp; Platnick, 1977" authorityName="Forster &amp; Platnick" authorityYear="1977" box="[140,576,343,369]" class="Arachnida" family="Symphytognathidae" genus="Curimagua" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[140,269,343,369]" italics="true" pageId="60" pageNumber="69">Curimagua</emphasis>
<bibRefCitation author="Forster RR &amp; Platnick N" box="[278,576,343,369]" pageId="60" pageNumber="69" pagination="1 - 29" refId="ref45702" refString="Forster RR, Platnick N (1977) A review of the spider family Symphytognathidae (Arachnida, Araneae). American Museum Novitates 2619: 1 - 29." type="journal article" year="1977">Forster &amp; Platnick, 1977</bibRefCitation>
</taxonomicName>
by having the chelicerae fused only near the base (
<figureCitation box="[208,317,378,404]" captionStart="Figure 78" captionStartId="150.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@150.[140,1108,164,1186]" captionTargetPageId="150" captionText="Figure 78. SEM. A-D, Crassignatha pianma sp. n. from Chang Yan He; E, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; F, Crassignatha quanqu sp. n. from Bang Bie village. A, female chelicera removed from prosoma, note single asymetrically bifid tooth; B, labrum, anterior, female; C, labrum, lateral, female; D, male metatarsus-tarsus joint, arrow indicates tarsal organ; E, epiandrous region; F, prosoma, lateral, male. Arrows in B and C indicate maximum extension of labrum." figureDoi="http://doi.org/10.5281/zenodo.3768896" httpUri="https://zenodo.org/record/3768896/files/figure.png" pageId="60" pageNumber="69">Fig. 78A</figureCitation>
); distinguished from
<taxonomicName authorityName="Marples" authorityYear="1951" box="[574,627,379,404]" class="Arachnida" family="Symphytognathidae" genus="Patu" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[574,627,379,404]" italics="true" pageId="60" pageNumber="69">Patu</emphasis>
</taxonomicName>
by the sculpturing of the carapace (
<figureCitation captionStart="Figure 77" captionStartId="149.[140,212,1205,1227]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@149.[140,1109,164,1186]" captionTargetPageId="149" captionText="Figure 77. Crassignatha pianma sp. n. from Chang Yan He, SEM. A-E, male; F, female. A, palp, prolateral; B, palp, retrolateral; C, palp, dorsal; D, distal ventral part of tibia II showing presumed mating claspers; E, F, prosoma, lateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768894" httpUri="https://zenodo.org/record/3768894/files/figure.png" pageId="60" pageNumber="69">Fig. 77</figureCitation>
E-F;
<bibRefCitation author="Wunderlich J" box="[230,443,413,440]" pageId="60" pageNumber="69" pagination="543 - 551" refId="ref47317" refString="Wunderlich J (1995) Drei bisher unbekannte Arten und Gattungen der Familie Anapidae (s. l.) aus Sud-Afrika, Brasilien und Malaysia (Arachnida: Araneae). Beitrage zur Araneologie 4: 543 - 551 (date on volume 1994; actually published 1995)." type="journal article" year="1995">Wunderlich 1995</bibRefCitation>
: fig. 15); usually further distinguished from
<taxonomicName authorityName="Marples" authorityYear="1951" box="[972,1025,414,439]" class="Arachnida" family="Symphytognathidae" genus="Patu" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[972,1025,414,439]" italics="true" pageId="60" pageNumber="69">Patu</emphasis>
</taxonomicName>
by the abdominal scutum in the male wrapping around the posterior (
<figureCitation box="[899,1008,448,475]" captionStart="Figure 74" captionStartId="146.[140,212,1125,1147]" captionTargetBox="[163,1101,188,1093]" captionTargetId="figure@146.[140,1109,164,1105]" captionTargetPageId="146" captionText="Figure 74. Crassignatha pianma sp. n. from Chang Yan He. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." figureDoi="http://doi.org/10.5281/zenodo.3768888" httpUri="https://zenodo.org/record/3768888/files/figure.png" pageId="60" pageNumber="69">Fig. 74A</figureCitation>
; scutum absent in
<taxonomicName authority=", Wunderlich 2004" authorityName=", Wunderlich" authorityYear="2004" box="[258,605,484,510]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="haeneli">
<emphasis box="[258,375,484,510]" italics="true" pageId="60" pageNumber="69">C. haeneli</emphasis>
,
<bibRefCitation author="Wunderlich J" box="[392,605,484,510]" pageId="60" pageNumber="69" pagination="1020 - 1111" refId="ref47368" refString="Wunderlich J (2004) The fossil spiders of the family Anapidae s. l. (Aeaneae) [sic] in Baltic, Dominican and Mexican amber and their extant relatives, with the description of the new subfamily Comarominae. Beitrage zur Araneologie 3: 1020 - 1111." type="journal article" year="2004">Wunderlich 2004</bibRefCitation>
</taxonomicName>
); from
<taxonomicName authorityName="Forster &amp; Platnick" authorityYear="1977" box="[695,824,484,510]" class="Arachnida" family="Symphytognathidae" genus="Curimagua" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[695,824,484,510]" italics="true" pageId="60" pageNumber="69">Curimagua</emphasis>
</taxonomicName>
by having eyes in three diads (two triads in
<taxonomicName authorityName="Forster &amp; Platnick" authorityYear="1977" box="[393,522,519,545]" class="Arachnida" family="Symphytognathidae" genus="Curimagua" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[393,522,519,545]" italics="true" pageId="60" pageNumber="69">Curimagua</emphasis>
</taxonomicName>
;
<bibRefCitation author="Forster RR &amp; Platnick N" box="[541,862,519,545]" pageId="60" pageNumber="69" pagination="1 - 29" refId="ref45702" refString="Forster RR, Platnick N (1977) A review of the spider family Symphytognathidae (Arachnida, Araneae). American Museum Novitates 2619: 1 - 29." type="journal article" year="1977">Forster and Platnick 1977</bibRefCitation>
) and one (entire or bifid) or two cheliceral teeth (
<figureCitation box="[497,607,554,580]" captionStart="Figure 78" captionStartId="150.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@150.[140,1108,164,1186]" captionTargetPageId="150" captionText="Figure 78. SEM. A-D, Crassignatha pianma sp. n. from Chang Yan He; E, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; F, Crassignatha quanqu sp. n. from Bang Bie village. A, female chelicera removed from prosoma, note single asymetrically bifid tooth; B, labrum, anterior, female; C, labrum, lateral, female; D, male metatarsus-tarsus joint, arrow indicates tarsal organ; E, epiandrous region; F, prosoma, lateral, male. Arrows in B and C indicate maximum extension of labrum." figureDoi="http://doi.org/10.5281/zenodo.3768896" httpUri="https://zenodo.org/record/3768896/files/figure.png" pageId="60" pageNumber="69">Figs 78A</figureCitation>
, 91E;
<taxonomicName authorityName="Forster &amp; Platnick" authorityYear="1977" box="[686,815,554,580]" class="Arachnida" family="Symphytognathidae" genus="Curimagua" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[686,815,554,580]" italics="true" pageId="60" pageNumber="69">Curimagua</emphasis>
</taxonomicName>
is toothless;
<bibRefCitation author="Forster RR &amp; Platnick N" pageId="60" pageNumber="69" pagination="1 - 29" refId="ref45702" refString="Forster RR, Platnick N (1977) A review of the spider family Symphytognathidae (Arachnida, Araneae). American Museum Novitates 2619: 1 - 29." type="journal article" year="1977">Forster and Platnick 1977</bibRefCitation>
). The poorly known genus
<taxonomicName authority="Simon, 1905" authorityName="Simon" authorityYear="1905" box="[625,919,589,616]" class="Arachnida" family="Symphytognathidae" genus="Anapogonia" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[625,759,590,616]" italics="true" pageId="60" pageNumber="69">Anapogonia</emphasis>
Simon, 1905
</taxonomicName>
was not considered for this diagnosis.
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="69" type="description">
<paragraph blockId="60.[140,1108,167,1637]" pageId="60" pageNumber="69">
<emphasis bold="true" box="[185,340,659,685]" pageId="60" pageNumber="69">
<emphasis box="[185,335,659,685]" inLineHeading="true" pageId="60" pageNumber="69" reason="1">Description</emphasis>
.
</emphasis>
Tiny ecribellate araneoid spiders. Total length 0.6-1.3. Six eyes in three doublets (
<figureCitation box="[354,454,695,721]" captionStart="Figure 82" captionStartId="154.[140,212,1203,1225]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@154.[140,1108,164,1184]" captionTargetPageId="154" captionText="Figure 82. Crassignatha yinzhi from Xiao Hei Shan Nature Reserve, SEM of male. A, palp, prolateral; B, palp, retrolateral; C, palp, prolateral, detail; D, palp, dorsal; E, prosoma, lateral; F, prosoma, anterior, dorsal to the left. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768900" httpUri="https://zenodo.org/record/3768900/files/figure.png" pageId="60" pageNumber="69">Fig. 82F</figureCitation>
, 91C). Carapace with fields of tubercles and pores (
<figureCitation captionStart="Figure 77" captionStartId="149.[140,212,1205,1227]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@149.[140,1109,164,1186]" captionTargetPageId="149" captionText="Figure 77. Crassignatha pianma sp. n. from Chang Yan He, SEM. A-E, male; F, female. A, palp, prolateral; B, palp, retrolateral; C, palp, dorsal; D, distal ventral part of tibia II showing presumed mating claspers; E, F, prosoma, lateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768894" httpUri="https://zenodo.org/record/3768894/files/figure.png" pageId="60" pageNumber="69">Figs 77E, F</figureCitation>
, 91A-D). Head region and clypeus raised in male, clypeus with two vertical clusters of sulci (
<figureCitation box="[411,520,765,791]" captionStart="Figure 82" captionStartId="154.[140,212,1203,1225]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@154.[140,1108,164,1184]" captionTargetPageId="154" captionText="Figure 82. Crassignatha yinzhi from Xiao Hei Shan Nature Reserve, SEM of male. A, palp, prolateral; B, palp, retrolateral; C, palp, prolateral, detail; D, palp, dorsal; E, prosoma, lateral; F, prosoma, anterior, dorsal to the left. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768900" httpUri="https://zenodo.org/record/3768900/files/figure.png" pageId="60" pageNumber="69">Fig. 82F</figureCitation>
). Sternum truncate posteriorly. Chelicerae fused near base, fang furrow absent, usually with a single anterior tooth bearing a small mesal apex (
<figureCitation box="[290,399,836,862]" captionStart="Figure 78" captionStartId="150.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@150.[140,1108,164,1186]" captionTargetPageId="150" captionText="Figure 78. SEM. A-D, Crassignatha pianma sp. n. from Chang Yan He; E, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; F, Crassignatha quanqu sp. n. from Bang Bie village. A, female chelicera removed from prosoma, note single asymetrically bifid tooth; B, labrum, anterior, female; C, labrum, lateral, female; D, male metatarsus-tarsus joint, arrow indicates tarsal organ; E, epiandrous region; F, prosoma, lateral, male. Arrows in B and C indicate maximum extension of labrum." figureDoi="http://doi.org/10.5281/zenodo.3768896" httpUri="https://zenodo.org/record/3768896/files/figure.png" pageId="60" pageNumber="69">Fig. 78A</figureCitation>
) except in
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[539,661,836,862]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="longtou">
<emphasis box="[539,661,836,862]" italics="true" pageId="60" pageNumber="69">C. longtou</emphasis>
</taxonomicName>
, which has two subequal teeth (Fig. 91E). Labrum weakly sclerotized, without spur (
<figureCitation box="[714,800,871,897]" captionStart="Figure 78" captionStartId="150.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@150.[140,1108,164,1186]" captionTargetPageId="150" captionText="Figure 78. SEM. A-D, Crassignatha pianma sp. n. from Chang Yan He; E, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; F, Crassignatha quanqu sp. n. from Bang Bie village. A, female chelicera removed from prosoma, note single asymetrically bifid tooth; B, labrum, anterior, female; C, labrum, lateral, female; D, male metatarsus-tarsus joint, arrow indicates tarsal organ; E, epiandrous region; F, prosoma, lateral, male. Arrows in B and C indicate maximum extension of labrum." figureDoi="http://doi.org/10.5281/zenodo.3768896" httpUri="https://zenodo.org/record/3768896/files/figure.png" pageId="60" pageNumber="69">Fig. 78</figureCitation>
B-C). Female palp absent. Metatarsus-tarsus joint without synaphrid-like distal constriction (see
<bibRefCitation author="Lopardo L &amp; Hormiga G &amp; Melic A" pageId="60" pageNumber="69" pagination="1 - 26" refId="ref46353" refString="Lopardo L, Hormiga G, Melic A (2007) Spinneret spigot morphology in synaphrid spiders (Araneae, Synaphridae), with comments on the systematics of the family and description of a new speices of Synaphris Simon 1894 from Spain. American Museum Novitates 3556: 1 - 26." type="journal article" year="2007">Lopardo et al. 2007</bibRefCitation>
). Tarsal organ near proximal margin, round, on slightly raised base (
<figureCitation captionStart="Figure 78" captionStartId="150.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@150.[140,1108,164,1186]" captionTargetPageId="150" captionText="Figure 78. SEM. A-D, Crassignatha pianma sp. n. from Chang Yan He; E, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; F, Crassignatha quanqu sp. n. from Bang Bie village. A, female chelicera removed from prosoma, note single asymetrically bifid tooth; B, labrum, anterior, female; C, labrum, lateral, female; D, male metatarsus-tarsus joint, arrow indicates tarsal organ; E, epiandrous region; F, prosoma, lateral, male. Arrows in B and C indicate maximum extension of labrum." figureDoi="http://doi.org/10.5281/zenodo.3768896" httpUri="https://zenodo.org/record/3768896/files/figure.png" pageId="60" pageNumber="69">Fig. 78D</figureCitation>
). Tibiae with two dorsal rows of trichobothria, metatarsi I and II (plus III in
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="longtou">
<emphasis italics="true" pageId="60" pageNumber="69">C. longtou</emphasis>
</taxonomicName>
) with trichobothrium. Male tibia II with field of 2-4 thick setae on ventral distal part. Abdominal setae long and sparse. Male abdomen usually with scutum laterally and posteriorly; female without scutum, although a sclerotized ring around spinnerets may be present (
<figureCitation box="[471,580,1117,1143]" captionStart="Figure 80" captionStartId="152.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@152.[140,1109,164,1270]" captionTargetPageId="152" captionText="Figure 80. Crassignatha yinzhi from Xiao Hei Shan Nature Reserve. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." figureDoi="http://doi.org/10.5281/zenodo.3813021" httpUri="https://zenodo.org/record/3813021/files/figure.png" pageId="60" pageNumber="69">Fig. 80B</figureCitation>
). Spinnerets with ventral orientation. Adult male retains flagelliform aggregate triplet (
<figureCitation box="[654,762,1152,1179]" captionStart="Figure 85" captionStartId="157.[140,212,1202,1224]" captionTargetBox="[140,1108,165,1182]" captionTargetId="figure@157.[140,1108,164,1182]" captionTargetPageId="157" captionText="Figure 85. Crassignatha quanqu sp. n. from Bang Bie village, SEM of spinnerets. A-E, female; F, male. A, spinnerets; B, ALS; C, PMS; D, F, PLS; E, base of ALS showing lack of colulus. AC: aciniform gland spigot; AG: aggregate gland spigot; ALS: anterior lateral spinneret; CY: cylindrical gland spigot; FL:flagelliform gland spigot; MAP: major ampullate gland spigot; mAP: minor ampullate gland spigot; n: nubbin; PI: piriform gland spigot; PLS: posterior lateral spinneret; PMS: posterior median spinneret; t: tartipore." figureDoi="http://doi.org/10.5281/zenodo.3768906" httpUri="https://zenodo.org/record/3768906/files/figure.png" pageId="60" pageNumber="69">Fig. 85F</figureCitation>
). Colulus absent (
<figureCitation box="[985,1095,1152,1179]" captionStart="Figure 85" captionStartId="157.[140,212,1202,1224]" captionTargetBox="[140,1108,165,1182]" captionTargetId="figure@157.[140,1108,164,1182]" captionTargetPageId="157" captionText="Figure 85. Crassignatha quanqu sp. n. from Bang Bie village, SEM of spinnerets. A-E, female; F, male. A, spinnerets; B, ALS; C, PMS; D, F, PLS; E, base of ALS showing lack of colulus. AC: aciniform gland spigot; AG: aggregate gland spigot; ALS: anterior lateral spinneret; CY: cylindrical gland spigot; FL:flagelliform gland spigot; MAP: major ampullate gland spigot; mAP: minor ampullate gland spigot; n: nubbin; PI: piriform gland spigot; PLS: posterior lateral spinneret; PMS: posterior median spinneret; t: tartipore." figureDoi="http://doi.org/10.5281/zenodo.3768906" httpUri="https://zenodo.org/record/3768906/files/figure.png" pageId="60" pageNumber="69">Fig. 85E</figureCitation>
). Epiandrous gland spigots absent (
<figureCitation box="[533,639,1188,1214]" captionStart="Figure 78" captionStartId="150.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@150.[140,1108,164,1186]" captionTargetPageId="150" captionText="Figure 78. SEM. A-D, Crassignatha pianma sp. n. from Chang Yan He; E, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; F, Crassignatha quanqu sp. n. from Bang Bie village. A, female chelicera removed from prosoma, note single asymetrically bifid tooth; B, labrum, anterior, female; C, labrum, lateral, female; D, male metatarsus-tarsus joint, arrow indicates tarsal organ; E, epiandrous region; F, prosoma, lateral, male. Arrows in B and C indicate maximum extension of labrum." figureDoi="http://doi.org/10.5281/zenodo.3768896" httpUri="https://zenodo.org/record/3768896/files/figure.png" pageId="60" pageNumber="69">Fig. 78E</figureCitation>
).
</paragraph>
<paragraph blockId="60.[140,1108,167,1637]" pageId="60" pageNumber="69">
Male palp: Palpal tibia without trichobothria (
<figureCitation box="[723,830,1223,1249]" captionStart="Figure 77" captionStartId="149.[140,212,1205,1227]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@149.[140,1109,164,1186]" captionTargetPageId="149" captionText="Figure 77. Crassignatha pianma sp. n. from Chang Yan He, SEM. A-E, male; F, female. A, palp, prolateral; B, palp, retrolateral; C, palp, dorsal; D, distal ventral part of tibia II showing presumed mating claspers; E, F, prosoma, lateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768894" httpUri="https://zenodo.org/record/3768894/files/figure.png" pageId="60" pageNumber="69">Fig. 77C</figureCitation>
). Cymbium with dorsal tooth near distal margin (
<figureCitation box="[442,548,1258,1285]" captionStart="Figure 77" captionStartId="149.[140,212,1205,1227]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@149.[140,1109,164,1186]" captionTargetPageId="149" captionText="Figure 77. Crassignatha pianma sp. n. from Chang Yan He, SEM. A-E, male; F, female. A, palp, prolateral; B, palp, retrolateral; C, palp, dorsal; D, distal ventral part of tibia II showing presumed mating claspers; E, F, prosoma, lateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768894" httpUri="https://zenodo.org/record/3768894/files/figure.png" pageId="60" pageNumber="69">Fig. 77B</figureCitation>
). Tegulum large and bulbous. Plate-like median apophysis on prolateral part of bulb. Membranous apophysis arises from near anterior part of median apophysis (
<figureCitation box="[440,544,1328,1355]" captionStart="Figure 82" captionStartId="154.[140,212,1203,1225]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@154.[140,1108,164,1184]" captionTargetPageId="154" captionText="Figure 82. Crassignatha yinzhi from Xiao Hei Shan Nature Reserve, SEM of male. A, palp, prolateral; B, palp, retrolateral; C, palp, prolateral, detail; D, palp, dorsal; E, prosoma, lateral; F, prosoma, anterior, dorsal to the left. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768900" httpUri="https://zenodo.org/record/3768900/files/figure.png" pageId="60" pageNumber="69">Fig. 82C</figureCitation>
). Embolus usually thick, rigid, more or less spiral, rarely long and flexible.
</paragraph>
<paragraph blockId="60.[140,1108,167,1637]" pageId="60" pageNumber="69">
<taxonomicName authority=": Epigynum" authorityName="Epigynum" box="[185,383,1399,1425]" class="Polychaeta" family="Polychaeta (awaiting allocation)" genus="Vulva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polychaeta (awaiting allocation)" pageId="60" pageNumber="69" phylum="Annelida" rank="genus">Vulva: Epigynum</taxonomicName>
present, usually a short rounded scape (
<figureCitation box="[844,928,1399,1425]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="60" pageNumber="69">Fig. 79</figureCitation>
), rarely a transverse bulge (Fig. 91F). Two round spermathecae separated by about their diameter. Copulatory ducts follow a path to near apex of scape (
<figureCitation box="[778,883,1469,1496]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="60" pageNumber="69">Fig. 76B</figureCitation>
); fertilization ducts inconspicuous.
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="69" type="reference_group">
<paragraph blockId="60.[140,1108,167,1637]" pageId="60" pageNumber="69">
<emphasis bold="true" box="[185,279,1539,1565]" pageId="60" pageNumber="69">
<emphasis box="[185,274,1539,1565]" inLineHeading="true" pageId="60" pageNumber="69" reason="1">Species</emphasis>
.
</emphasis>
<bibRefCitation author="Wunderlich J" box="[287,507,1539,1566]" pageId="60" pageNumber="69" pagination="543 - 551" refId="ref47317" refString="Wunderlich J (1995) Drei bisher unbekannte Arten und Gattungen der Familie Anapidae (s. l.) aus Sud-Afrika, Brasilien und Malaysia (Arachnida: Araneae). Beitrage zur Araneologie 4: 543 - 551 (date on volume 1994; actually published 1995)." type="journal article" year="1995">Wunderlich (1995)</bibRefCitation>
established
<taxonomicName authorityName="Wunderlich" authorityYear="1995" box="[649,787,1540,1566]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis box="[649,787,1540,1566]" italics="true" pageId="60" pageNumber="69">Crassignatha</emphasis>
</taxonomicName>
to accommodate
<taxonomicName authorityName=", Wunderlich" authorityYear="2004" box="[995,1107,1540,1566]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="haeneli">
<emphasis box="[995,1107,1540,1566]" italics="true" pageId="60" pageNumber="69">C. haeneli</emphasis>
</taxonomicName>
based on a single male specimen from
<collectingCountry box="[588,691,1575,1601]" name="Malaysia" pageId="60" pageNumber="69">Malaysia</collectingCountry>
. Seven new species are added to the genus:
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[219,336,1610,1636]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="pianma">
<emphasis box="[219,336,1610,1636]" italics="true" pageId="60" pageNumber="69">C. pianma</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[351,452,1610,1636]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="yinzhi">
<emphasis box="[351,452,1610,1636]" italics="true" pageId="60" pageNumber="69">C. yinzhi</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[467,584,1610,1636]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="quanqu">
<emphasis box="[467,584,1610,1636]" italics="true" pageId="60" pageNumber="69">C. quanqu</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[599,693,1610,1636]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="yamu">
<emphasis box="[599,693,1610,1636]" italics="true" pageId="60" pageNumber="69">C. yamu</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[707,796,1610,1636]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="ertou">
<emphasis box="[707,796,1610,1636]" italics="true" pageId="60" pageNumber="69">C. ertou</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[810,898,1610,1636]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="gudu">
<emphasis box="[810,898,1610,1636]" italics="true" pageId="60" pageNumber="69">C. gudu</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[962,1076,1610,1636]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="69" phylum="Arthropoda" rank="species" species="longtou">
<emphasis box="[962,1076,1610,1636]" italics="true" pageId="60" pageNumber="69">C. longtou</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>