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<document ID-DOI="10.11646/zootaxa.5264.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836172" ID-ZooBank="47E141C8-F69A-422E-B7A7-E1AF4AD201BE" approvalRequired="6" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="3" checkinTime="1681726002599" checkinUser="plazi" docAuthor="Nozaki, Tsubasa" docDate="2023" docId="03AB87F4FFB18B2AFF21FB01FEB83620" docLanguage="en" docName="zootaxa.5264.1.4.pdf" docOrigin="Zootaxa 5264 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrikothelyna limbata Nozaki 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="71" masterDocId="FF92FF8CFFB48B2DFFB6FFCEFFF93055" masterDocTitle="Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan" masterLastPageNumber="76" masterPageNumber="64" pageNumber="69" updateTime="1681726413705" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nozaki, Tsubasa</mods:namePart>
<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan &amp; The Kyushu University Museum, Fukuoka 812 - 8581, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5264</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>64</mods:start>
<mods:end>76</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.1.4</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.1.4</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7836172</mods:identifier>
<mods:identifier type="ZooBank">47E141C8-F69A-422E-B7A7-E1AF4AD201BE</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7836199" ID-Zenodo-Dep="7836199" LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620" httpUri="http://treatment.plazi.org/id/03AB87F4FFB18B2AFF21FB01FEB83620" lastPageId="7" lastPageNumber="71" pageId="5" pageNumber="69">
<subSubSection box="[151,540,1231,1258]" pageId="5" pageNumber="69" type="nomenclature">
<paragraph blockId="5.[151,672,1231,1330]" box="[151,540,1231,1258]" pageId="5" pageNumber="69">
<heading bold="true" box="[151,540,1231,1258]" fontSize="11" level="1" pageId="5" pageNumber="69" reason="1">
<emphasis bold="true" box="[151,540,1231,1258]" pageId="5" pageNumber="69">
<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[151,439,1231,1258]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="69" phylum="Arthropoda" rank="species" species="limbata" status="sp. nov.">
<emphasis bold="true" box="[151,439,1231,1258]" italics="true" pageId="5" pageNumber="69">Andrikothelyna limbata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[446,540,1231,1257]" pageId="5" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="69" type="description">
<paragraph blockId="5.[151,672,1231,1330]" box="[151,407,1267,1293]" pageId="5" pageNumber="69">
(
<figureCitation box="[159,316,1267,1293]" captionStart="FIGURES 18" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 18. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="5" pageNumber="69">Figs 23, 67</figureCitation>
,
<figureCitation box="[328,400,1267,1293]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="5" pageNumber="69">2028</figureCitation>
)
</paragraph>
<paragraph blockId="5.[151,672,1231,1330]" box="[151,672,1303,1330]" pageId="5" pageNumber="69">(Japanese name: tsumaki-toguro-hanekakushi)</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="69" type="diagnosis">
<paragraph blockId="5.[151,1437,1375,2013]" pageId="5" pageNumber="69">
<emphasis bold="true" box="[151,274,1375,1401]" pageId="5" pageNumber="69">Diagnosis.</emphasis>
<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[284,554,1375,1401]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="69" phylum="Arthropoda" rank="species" species="limbata" status="sp. nov.">
<emphasis box="[284,554,1375,1401]" italics="true" pageId="5" pageNumber="69">Andrikothelyna limbata</emphasis>
</taxonomicName>
<emphasis bold="true" box="[565,658,1375,1401]" pageId="5" pageNumber="69">
<taxonomicNameLabel box="[565,658,1375,1401]" pageId="5" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName authorityName="Nozaki" authorityYear="2023" baseAuthorityName="Pace" baseAuthorityYear="2013" box="[820,941,1376,1402]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="69" phylum="Arthropoda" rank="species" species="orientis">
<emphasis box="[820,941,1376,1402]" italics="true" pageId="5" pageNumber="69">A. orientis</emphasis>
</taxonomicName>
but can be distinguished based on yellow semicircular maculation on the posterior margin of elytra (
<figureCitation box="[816,919,1411,1437]" captionStart="FIGURES 18" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 18. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="5" pageNumber="69">Figs 23</figureCitation>
). This new species also may be distinguished from other species of the genus based on the following characteristics: posterior margin of abdominal tergite VIII having four teeth (
<figureCitation box="[360,445,1483,1509]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="5" pageNumber="69">Fig. 20</figureCitation>
); median lobe of male aedeagus, bulb, and flagellum spiraled three-dimensionally (
<figureCitation captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="5" pageNumber="69">Figs 2325</figureCitation>
), flagellum having cluster of spines at apex; extremely narrow paramere (
<figureCitation box="[1092,1180,1519,1545]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="5" pageNumber="69">Fig. 26</figureCitation>
); basal part of female spermatheca (
<figureCitation box="[306,434,1555,1581]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="5" pageNumber="69">Figs 2728</figureCitation>
) bent at the middle.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="70" pageId="5" pageNumber="69" type="materials_examined">
<paragraph blockId="5.[151,1437,1375,2013]" lastBlockId="6.[151,1437,150,573]" lastPageId="6" lastPageNumber="70" pageId="5" pageNumber="69">
<emphasis bold="true" box="[199,433,1591,1617]" pageId="5" pageNumber="69">Material examined.</emphasis>
<materialsCitation collectingDate="2016-06-03" collectionCode="KUM" pageId="5" pageNumber="69" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[441,557,1591,1617]" pageId="5" pageNumber="69">
<typeStatus box="[441,552,1591,1617]" pageId="5" pageNumber="69">Holotype</typeStatus>
.
</emphasis>
Male, “JAPAN Kyushu / Kaijin Shrine / TSUSHIMA Is. /
<date box="[1224,1352,1591,1618]" pageId="5" pageNumber="69" value="2016-06-03">
<collectingDate box="[1224,1352,1591,1618]" pageId="5" pageNumber="69" value="2016-06-03">3. VI. 2016</collectingDate>
</date>
/ Tateo ITO leg.” (
<collectionCode box="[278,348,1627,1653]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectorName="Honshu &amp; Gifu-ken" country="Japan" county="Kani-shi" elevation="285" latitude="33.38" location="Sengen-yama" longLatPrecision="723" longitude="137.12" municipality="Kukuri" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis bold="true" box="[368,839,1627,1653]" pageId="5" pageNumber="69">
<typeStatus box="[368,490,1627,1653]" pageId="5" pageNumber="69">Paratypes</typeStatus>
.
<collectingCountry box="[503,597,1627,1653]" name="Japan" pageId="5" pageNumber="69">JAPAN</collectingCountry>
:
<collectorName box="[611,709,1627,1653]" pageId="5" pageNumber="69">Honshu</collectorName>
:
<collectorName box="[722,834,1627,1653]" pageId="5" pageNumber="69">Gifu-ken</collectorName>
:
</emphasis>
1³,
<specimenCount box="[907,942,1627,1653]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB18B28FC0BF995FBA13620" box="[957,1112,1627,1653]" country="Japan" county="Kani-shi" latitude="33.38" longLatPrecision="723" longitude="137.12" municipality="Kukuri" name="Sengen-yama" pageId="5" pageNumber="69">Sengen-yama</location>
,
<collectingMunicipality box="[1125,1207,1627,1653]" pageId="5" pageNumber="69">Kukuri</collectingMunicipality>
,
<collectingCounty box="[1220,1316,1627,1653]" pageId="5" pageNumber="69">Kani-shi</collectingCounty>
[
<geoCoordinate box="[1332,1432,1627,1653]" degrees="33.38" direction="north" orientation="latitude" pageId="5" pageNumber="69" precision="555" value="33.38">33.38°N</geoCoordinate>
,
<geoCoordinate box="[151,259,1663,1689]" degrees="137.12" direction="east" orientation="longitude" pageId="5" pageNumber="69" precision="555" value="137.12">137.12°E</geoCoordinate>
] (alt.
<quantity box="[321,449,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="2.85" metricValueMax="3.7" metricValueMin="2.0" pageId="5" pageNumber="69" unit="m" value="285.0" valueMax="370.0" valueMin="200.0">
<elevation box="[321,449,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="2.85" metricValueMax="3.7" metricValueMin="2.0" pageId="5" pageNumber="69" unit="m" value="285.0" valueMax="370.0" valueMin="200.0">200370 m</elevation>
</quantity>
)
</materialsCitation>
<materialsCitation collectingDate="2018-08-07" collectionCode="KUM" collectorName="Masato Ando &amp; Kawakita Nature &amp; Athletic Park" country="Japan" county="Yawata-shi" location="Kyoto-fu" municipality="Hayashinomoto" pageId="5" pageNumber="69" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
,
<date box="[468,602,1663,1689]" pageId="5" pageNumber="69" value="2018-08-07">
<collectingDate box="[468,602,1663,1689]" pageId="5" pageNumber="69" value="2018-08-07">7.VIII.2018</collectingDate>
</date>
,
<collectorName box="[612,761,1663,1690]" pageId="5" pageNumber="69">Masato Ando</collectorName>
leg. (
<collectionCode box="[821,892,1663,1689]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
).
<emphasis bold="true" box="[909,1026,1663,1689]" pageId="5" pageNumber="69">
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB18B28FC3BF9B1FC0436CC" box="[909,1021,1663,1689]" country="Japan" county="Yawata-shi" municipality="Hayashinomoto" name="Kyoto-fu" pageId="5" pageNumber="69">Kyoto-fu</location>
:
</emphasis>
2³,
<specimenCount box="[1079,1115,1663,1689]" pageId="5" pageNumber="69" type="female">2♀</specimenCount>
,
<collectorName box="[1126,1315,1663,1690]" pageId="5" pageNumber="69">Kawakita Nature</collectorName>
&amp;
<collectorName pageId="5" pageNumber="69">Athletic Park</collectorName>
,
<collectingMunicipality box="[215,395,1699,1725]" pageId="5" pageNumber="69">Hayashinomoto</collectingMunicipality>
, 12,
<collectingCounty box="[443,571,1699,1725]" pageId="5" pageNumber="69">Yawata-shi</collectingCounty>
</materialsCitation>
,
<date box="[580,683,1699,1726]" pageId="5" pageNumber="69" value="2015-05-05">
5.
<collectionCode box="[597,619,1699,1726]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="69" type="Museum">V</collectionCode>
<materialsCitation box="[619,683,1699,1726]" collectingDate="2018-08-07" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Yawata-shi" location="Kyoto-fu" municipality="Hayashinomoto" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" typeStatus="paratype">.2015</materialsCitation>
</date>
,
<collectorName box="[693,790,1700,1726]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[850,921,1699,1725]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
); 2³,
<specimenCount box="[986,1021,1699,1725]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
, same locality,
<materialsCitation collectingDate="2015-07-11" collectionCode="KUM" collectorName="Tateo Ito &amp; Kawakita Nature &amp; Athletic Park" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<date box="[1191,1329,1699,1726]" pageId="5" pageNumber="69" value="2015-07-11">
<collectingDate box="[1191,1329,1699,1726]" pageId="5" pageNumber="69" value="2015-07-11">11.VII.2015</collectingDate>
</date>
,
<collectorName box="[1338,1436,1700,1726]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[208,279,1735,1761]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
); 3³,
<specimenCount box="[350,387,1735,1761]" pageId="5" pageNumber="69" type="female">4♀</specimenCount>
,
<collectorName box="[400,591,1735,1762]" pageId="5" pageNumber="69">Kawakita Nature</collectorName>
&amp;
<collectorName box="[627,780,1735,1761]" pageId="5" pageNumber="69">Athletic Park</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB18B28FCAEF909FC6036B4" box="[792,921,1735,1761]" country="Japan" county="Otokuni-gun" municipality="Oyamazaki-cho" name="Motohiraki" pageId="5" pageNumber="69">Motohiraki</location>
,
<collectingMunicipality box="[933,1113,1735,1761]" pageId="5" pageNumber="69">Oyamazaki-cho</collectingMunicipality>
,
<collectingCounty box="[1125,1271,1735,1761]" pageId="5" pageNumber="69">Otokuni-gun</collectingCounty>
</materialsCitation>
,
<materialsCitation collectingDate="2015-08-19" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date box="[1283,1432,1735,1762]" pageId="5" pageNumber="69" value="2015-08-19">
<collectingDate box="[1283,1432,1735,1762]" pageId="5" pageNumber="69" value="2015-08-19">19.VIII.2015</collectingDate>
</date>
,
<collectorName box="[151,252,1772,1798]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (cI); 1³,
<specimenCount box="[415,450,1771,1797]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
, same locality
</materialsCitation>
,
<materialsCitation box="[627,1202,1771,1798]" collectingDate="2015-08-22" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date box="[627,776,1771,1798]" pageId="5" pageNumber="69" value="2015-08-22">
<collectingDate box="[627,776,1771,1798]" pageId="5" pageNumber="69" value="2015-08-22">22.VIII.2015</collectingDate>
</date>
,
<collectorName box="[788,888,1772,1798]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (cI);
<specimenCount box="[1001,1036,1771,1797]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
, same locality
</materialsCitation>
,
<materialsCitation collectingDate="2015-08-26" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="1" typeStatus="paratype">
<date box="[1214,1362,1771,1798]" pageId="5" pageNumber="69" value="2015-08-26">
<collectingDate box="[1214,1362,1771,1798]" pageId="5" pageNumber="69" value="2015-08-26">26.VIII.2015</collectingDate>
</date>
,
<collectorName pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[253,324,1807,1833]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
); 1³, same locality
</materialsCitation>
,
<materialsCitation box="[568,1173,1807,1834]" collectingDate="2015-11-02" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date box="[568,684,1807,1834]" pageId="5" pageNumber="69" value="2015-11-02">
<collectingDate box="[568,684,1807,1834]" pageId="5" pageNumber="69" value="2015-11-02">2.XI.2015</collectingDate>
</date>
,
<collectorName box="[699,802,1808,1834]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[873,944,1807,1833]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
);
<specimenCount box="[967,1002,1807,1833]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
, same locality
</materialsCitation>
,
<materialsCitation collectingDate="2015-09-28" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="1" typeStatus="paratype">
<date box="[1188,1318,1807,1834]" pageId="5" pageNumber="69" value="2015-09-28">
<collectingDate box="[1188,1318,1807,1834]" pageId="5" pageNumber="69" value="2015-09-28">28.IX.2015</collectingDate>
</date>
,
<collectorName box="[1333,1436,1808,1834]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[210,281,1843,1869]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
); 2³, same locality
</materialsCitation>
,
<materialsCitation box="[522,1126,1843,1872]" collectingDate="2015-10-27" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="1" typeStatus="paratype">
<date box="[522,643,1843,1870]" pageId="5" pageNumber="69" value="2015-10-27">
<collectingDate box="[522,643,1843,1870]" pageId="5" pageNumber="69" value="2015-10-27">27.X.2015</collectingDate>
</date>
,
<collectorName box="[657,759,1844,1870]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[828,899,1843,1869]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
); 1³, same locality
</materialsCitation>
,
<materialsCitation collectingDate="2016-03-17" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="1" typeStatus="paratype">
<date box="[1140,1269,1843,1869]" pageId="5" pageNumber="69" value="2016-03-17">
<collectingDate box="[1140,1269,1843,1869]" pageId="5" pageNumber="69" value="2016-03-17">17.III.2016</collectingDate>
</date>
,
<collectorName box="[1282,1385,1844,1870]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[159,230,1879,1905]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
); 1³, same locality
</materialsCitation>
,
<date box="[471,588,1879,1905]" pageId="5" pageNumber="69" value="2020-05-14">
14.
<collectionCode box="[502,524,1879,1905]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="69" type="Museum">V</collectionCode>
<materialsCitation box="[523,588,1879,1905]" collectingDate="2016-03-17" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="2" specimenCount-female="2" typeStatus="paratype">.2020</materialsCitation>
</date>
,
<collectorName box="[602,705,1880,1906]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (cI); 6³,
<specimenCount box="[876,912,1879,1905]" pageId="5" pageNumber="69" type="female">2♀</specimenCount>
, same locality,
<date box="[1094,1210,1879,1905]" pageId="5" pageNumber="69" value="2020-06-01">
1.
<collectionCode box="[1107,1141,1879,1905]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="69">VI</collectionCode>
<materialsCitation box="[1141,1210,1879,1905]" collectingDate="2016-03-17" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="2" specimenCount-female="2" typeStatus="paratype">.2020</materialsCitation>
</date>
,
<collectorName box="[1224,1327,1880,1906]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (cI); 1 unsexed, 1³,
<specimenCount box="[333,370,1915,1941]" pageId="5" pageNumber="69" type="female">2♀</specimenCount>
, same locality,
<date box="[549,679,1915,1941]" pageId="5" pageNumber="69" value="2020-06-17">
17.
<collectionCode box="[580,613,1915,1941]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="69">VI</collectionCode>
<materialsCitation box="[613,679,1915,1941]" collectingDate="2016-03-17" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Otokuni-gun" location="Motohiraki" municipality="Oyamazaki-cho" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" typeStatus="paratype">.2020</materialsCitation>
</date>
,
<collectorName box="[693,794,1916,1942]" pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[862,933,1915,1941]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
); 1³,
<specimenCount box="[1006,1041,1915,1941]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
, same locality,
<materialsCitation collectingDate="2020-07-16" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" location="Kawara" municipality="Maidzuru-shi" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date box="[1222,1361,1915,1941]" pageId="5" pageNumber="69" value="2020-07-16">
<collectingDate box="[1222,1361,1915,1941]" pageId="5" pageNumber="69" value="2020-07-16">16.VII.2020</collectingDate>
</date>
,
<collectorName pageId="5" pageNumber="69">Tateo Ito</collectorName>
leg. (
<collectionCode box="[253,324,1951,1977]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
);
<specimenCount box="[348,383,1951,1977]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB18B28FE27F86EFE1237EF" box="[401,491,1952,1978]" country="Japan" municipality="Maidzuru-shi" name="Kawara" pageId="5" pageNumber="69">Kawara</location>
,
<collectingMunicipality box="[505,659,1951,1977]" pageId="5" pageNumber="69">Maidzuru-shi</collectingMunicipality>
, (from fallen tree)
</materialsCitation>
,
<date box="[892,1022,1951,1977]" pageId="5" pageNumber="69" value="2007-06-27">
27.
<collectionCode box="[925,957,1951,1977]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="69">VI</collectionCode>
<materialsCitation box="[958,1022,1951,1977]" collectorName="Yuzo Kuroda &amp; Okayamaken" country="Japan" county="Okayama-shi" location="Handayama" municipality="Kita-ku" pageId="5" pageNumber="69" specimenCount="2" specimenCount-female="2" typeStatus="paratype">.2007</materialsCitation>
</date>
,
<collectorName box="[1037,1189,1951,1978]" pageId="5" pageNumber="69">Yuzo Kuroda</collectorName>
leg. (cI).
<emphasis bold="true" pageId="5" pageNumber="69">
<collectorName pageId="5" pageNumber="69">Okayamaken</collectorName>
:
</emphasis>
<specimenCount box="[211,247,1987,2013]" pageId="5" pageNumber="69" type="female">2♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB18B28FEB2F80DFE743788" box="[260,397,1987,2013]" country="Japan" county="Okayama-shi" municipality="Kita-ku" name="Handayama" pageId="5" pageNumber="69">Handayama</location>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB18B28FE21F80DFE0D3788" box="[407,500,1987,2013]" country="Japan" county="Okayama-shi" municipality="Kita-ku" name="Hokaiin" pageId="5" pageNumber="69">Hokaiin</location>
,
<collectingMunicipality box="[511,599,1987,2013]" pageId="5" pageNumber="69">Kita-ku</collectingMunicipality>
,
<collectingCounty box="[610,762,1987,2013]" pageId="5" pageNumber="69">Okayama-shi</collectingCounty>
,
<date box="[773,889,1987,2013]" pageId="5" pageNumber="69" value="2004-06-02">
2.
<collectionCode box="[791,823,1987,2013]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="69">VI</collectionCode>
<materialsCitation box="[823,889,1987,2013]" collectionCode="KUM" collectorName="Seidai Nagashima" country="Japan" county="Okayama-shi" location="Kasaiyama" municipality="Kita-ku" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" typeStatus="paratype">.2004</materialsCitation>
</date>
,
<collectorName box="[900,1103,1987,2013]" pageId="5" pageNumber="69">Seidai Nagashima</collectorName>
leg. (
<collectionCode box="[1166,1237,1987,2013]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="69" type="Herbarium">KUM</collectionCode>
);
<specimenCount box="[1256,1291,1987,2013]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB18B28FAAFF80DFA613788" box="[1305,1432,1987,2013]" country="Japan" county="Okayama-shi" municipality="Kita-ku" name="Kasaiyama" pageId="5" pageNumber="69">Kasaiyama</location>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFF21FF59FF0E30E4" box="[151,247,151,177]" country="Japan" county="Okayama-shi" municipality="Kita-ku" name="Hataayu" pageId="6" pageNumber="70">Hataayu</location>
,
<collectingMunicipality box="[258,346,151,177]" pageId="6" pageNumber="70">Kita-ku</collectingMunicipality>
,
<collectingCounty box="[356,508,151,177]" pageId="6" pageNumber="70">Okayama-shi</collectingCounty>
,
<materialsCitation collectingDate="1974-01-15" collectionCode="KUM" collectorName="Osamu Yamaji &amp; Kochi-ken" country="Japan" county="Shiman-cho" location="Shikoku" municipality="Kubogawa" pageId="6" pageNumber="70" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date box="[519,629,150,177]" pageId="6" pageNumber="70" value="1974-01-15">
<collectingDate box="[519,629,150,177]" pageId="6" pageNumber="70" value="1974-01-15">15.I.1974</collectingDate>
</date>
,
<collectorName box="[639,804,151,177]" pageId="6" pageNumber="70">Osamu Yamaji</collectorName>
leg. (
<collectionCode box="[866,936,151,177]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
).
<emphasis bold="true" box="[954,1203,150,176]" pageId="6" pageNumber="70">
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFC0CFF58FBDB30E5" box="[954,1058,150,176]" country="Japan" county="Shiman-cho" municipality="Kubogawa" name="Shikoku" pageId="6" pageNumber="70">Shikoku</location>
:
<collectorName box="[1069,1198,150,176]" pageId="6" pageNumber="70">Kochi-ken</collectorName>
:
</emphasis>
<specimenCount box="[1209,1244,151,177]" pageId="6" pageNumber="70" type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFB5FFF59FA6030E4" box="[1257,1433,151,177]" country="Japan" county="Shiman-cho" municipality="Kubogawa" name="Towa Uragoshi" pageId="6" pageNumber="70">Tôwa Uragoshi</location>
,
<collectingMunicipality box="[151,275,187,213]" pageId="6" pageNumber="70">Kubogawa</collectingMunicipality>
,
<collectingCounty box="[288,426,187,213]" pageId="6" pageNumber="70">Shiman-chô</collectingCounty>
</materialsCitation>
,
<date box="[438,541,187,213]" pageId="6" pageNumber="70" value="2010-05-08">
8.
<collectionCode box="[454,476,187,213]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="70" type="Museum">V</collectionCode>
<materialsCitation box="[476,541,187,213]" collectionCode="KUM" collectorName="Takasuke Miyata &amp; Toshie Miyata" country="Japan" county="Tsushima-shi" location="Kyushu" municipality="Minemachikisaka" pageId="6" pageNumber="70" specimenCount="1" typeStatus="paratype">.2010</materialsCitation>
</date>
,
<collectorName box="[553,746,187,213]" pageId="6" pageNumber="70">Takasuke Miyata</collectorName>
&amp;
<collectorName box="[783,945,187,213]" pageId="6" pageNumber="70">Toshie Miyata</collectorName>
leg. (car-net) (
<collectionCode box="[1114,1185,187,213]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
).
<emphasis bold="true" pageId="6" pageNumber="70">
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFB00FF75FAE13080" box="[1206,1304,187,213]" country="Japan" county="Tsushima-shi" municipality="Minemachikisaka" name="Kyushu" pageId="6" pageNumber="70">Kyushu</location>
:
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFA92FF74FF3E30AC" country="Japan" county="Tsushima-shi" municipality="Minemachikisaka" name="Nagasakiken" pageId="6" pageNumber="70">Nagasakiken</location>
:
</emphasis>
3³,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFEB3FF11FE0930AC" box="[261,496,223,249]" country="Japan" county="Tsushima-shi" municipality="Minemachikisaka" name="Kaijin Shinto Shrine" pageId="6" pageNumber="70">Kaijin Shinto Shrine</location>
,
<collectingMunicipality box="[508,710,223,249]" pageId="6" pageNumber="70">Minemachikisaka</collectingMunicipality>
,
<collectingCounty box="[721,873,223,249]" pageId="6" pageNumber="70">Tsushima-shi</collectingCounty>
,
<date box="[884,1000,223,249]" pageId="6" pageNumber="70" value="2016-06-03">
3.
<collectionCode box="[901,934,223,249]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="70">VI</collectionCode>
<materialsCitation box="[933,1000,223,249]" collectingDate="2016-06-03" collectionCode="KUM" collectorName="Tateo Ito" country="Japan" county="Minamisatsuma-shi" location="Kagoshima-ken" municipality="Oura-cho" pageId="6" pageNumber="70" specimenCount="1" specimenCount-female="1" typeStatus="paratype">.2016</materialsCitation>
</date>
,
<collectorName box="[1012,1112,223,249]" pageId="6" pageNumber="70">Tateo Ito</collectorName>
leg. (same data as holotype) (
<collectionCode box="[161,231,259,285]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
).
<emphasis bold="true" box="[250,448,258,284]" pageId="6" pageNumber="70">
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFF4CFECCFE423149" box="[250,443,258,284]" country="Japan" county="Minamisatsuma-shi" municipality="Oura-cho" name="Kagoshima-ken" pageId="6" pageNumber="70">Kagoshima-ken</location>
:
</emphasis>
1³,
<specimenCount box="[507,541,259,285]" pageId="6" pageNumber="70" type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFD9BFECAFD5B3148" box="[557,674,259,287]" country="Japan" county="Minamisatsuma-shi" municipality="Oura-cho" name="Oura river" pageId="6" pageNumber="70">Ôura river</location>
estuary,
<collectingMunicipality box="[776,885,260,287]" pageId="6" pageNumber="70">Ôura-chô</collectingMunicipality>
,
<collectingCounty box="[897,1121,259,285]" pageId="6" pageNumber="70">Minamisatsuma-shi</collectingCounty>
,
<date box="[1134,1237,259,285]" pageId="6" pageNumber="70" value="1994-05-04">
4.
<collectionCode box="[1152,1174,259,285]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="70" type="Museum">V</collectionCode>
<materialsCitation box="[1174,1237,259,285]" collectionCode="KUM" collectorName="Kaoru Haga &amp; Amami-Oshima Is." country="Japan" county="Mt. Yuwan" location="Ryukyu" municipality="Kagoshima-ken" pageId="6" pageNumber="70" specimenCount="1" specimenCount-female="1" typeStatus="paratype">.1994</materialsCitation>
</date>
,
<collectorName box="[1249,1387,259,285]" pageId="6" pageNumber="70">Kaoru Haga</collectorName>
leg. (
<collectionCode box="[160,230,295,321]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
).
<emphasis bold="true" box="[250,803,294,324]" pageId="6" pageNumber="70">
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFF4CFEE9FEA53114" box="[250,348,295,321]" country="Japan" county="Mt. Yuwan" municipality="Kagoshima-ken" name="Ryukyu" pageId="6" pageNumber="70">Ryukyu</location>
:
<collectingMunicipality box="[361,554,294,320]" pageId="6" pageNumber="70">Kagoshima-ken</collectingMunicipality>
:
<collectorName box="[565,797,295,324]" pageId="6" pageNumber="70">Amami-Ôshima Is.</collectorName>
:
</emphasis>
<specimenCount box="[811,845,295,321]" pageId="6" pageNumber="70" type="female">1♀</specimenCount>
,
<collectingCounty box="[860,989,295,321]" pageId="6" pageNumber="70">Mt. Yuwan</collectingCounty>
.
<materialsCitation collectingDate="1971-04-19" collectionCode="EUM" collectorName="M. Sakai" country="Japan" location="Hatsuno" pageId="6" pageNumber="70" specimenCount="1" typeStatus="paratype">
<date box="[1000,1126,295,321]" pageId="6" pageNumber="70" value="1971-04-19">
<collectingDate box="[1000,1126,295,321]" pageId="6" pageNumber="70" value="1971-04-19">19.IV.1971</collectingDate>
</date>
,
<collectorName box="[1139,1240,295,321]" pageId="6" pageNumber="70">M. Sakai</collectorName>
leg. (
<collectionCode box="[1306,1374,295,321]" pageId="6" pageNumber="70">EUM</collectionCode>
); 1³,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFF21FE85FF0E3130" box="[151,247,331,357]" country="Japan" name="Hatsuno" pageId="6" pageNumber="70">Hatsuno</location>
</materialsCitation>
,
<date box="[259,362,331,357]" pageId="6" pageNumber="70" value="1977-05-04">
4.
<collectionCode box="[277,299,331,357]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="70" type="Museum">V</collectionCode>
<materialsCitation box="[299,362,331,357]" collectionCode="EUM" collectorName="M. Sakai &amp; A. Oda &amp; Ishigaki Is." country="Japan" county="Ishigaki-shi" location="Okinawa-ken" municipality="Shiraho" pageId="6" pageNumber="70" specimenCount="4" specimenCount-female="4" typeStatus="paratype">.1977</materialsCitation>
</date>
,
<collectorName box="[374,468,331,357]" pageId="6" pageNumber="70">M.Sakai</collectorName>
&amp;
<collectorName box="[503,585,331,357]" pageId="6" pageNumber="70">A. Oda</collectorName>
leg. (
<collectionCode box="[649,717,331,357]" pageId="6" pageNumber="70">EUM</collectionCode>
).
<emphasis bold="true" box="[736,1057,330,357]" pageId="6" pageNumber="70">
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFD56FE84FC7F3131" box="[736,902,330,356]" country="Japan" county="Ishigaki-shi" municipality="Shiraho" name="Okinawa-ken" pageId="6" pageNumber="70">Okinawa-ken</location>
:
<collectorName box="[914,1051,330,357]" pageId="6" pageNumber="70">Ishigaki Is.</collectorName>
:
</emphasis>
10³,
<specimenCount box="[1127,1164,331,357]" pageId="6" pageNumber="70" type="female">4♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFB2FFE85FA613130" box="[1177,1432,331,357]" country="Japan" county="Ishigaki-shi" municipality="Shiraho" name="Reverbed of Toro Riv." pageId="6" pageNumber="70">Reverbed of Tôro Riv.</location>
,
<collectingMunicipality box="[151,241,367,393]" pageId="6" pageNumber="70">Shiraho</collectingMunicipality>
,
<collectingCounty box="[254,393,367,393]" pageId="6" pageNumber="70">Ishigaki-shi</collectingCounty>
.,
<date box="[408,568,367,393]" pageId="6" pageNumber="70" value="2021-05-28" valueMax="2021-05-29" valueMin="2021-05-28">
2829.
<collectionCode box="[485,506,367,393]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="70" type="Museum">V</collectionCode>
<materialsCitation box="[506,568,367,393]" collectionCode="KUM" collectorName="N. Tsuji &amp; S. Imada" country="Japan" county="Ishigaki-shi" location="Inoda Auto Camping Ground" municipality="Tozato" pageId="6" pageNumber="70" specimenCount="1" typeStatus="paratype">.2021</materialsCitation>
</date>
,
<collectorName box="[580,672,367,393]" pageId="6" pageNumber="70">N. Tsuji</collectorName>
&amp;
<collectorName box="[710,811,367,393]" pageId="6" pageNumber="70">S. Imada</collectorName>
leg. (
<collectionCode box="[877,947,367,393]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
, one pair for cI); 2³,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFB19FEA1FF1631F8" country="Japan" county="Ishigaki-shi" municipality="Tozato" name="Inoda Auto Camping Ground" pageId="6" pageNumber="70">Inoda Auto Camping Ground</location>
,
<collectingMunicipality box="[252,333,403,429]" pageId="6" pageNumber="70">Tôzato</collectingMunicipality>
,
<collectingCounty box="[345,480,403,429]" pageId="6" pageNumber="70">Ishigaki-shi</collectingCounty>
,
<materialsCitation box="[493,1205,403,429]" collectingDate="2020-03-19" collectionCode="KUM" collectorName="Takuto Hashizume" country="Japan" county="Ishigaki-shi" location="Inoda Auto Camping Ground" municipality="Tozato" pageId="6" pageNumber="70" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date box="[493,621,403,429]" pageId="6" pageNumber="70" value="2020-03-19">
<collectingDate box="[493,621,403,429]" pageId="6" pageNumber="70" value="2020-03-19">19.III.2020</collectingDate>
</date>
,
<collectorName box="[634,845,403,429]" pageId="6" pageNumber="70">Takuto Hashizume</collectorName>
leg. (
<collectionCode box="[912,983,403,429]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
);
<specimenCount box="[1004,1039,403,429]" pageId="6" pageNumber="70" type="female">1♀</specimenCount>
, same locality
</materialsCitation>
,
<materialsCitation collectingDate="2020-03-22" collectionCode="KUM" collectorName="Takuto Hashizume" country="Japan" county="Ishigaki-shi" location="Inoda Auto Camping Ground" municipality="Tozato" pageId="6" pageNumber="70" specimenCount="1" typeStatus="paratype">
<date box="[1218,1347,403,429]" pageId="6" pageNumber="70" value="2020-03-22">
<collectingDate box="[1218,1347,403,429]" pageId="6" pageNumber="70" value="2020-03-22">22.III.2020</collectingDate>
</date>
,
<collectorName pageId="6" pageNumber="70">Takuto Hashizume</collectorName>
leg. (
<collectionCode box="[342,413,439,465]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
); 1 unsexed, same locality
</materialsCitation>
,
<materialsCitation box="[724,1432,439,465]" collectingDate="2020-03-23" collectionCode="KUM" collectorName="Takuto Hashizume" country="Japan" county="Ishigaki-shi" location="Inoda Auto Camping Ground" municipality="Tozato" pageId="6" pageNumber="70" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date box="[724,852,439,465]" pageId="6" pageNumber="70" value="2020-03-23">
<collectingDate box="[724,852,439,465]" pageId="6" pageNumber="70" value="2020-03-23">23.III.2020</collectingDate>
</date>
,
<collectorName box="[864,1076,439,465]" pageId="6" pageNumber="70">Takuto Hashizume</collectorName>
leg. (
<collectionCode box="[1140,1211,439,465]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
);
<specimenCount box="[1232,1267,439,465]" pageId="6" pageNumber="70" type="female">1♀</specimenCount>
, same locality
</materialsCitation>
,
<materialsCitation box="[151,1174,474,503]" collectingDate="2020-10-26" collectionCode="KUM" collectorName="Tsubasa Nozaki" country="Japan" latitude="24.46" location="Inoda" longLatPrecision="750" longitude="124.25" municipality="Ishigaki-shi" pageId="6" pageNumber="70" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<date box="[151,272,475,501]" pageId="6" pageNumber="70" value="2020-10-26">
<collectingDate box="[151,272,475,501]" pageId="6" pageNumber="70" value="2020-10-26">26.X.2020</collectingDate>
</date>
,
<collectorName box="[284,463,475,501]" pageId="6" pageNumber="70">Tsubasa Nozaki</collectorName>
leg. (
<collectionCode box="[528,599,475,501]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
); 6³,
<specimenCount box="[669,706,475,501]" pageId="6" pageNumber="70" type="female">2♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AB87F4FFB18B2AFF21FB01FEB83620:8EDD6039FFB28B2BFD79FE15FCEB31A0" box="[719,786,475,501]" country="Japan" latitude="24.46" longLatPrecision="750" longitude="124.25" municipality="Ishigaki-shi" name="Inoda" pageId="6" pageNumber="70">Inoda</location>
,
<collectingMunicipality box="[798,930,475,501]" pageId="6" pageNumber="70">Ishigaki-shi</collectingMunicipality>
[
<geoCoordinate box="[945,1045,475,501]" degrees="24.46" direction="north" orientation="latitude" pageId="6" pageNumber="70" precision="555" value="24.46">24.46°N</geoCoordinate>
,
<geoCoordinate box="[1057,1168,474,501]" degrees="124.25" direction="east" orientation="longitude" pageId="6" pageNumber="70" precision="555" value="124.25">124.25°E</geoCoordinate>
]
</materialsCitation>
,
<date box="[1187,1304,475,501]" pageId="6" pageNumber="70" value="2021-05-30">
30.
<collectionCode box="[1220,1242,475,501]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="70" type="Museum">V</collectionCode>
.2021
</date>
,
<collectorName box="[1316,1407,475,501]" pageId="6" pageNumber="70">N. Tsuji</collectorName>
&amp;
<collectorName box="[151,251,511,537]" pageId="6" pageNumber="70">S. Imada</collectorName>
leg. (
<collectionCode box="[315,385,511,537]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="70" type="Herbarium">KUM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="71" pageId="6" pageNumber="70" type="description">
<paragraph blockId="6.[151,1437,150,573]" box="[199,1169,546,573]" pageId="6" pageNumber="70">
<emphasis bold="true" box="[199,414,546,573]" pageId="6" pageNumber="70">Description. Body</emphasis>
(
<figureCitation box="[429,585,547,573]" captionStart="FIGURES 18" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 18. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="6" pageNumber="70">Figs 23, 67</figureCitation>
) robust; dorsal surface glossy, distinctly reticulated.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836182" ID-Zenodo-Dep="7836182" httpUri="https://zenodo.org/record/7836182/files/figure.png" pageId="6" pageNumber="70" startId="6.[151,264,1900,1924]" targetBox="[197,1383,622,1849]" targetPageId="6">
<paragraph blockId="6.[151,1437,1900,1960]" pageId="6" pageNumber="70">
<emphasis bold="true" box="[151,340,1900,1924]" pageId="6" pageNumber="70">FIGURES 1619.</emphasis>
<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[346,618,1900,1924]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="70" phylum="Arthropoda" rank="species" species="rubiginosa" status="sp. nov.">
<emphasis box="[346,618,1900,1924]" italics="true" pageId="6" pageNumber="70">Andrikothelyna rubiginosa</emphasis>
</taxonomicName>
<emphasis bold="true" box="[624,704,1901,1925]" pageId="6" pageNumber="70">
<taxonomicNameLabel box="[624,704,1901,1925]" pageId="6" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female. 16―Meso- and metaventrite; 17―abdominal tergite VIII; 18― abdominal sternite VIII; 19―spermatheca.
</paragraph>
</caption>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,272,150,176]" pageId="7" pageNumber="71">Color.</emphasis>
Body dark brown to black, pronotum brownish red to dark brown, posterior margin of each elytron with yellow semicircular maculation; antennae dark brown with segments I and II brownish yellow and the third yellowish brown, maxillary palpi dark brown to brownish yellow; forelegs reddish yellow to yellow, meso- and metalegs reddish brown to yellowish brown.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,270,295,321]" pageId="7" pageNumber="71">Head.</emphasis>
Eyes large, as long as 1/3 of head; infraorbital carina developed.Antennae longer than head and pronotum combined; segments IIX slightly dilated; segments VX distinctly transverse; segment XI elongate, round at apex; approximate relative length of segments from basal to apex: 16: 18: 13: 10: 11: 10: 11: 10: 10: 11: 38.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,295,403,429]" pageId="7" pageNumber="71">Thorax.</emphasis>
Pronotum, postero-lateral angle obtuse. Elytra, postero-lateral angle gently sinuate; longitudinal row of granules along suture strong; near lateral margin gently carinate, granulated, and curved upward (
<figureCitation box="[1318,1423,439,465]" captionStart="FIGURES 18" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 18. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="7" pageNumber="71">Figs 67</figureCitation>
). Mesoventral process, apex varying truncated to round.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,323,510,536]" pageId="7" pageNumber="71">Abdomen.</emphasis>
Tergite VIII (
<figureCitation box="[475,555,511,537]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="7" pageNumber="71">Fig. 20</figureCitation>
) weakly granulated with two medial granules; posterior margin with four teeth; lateral teeth long, less curved, medial teeth slightly shorter than lateral teeth; medial grub distinctly shallower than lateral grub, depth 1/3 of lateral grub.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,262,619,645]" pageId="7" pageNumber="71">Legs.</emphasis>
Moderately slender in general; tibiae covered with fine setae. Approximate relative length of mesotarsal segments from basal to apex: 32: 10: 11: 34, and that of metatarsal segments: 42: 11: 10: 10: 28.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,267,691,717]" pageId="7" pageNumber="71">Male.</emphasis>
Tergite VII with two posteromedial granules; posterior margin of sternite VIII (
<figureCitation box="[1169,1251,691,717]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="7" pageNumber="71">Fig. 21</figureCitation>
) without medial notch and finely sinuate. Median lobe of aedeagus (
<figureCitation box="[748,879,726,753]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="7" pageNumber="71">Figs 2325</figureCitation>
) length about
<quantity box="[1047,1151,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="71" unit="mm" value="0.75">0.75 mm</quantity>
, very narrow; bulb itself spiraled three-dimensionally; flagellum spiraled, having flag-like appendage at base and cluster of spines at apex. Paramere (
<figureCitation box="[272,354,799,825]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="7" pageNumber="71">Fig. 26</figureCitation>
) very narrow; velum remarkably elongate; apical lobe of paramerite digitate.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,293,835,861]" pageId="7" pageNumber="71">Female.</emphasis>
Tergite VII without or with one posteromedial granule; posterior margin of sternite VIII (
<figureCitation box="[1291,1372,835,861]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="7" pageNumber="71">Fig. 22</figureCitation>
) with medial notch. Spermatheca (
<figureCitation box="[470,601,871,897]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="7" pageNumber="71">Figs 2728</figureCitation>
), apical part short and subspherical; basal part smooth and bent at middle.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,382,907,933]" pageId="7" pageNumber="71">Measurements.</emphasis>
(population of Honshu and Kyushu, n = 5): BL ≈ 1.362.96; AL, 0.670.86; PL, 0.30.42; PW, 0.590.8; EL, 0.320.46; EW, 0.660.94; HTL, 0.360.52; PW/PL, 1.851.95; EW/EL, 22.08; AL/PL, 1.882.21. Measurement (population of Ishigaki Is., n = 5): BL ≈ 1.842.32; AL, 0.620.8; PL, 0.320.42; PW, 0.560.74; EL, 0.330.44; EW, 0.620.88; HTL, 0.340.48; PW/PL, 1.751.98; EW/EL, 1.882; AL/PL, 1.922.07.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,319,1050,1076]" pageId="7" pageNumber="71">Variation.</emphasis>
Sexual dimorphism was evident in the number of posteromedial granules on tergite VII and a presence of a notch on sternite VIII.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
Geographical variations were detected in body coloration, the degree of gloss on the dorsal surface, and the shape of the spermatheca in this species. The population from Honshu and Kyushu (
<figureCitation box="[1129,1236,1159,1185]" captionStart="FIGURES 18" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 18. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="7" pageNumber="71">Figs 2, 6</figureCitation>
,
<figureCitation box="[1250,1279,1159,1185]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="7" pageNumber="71">27</figureCitation>
) revealed the following characteristics: body color dark brown; pronotum brownish red; maxillary palpi dark brown; forelegs reddish yellow; meso- and metalegs reddish brown; body surface less glossy; spermatheca weakly sclerotized, whereas the population from Ryukyu (
<figureCitation box="[584,686,1267,1293]" captionStart="FIGURES 18" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 18. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="7" pageNumber="71">Figs 3, 7</figureCitation>
,
<figureCitation box="[698,727,1267,1293]" captionStart="FIGURES 2028" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 2028. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 2325―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="7" pageNumber="71">28</figureCitation>
) exhibited the following characteristics: body black; pronotum dark brown; maxillary palpi brownish yellow; forelegs yellow; meso- and metalegs yellowish brown; body surface glossier compared to that of the other populations, antennae longer than that in the other.
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">Regardless of the region, intra-species variations were found in the following characters: the apical shape of the mesoventral process and the presence of a posteromedial granule of female tergite VII.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="71" type="etymology">
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,334,1447,1473]" pageId="7" pageNumber="71">Etymology.</emphasis>
The specific name of the species is derived from the Latin adjective “
<emphasis box="[1084,1180,1447,1473]" italics="true" pageId="7" pageNumber="71">limbâtus</emphasis>
,” meaning “bordered,” and refers to yellow maculation on the posterior edge of the elytra.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="71" type="distribution">
<paragraph blockId="7.[151,1437,150,1653]" box="[199,1218,1518,1547]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,353,1518,1544]" pageId="7" pageNumber="71">Distribution.</emphasis>
<collectingCountry box="[359,427,1519,1545]" name="Japan" pageId="7" pageNumber="71">Japan</collectingCountry>
: Honshu, Shikoku, Kyushu, Ryukyu (Amami Ôshima Is., Ishigaki Is.).
</paragraph>
<paragraph blockId="7.[151,1437,150,1653]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[199,331,1554,1580]" pageId="7" pageNumber="71">Bionomics.</emphasis>
On Ishigaki Island, this new species was spotted walking on the dead branches of the living trunks of beach hibiscus,
<taxonomicName box="[359,575,1591,1617]" class="Magnoliopsida" family="Malvaceae" genus="Talipariti" kingdom="Plantae" order="Malvales" pageId="7" pageNumber="71" phylum="Tracheophyta" rank="species" species="tiliaceum">
<emphasis box="[359,575,1591,1617]" italics="true" pageId="7" pageNumber="71">Talipariti tiliaceum</emphasis>
</taxonomicName>
(L.) Fryxell (
<figureCitation box="[727,855,1591,1617]" captionStart="FIGURES 2930" captionStartId="9.[151,264,678,702]" captionTargetBox="[151,1436,181,655]" captionTargetId="figure-32@9.[151,1436,181,655]" captionTargetPageId="9" captionText="FIGURES 2930. Habitat of Andrikothelyna limbata sp. nov. 29―dead branch of beach hibiscus, Talipariti tiliaceum from which the species is collected; 30―the zoomed image of the arrowed area." figureDoi="http://doi.org/10.5281/zenodo.7836191" httpUri="https://zenodo.org/record/7836191/files/figure.png" pageId="7" pageNumber="71">Figs 2930</figureCitation>
) and observed walking on the branches at night and were collected.
</paragraph>
</subSubSection>
</treatment>
</document>