283 lines
41 KiB
XML
283 lines
41 KiB
XML
<document id="4DED798BAB79EF6F4563292022E886F1" ID-DOI="10.11646/zootaxa.5352.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8407445" ID-ZooBank="0483BDD2-C037-4BBB-98F4-81BA67944984" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1696430284000" checkinUser="plazi" docAuthor="Loktionov, Valery M., Lelej, Arkady S., Vasilenko, Dmitry V. & Perkovsky, Evgeny E." docDate="2023" docId="03AB87D6A328B761FF555C71FD15FED0" docLanguage="en" docName="zootaxa.5352.3.6.pdf" docOrigin="Zootaxa 5352 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5352.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paleogenia waichertae Loktionov, Lelej et Perkovsky 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="430" masterDocId="FF92FFAEA329B765FFC25A63FF86FF91" masterDocTitle="Discovery of a new extinct spider wasp (Hymenoptera, Pompilidae) from Eocene Rovno amber" masterLastPageNumber="432" masterPageNumber="426" pageNumber="427" updateTime="1696430675636" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0704017A3BF55311C2BB55F3575220F5">Discovery of a new extinct spider wasp (Hymenoptera, Pompilidae) from Eocene Rovno amber</mods:title>
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<mods:name id="CBC2878E2623971C1481D000435D883A" type="personal">
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<mods:roleTerm id="29192B7CA08F48123A1B01B6A7A24E92">Author</mods:roleTerm>
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<mods:namePart id="43F98FE2035175C562FA77EBD4E9A76E">Loktionov, Valery M.</mods:namePart>
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<mods:affiliation id="641C2F301688B959A1BB986B46937B55">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia.</mods:affiliation>
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<mods:namePart id="3D901E2370A07193AB6B91EABE640DC0">Lelej, Arkady S.</mods:namePart>
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<mods:affiliation id="A980C318C46B9E52B1C34A7794D99C93">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia.</mods:affiliation>
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<mods:namePart id="37AEFA4B61DC913A12E366A7B633B9C8">Vasilenko, Dmitry V.</mods:namePart>
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<mods:affiliation id="40CBB60DED27779896A3B6538B278F08">Cherepovets State University, Lunacharsky Prospect 5, Cherepovets, Vologda Region, 162600, Russia. & Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya 123, Moscow, 117647, Russia.</mods:affiliation>
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<mods:namePart id="A0B5B6FF546CCA5F2B1D4B5A39468EB6">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="292801F90603334979DDFA9D73290E9B">Natural History Museum of Denmark, Universitetsparken 15, Copenhagen, 2100, Denmark.</mods:affiliation>
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<mods:date id="CA0FDDE9DA7AD4C1CAE409340D993A9F">2023</mods:date>
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<mods:number id="27EB6E42C23F88113C33FF0E563D55C9">2023-10-04</mods:number>
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<mods:classification id="F2D426A668293AEABC7D81F0AA89F042">journal article</mods:classification>
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<treatment id="03AB87D6A328B761FF555C71FD15FED0" LSID="urn:lsid:plazi:treatment:03AB87D6A328B761FF555C71FD15FED0" httpUri="http://treatment.plazi.org/id/03AB87D6A328B761FF555C71FD15FED0" lastPageId="4" lastPageNumber="430" pageId="1" pageNumber="427">
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<subSubSection id="C318654BA328B764FF555C71FC14F9BC" box="[151,914,1554,1581]" pageId="1" pageNumber="427" type="nomenclature">
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<paragraph id="8BBD36C0A328B764FF555C71FC14F9BC" blockId="1.[151,914,1554,1617]" box="[151,914,1554,1581]" pageId="1" pageNumber="427">
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<heading id="D0F581ACA328B764FF555C71FC14F9BC" bold="true" box="[151,914,1554,1581]" fontSize="11" level="1" pageId="1" pageNumber="427" reason="1">
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<emphasis id="B976EAD2A328B764FF555C71FC14F9BC" bold="true" box="[151,914,1554,1581]" pageId="1" pageNumber="427">
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<taxonomicName id="4C024D43A328B764FF555C71FCAEF9BC" authority="Loktionov, Lelej et Perkovsky" authorityName="Loktionov, Lelej et Perkovsky" authorityYear="2023" box="[151,808,1554,1581]" class="Insecta" family="Pompilidae" genus="Paleogenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="species" species="waichertae" status="sp. nov.">
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<emphasis id="B976EAD2A328B764FF555C71FE2CF9BC" bold="true" box="[151,426,1554,1581]" italics="true" pageId="1" pageNumber="427">Paleogenia waichertae</emphasis>
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Loktionov, Lelej et Perkovsky
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="A24557A9A328B764FCF75C70FC14F9BC" box="[821,914,1555,1581]" pageId="1" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C318654BA328B764FF555C54FE80F9C0" box="[151,262,1591,1617]" pageId="1" pageNumber="427" type="description">
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<paragraph id="8BBD36C0A328B764FF555C54FE80F9C0" blockId="1.[151,914,1554,1617]" box="[151,262,1591,1617]" pageId="1" pageNumber="427">
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<figureCitation id="13392A45A328B764FF555C54FE80F9C0" box="[151,262,1591,1617]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–10" captionStartId-0="2.[151,264,1911,1935]" captionStartId-1="3.[151,264,1938,1962]" captionTargetBox-0="[151,1436,669,1887]" captionTargetBox-1="[152,1435,181,1914]" captionTargetId-0="figure-199@2.[151,1436,669,1887]" captionTargetId-1="figure-27@3.[152,1435,181,1914]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." captionText-1="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi-0="http://doi.org/10.5281/zenodo.8407447" figureDoi-1="http://doi.org/10.5281/zenodo.8407449" httpUri-0="https://zenodo.org/record/8407447/files/figure.png" httpUri-1="https://zenodo.org/record/8407449/files/figure.png" pageId="1" pageNumber="427">Figs 1–10</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C318654BA328B764FF555C1CFCF4F894" pageId="1" pageNumber="427" type="materials_examined">
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||
<paragraph id="8BBD36C0A328B764FF555C1CFCF4F894" blockId="1.[151,1437,1662,2013]" pageId="1" pageNumber="427">
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||
<emphasis id="B976EAD2A328B764FF555C1CFEC7F909" bold="true" box="[151,321,1662,1689]" pageId="1" pageNumber="427">Type material</emphasis>
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.
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<typeStatus id="54B98862A328B764FE915C1CFE38F908" box="[339,446,1663,1689]" pageId="1" pageNumber="427">Holotype</typeStatus>
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, ♁:
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<collectionCode id="ED13AE05A328B764FE365C1CFDB4F908" box="[500,562,1663,1689]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="427">SIZK</collectionCode>
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K-3286, Klesov,
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<collectingRegion id="49C6F822A328B764FCC65C1CFCC9F908" box="[772,847,1663,1689]" country="Ukraine" name="Rivne" pageId="1" pageNumber="427">Rovno</collectingRegion>
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amber, late Eocene. Syninclusions:
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<collectionCode id="ED13AE05A328B764FB345C1CFAB2F908" box="[1270,1332,1663,1689]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="427">SIZK</collectionCode>
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K-3287:
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<taxonomicName id="4C024D43A328B764FF555CC0FF6DF92C" box="[151,235,1699,1725]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
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(female of
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<taxonomicName id="4C024D43A328B764FE435CC0FDC3F92C" box="[385,581,1699,1725]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
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),
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<collectionCode id="ED13AE05A328B764FD9A5CC0FD10F92C" box="[600,662,1699,1725]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="427">SIZK</collectionCode>
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K-3288:
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<taxonomicName id="4C024D43A328B764FCCD5CC0FC50F92C" authority=", SIZK" authorityName="SIZK" box="[783,982,1699,1725]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="order">Coleoptera, SIZK</taxonomicName>
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K-3289:
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<taxonomicName id="4C024D43A328B764FB8F5CC0FB70F92C" authorityName="Borner" authorityYear="1904" box="[1101,1270,1699,1725]" class="Insecta" kingdom="Animalia" order="Archaeognatha" pageId="1" pageNumber="427" phylum="Arthropoda" rank="order">Archaeognatha</taxonomicName>
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(
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<taxonomicName id="4C024D43A328B764FACE5CC0FA16F92C" authorityName="Grassi" authorityYear="1888" box="[1292,1424,1699,1725]" class="Insecta" family="Machilidae" kingdom="Animalia" order="Archaeognatha" pageId="1" pageNumber="427" phylum="Arthropoda" rank="family">Machilidae</taxonomicName>
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), 2
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<taxonomicName id="4C024D43A328B764FF6E5CA4FEC2F970" authorityName="Anatomy Consortium" authorityYear="2013" box="[172,324,1735,1761]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
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(1
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<taxonomicName id="4C024D43A328B764FEB75CA4FDCFF970" baseAuthorityName="Debauche" baseAuthorityYear="1948" box="[373,585,1735,1761]" class="Insecta" family="Mymarommatidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="family">Mymarommatidae</taxonomicName>
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), 5 Acari (larvae of Parasitengona).
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<typeStatus id="54B98862A328B764FC345CA4FBDDF970" box="[1014,1115,1735,1761]" pageId="1" pageNumber="427">Paratype</typeStatus>
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, ♁:
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<collectionCode id="ED13AE05A328B764FB565CA4FB54F970" box="[1172,1234,1735,1761]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="427">SIZK</collectionCode>
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K-8178, Klesov,
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<collectingRegion id="49C6F822A328B764FF555C88FF64F894" box="[151,226,1771,1797]" country="Ukraine" name="Rivne" pageId="1" pageNumber="427">Rovno</collectingRegion>
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amber, late Eocene. Syninclusion:
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<collectionCode id="ED13AE05A328B764FDAF5C88FD2DF894" box="[621,683,1771,1797]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="427">SIZK</collectionCode>
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K-8179:
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<taxonomicName id="4C024D43A328B764FCD45C88FCE8F894" box="[790,878,1771,1797]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C318654BA328B764FF055D6DFADAF804" pageId="1" pageNumber="427" type="diagnosis">
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||
<paragraph id="8BBD36C0A328B764FF055D6DFADAF804" blockId="1.[151,1437,1662,2013]" pageId="1" pageNumber="427">
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<emphasis id="B976EAD2A328B764FF055D6DFEBCF8B9" bold="true" box="[199,314,1806,1832]" pageId="1" pageNumber="427">Diagnosis</emphasis>
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. Male of the new species clearly differs from those of all other congeners in the combination of the following characters: the tibial spur 0.8–0.9 × as long as the first tarsomere; the apical portion of the hypopygium well-exposed beyond the tergum 7 and the sternum 6, rod-shaped and slightly widened apically, bearing erect setae in ventral view (
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<figureCitation id="13392A45A328B764FE8C5D18FE11F804" box="[334,407,1914,1941]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1911,1935]" captionTargetBox="[151,1436,669,1887]" captionTargetId="figure-199@2.[151,1436,669,1887]" captionTargetPageId="2" captionText="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="1" pageNumber="427">Figs 5</figureCitation>
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,
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<figureCitation id="13392A45A328B764FE615D18FE35F804" box="[419,435,1915,1941]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="1" pageNumber="427">6</figureCitation>
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); the gena 0.2 × wider than the eye in lateral view (
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<figureCitation id="13392A45A328B764FC315D18FBBEF804" box="[1011,1080,1915,1941]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="1" pageNumber="427">Fig. 8</figureCitation>
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). The female is unknown.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C318654BA328B767FF055DFDFE4EFDAC" lastPageId="2" lastPageNumber="428" pageId="1" pageNumber="427" type="description">
|
||
<paragraph id="8BBD36C0A328B767FF055DFDFBE2FF68" blockId="1.[151,1437,1662,2013]" lastBlockId="2.[151,1437,150,645]" lastPageId="2" lastPageNumber="428" pageId="1" pageNumber="427">
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<emphasis id="B976EAD2A328B764FF055DFDFED7F829" bold="true" box="[199,337,1950,1976]" pageId="1" pageNumber="427">Description</emphasis>
|
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. MALE. Body length:
|
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<quantity id="4CFA9B25A328B764FD905DFCFD21F828" box="[594,679,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="427" unit="mm" value="3.2">3.2 mm</quantity>
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||
(
|
||
<typeStatus id="54B98862A328B764FD775DFCFC9BF828" box="[693,797,1951,1977]" pageId="1" pageNumber="427">holotype</typeStatus>
|
||
), 3.0 mm (
|
||
<typeStatus id="54B98862A328B764FC505DFCFC7FF828" box="[914,1017,1951,1977]" pageId="1" pageNumber="427">paratype</typeStatus>
|
||
). Head without noticeable erect setae. Eyes large; eye 0.48 × as wide as median interocular distance (head in frontal view) (
|
||
<figureCitation id="13392A45A328B764FB895DA0FB09F84C" box="[1099,1167,1987,2013]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="1" pageNumber="427">Fig. 6</figureCitation>
|
||
). Clypeus large, convex in dorsal view, trapezoid; 0.58 × as long as wide; its apical margin nearly straight; its posterolateral corners rounded (
|
||
<figureCitation id="13392A45A32BB767FF5D5AD8FF61FF44" box="[159,231,187,213]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="2" pageNumber="428">Fig. 6</figureCitation>
|
||
). Mandible apex bidentate. Gena narrow, 0.2 × as wide as eye (head in lateral view) (
|
||
<figureCitation id="13392A45A32BB767FB1C5AD8FAA0FF44" box="[1246,1318,187,213]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="2" pageNumber="428">Fig. 8</figureCitation>
|
||
). Antenna shortened, with 11 cylindrical flagellomeres (
|
||
<figureCitation id="13392A45A32BB767FD515ABCFD5DFF68" box="[659,731,223,249]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1911,1935]" captionTargetBox="[151,1436,669,1887]" captionTargetId="figure-199@2.[151,1436,669,1887]" captionTargetPageId="2" captionText="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="2" pageNumber="428">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="13392A45A32BB767FD255ABCFC94FF68" box="[743,786,223,249]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="2" pageNumber="428">6, 8</figureCitation>
|
||
); apical flagellomere pointed.
|
||
</paragraph>
|
||
<paragraph id="8BBD36C0A32BB767FF055B60FCDCFED0" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="428">
|
||
Metapostnotum well developed, long (
|
||
<figureCitation id="13392A45A32BB767FDBE5B60FD65FE8C" box="[636,739,259,285]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="2" pageNumber="428">Figs 8, 9</figureCitation>
|
||
). Propodeum smooth, with dense micropubescence, in lateral view gently sloping, its posterior surface nearly flat (
|
||
<figureCitation id="13392A45A32BB767FD255B44FCA9FED0" box="[743,815,295,321]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1911,1935]" captionTargetBox="[151,1436,669,1887]" captionTargetId="figure-199@2.[151,1436,669,1887]" captionTargetPageId="2" captionText="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="2" pageNumber="428">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="13392A45A32BB767FCFE5B44FCCAFED0" box="[828,844,295,321]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="2" pageNumber="428">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD36C0A32BB767FF055B28FCE5FE3C" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="428">
|
||
Tergum 1 not petiolate basally, gradually widening towards apex (
|
||
<figureCitation id="13392A45A32BB767FC775B28FC7DFEF4" box="[949,1019,331,357]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1911,1935]" captionTargetBox="[151,1436,669,1887]" captionTargetId="figure-199@2.[151,1436,669,1887]" captionTargetPageId="2" captionText="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="2" pageNumber="428">Fig. 3</figureCitation>
|
||
). Terga with dense micropubescence (
|
||
<figureCitation id="13392A45A32BB767FF5D5B0CFF61FE18" box="[159,231,367,393]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1911,1935]" captionTargetBox="[151,1436,669,1887]" captionTargetId="figure-199@2.[151,1436,669,1887]" captionTargetPageId="2" captionText="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="2" pageNumber="428">Fig. 3</figureCitation>
|
||
). Apical portion of hypopygium well exposed beoynd tergum 7 and sternum 6, rod-shaped and slightly widened apically, bearing erect setae in ventral view (
|
||
<figureCitation id="13392A45A32BB767FD325BF0FCBFFE3C" box="[752,825,402,429]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1911,1935]" captionTargetBox="[151,1436,669,1887]" captionTargetId="figure-199@2.[151,1436,669,1887]" captionTargetPageId="2" captionText="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="2" pageNumber="428">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="13392A45A32BB767FC875BF0FCD3FE3C" box="[837,853,403,429]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="2" pageNumber="428">6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD36C0A32BB767FF055BD4FAE2FE64" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="428">
|
||
Legs without noticeable erect spines (
|
||
<figureCitation id="13392A45A32BB767FDAA5BD4FD29FE40" box="[616,687,439,465]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1911,1935]" captionTargetBox="[151,1436,669,1887]" captionTargetId="figure-199@2.[151,1436,669,1887]" captionTargetPageId="2" captionText="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="2" pageNumber="428">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="13392A45A32BB767FD785BD4FD72FE40" box="[698,756,439,465]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="2" pageNumber="428">6, 10</figureCitation>
|
||
). Tibia with short apical spine-like setae. Tibial spur of meso- and metalegs 0.8–0.9 × as long as basal tarsomere (
|
||
<figureCitation id="13392A45A32BB767FD145BB8FC99FE64" box="[726,799,475,501]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1911,1935]" captionTargetBox="[151,1436,669,1887]" captionTargetId="figure-199@2.[151,1436,669,1887]" captionTargetPageId="2" captionText="FIGURES 1–5. Holotype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-3286. 1, habitus, total dorsolateral view. 2, head and antenna, frontolateral view. 3, propodeum and metasoma, dorso-lateral view. 4, meso- and metatibia and tarsi, lateral view (arrow points hypopygium). 5, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="2" pageNumber="428">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="13392A45A32BB767FCE95BB8FCCCFE64" box="[811,842,475,501]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1938,1962]" captionTargetBox="[152,1435,181,1914]" captionTargetId="figure-27@3.[152,1435,181,1914]" captionTargetPageId="3" captionText="FIGURES 6–10. Paratype of Paleogenia waichertae Loktionov, Lelej et Perkovsky, sp. nov., ♁, SIZK K-8178. 6, habitus, total ventral view. 7, habitus, total lateral view. 8, mesosoma, head and antenna, lateral view. 9, head, mesosoma and metasoma, lateral view. 10, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale)." figureDoi="http://doi.org/10.5281/zenodo.8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="2" pageNumber="428">10</figureCitation>
|
||
). Tarsal claws very small, inner tooth indistinct.
|
||
</paragraph>
|
||
<paragraph id="8BBD36C0A32BB767FF055B9CFD25FD88" blockId="2.[151,1437,150,645]" box="[199,675,511,537]" pageId="2" pageNumber="428">Wings crumpled (characters are invisible).</paragraph>
|
||
<paragraph id="8BBD36C0A32BB767FF055840FE4EFDAC" blockId="2.[151,1437,150,645]" box="[199,456,547,573]" pageId="2" pageNumber="428">FEMALE is unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318654BA32BB767FF055825FF66FD14" pageId="2" pageNumber="428" type="distribution">
|
||
<paragraph id="8BBD36C0A32BB767FF055825FF66FD14" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="428">
|
||
<emphasis id="B976EAD2A32BB767FF055825FE41FDF0" bold="true" box="[199,455,582,609]" pageId="2" pageNumber="428">Distribution and age</emphasis>
|
||
.
|
||
<collectingRegion id="49C6F822A32BB767FE185824FDA3FDF0" box="[474,549,583,609]" country="Ukraine" name="Rivne" pageId="2" pageNumber="428">Rovno</collectingRegion>
|
||
amber,
|
||
<collectingCountry id="F3157650A32BB767FD4A5824FD60FDF0" box="[648,742,583,609]" name="Ukraine" pageId="2" pageNumber="428">Ukraine</collectingCountry>
|
||
. Priabonian, 33.9-37.8 Ma, Upper Eocene (
|
||
<bibRefCitation id="EF934B31A32BB767FB2B5824FF55FD14" author="Colombo, W. D. & Perkovsky, E. E. & Vasilenko, D. V." pageId="2" pageNumber="428" pagination="429 - 434" refId="ref3045" refString="Colombo, W. D., Perkovsky, E. E. & Vasilenko, D. V. (2021) The first sclerodermine flat wasp (Hymenoptera: Bethylidae) from the upper Eocene Rovno amber Ukraine. Alcheringa, 45 (4), 429 - 434. https: // doi. org / 10.1080 / 03115518.2021.2006311" type="journal article" year="2021">
|
||
Colombo
|
||
<emphasis id="B976EAD2A32BB767FA9D5824FA1BFDF0" box="[1375,1437,583,609]" italics="true" pageId="2" pageNumber="428">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7D6648A32BB767FF555D14FC0EF86A" ID-DOI="http://doi.org/10.5281/zenodo.8407447" ID-Zenodo-Dep="8407447" httpUri="https://zenodo.org/record/8407447/files/figure.png" pageId="2" pageNumber="428" startId="2.[151,264,1911,1935]" targetBox="[151,1436,669,1887]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BBD36C0A32BB767FF555D14FC0EF86A" blockId="2.[151,1437,1911,2043]" pageId="2" pageNumber="428">
|
||
<emphasis id="B976EAD2A32BB767FF555D14FEB1F81E" bold="true" box="[151,311,1911,1935]" pageId="2" pageNumber="428">FIGURES 1–5</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C024D43A32BB767FE0E5D14FC78F81E" authority="Loktionov, Lelej et Perkovsky" authorityName="Loktionov, Lelej et Perkovsky" authorityYear="2023" box="[460,1022,1911,1935]" class="Insecta" family="Pompilidae" genus="Paleogenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="428" phylum="Arthropoda" rank="species" species="waichertae" status="sp. nov.">
|
||
<emphasis id="B976EAD2A32BB767FE0E5D14FD3CF81E" box="[460,698,1911,1935]" italics="true" pageId="2" pageNumber="428">Paleogenia waichertae</emphasis>
|
||
Loktionov, Lelej et Perkovsky
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A24557A9A32BB767FBC95D14FBE6F81F" box="[1035,1120,1911,1935]" pageId="2" pageNumber="428" rank="species">
|
||
<emphasis id="B976EAD2A32BB767FBC95D14FBDCF81E" bold="true" box="[1035,1114,1911,1935]" pageId="2" pageNumber="428">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, ♁, SIZK K-3286.
|
||
<emphasis id="B976EAD2A32BB767FAF35D14FAB8F81E" bold="true" box="[1329,1342,1911,1935]" pageId="2" pageNumber="428">1</emphasis>
|
||
, habitus, total dorsolateral view.
|
||
<emphasis id="B976EAD2A32BB767FE455DF8FE12F822" bold="true" box="[391,404,1947,1971]" pageId="2" pageNumber="428">2</emphasis>
|
||
, head and antenna, frontolateral view.
|
||
<emphasis id="B976EAD2A32BB767FCE35DF8FCA8F822" bold="true" box="[801,814,1947,1971]" pageId="2" pageNumber="428">3</emphasis>
|
||
, propodeum and metasoma, dorso-lateral view.
|
||
<emphasis id="B976EAD2A32BB767FADB5DF8FAA0F822" bold="true" box="[1305,1318,1947,1971]" pageId="2" pageNumber="428">4</emphasis>
|
||
, meso- and metatibia and tarsi, lateral view (arrow points hypopygium).
|
||
<emphasis id="B976EAD2A32BB767FCC65DDCFC97F846" bold="true" box="[772,785,1983,2007]" pageId="2" pageNumber="428">5</emphasis>
|
||
, apical portion of metasoma, hypopygium and legs part. (arrow points hypopygium), ventral view. Scale bar 1 mm (Figs 2–5 not to scale).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7D6648A32AB766FF555DF1FB5CF863" ID-DOI="http://doi.org/10.5281/zenodo.8407449" ID-Zenodo-Dep="8407449" httpUri="https://zenodo.org/record/8407449/files/figure.png" pageId="3" pageNumber="429" startId="3.[151,264,1938,1962]" targetBox="[152,1435,181,1914]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BBD36C0A32AB766FF555DF1FB5CF863" blockId="3.[151,1437,1938,2034]" pageId="3" pageNumber="429">
|
||
<emphasis id="B976EAD2A32AB766FF555DF1FEC5F83B" bold="true" box="[151,323,1938,1962]" pageId="3" pageNumber="429">FIGURES 6–10</emphasis>
|
||
. Paratype of
|
||
<taxonomicName id="4C024D43A32AB766FE135DF1FB87F83B" authority="Loktionov, Lelej et Perkovsky" authorityName="Loktionov, Lelej et Perkovsky" authorityYear="2023" box="[465,1025,1938,1962]" class="Insecta" family="Pompilidae" genus="Paleogenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="waichertae" status="sp. nov.">
|
||
<emphasis id="B976EAD2A32AB766FE135DF1FD38F83B" box="[465,702,1938,1962]" italics="true" pageId="3" pageNumber="429">Paleogenia waichertae</emphasis>
|
||
Loktionov, Lelej et Perkovsky
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A24557A9A32AB766FBCC5DF1FBE4F83B" box="[1038,1122,1938,1962]" pageId="3" pageNumber="429" rank="species">
|
||
<emphasis id="B976EAD2A32AB766FBCC5DF1FBDAF83B" bold="true" box="[1038,1116,1938,1962]" pageId="3" pageNumber="429">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, ♁, SIZK K-8178.
|
||
<emphasis id="B976EAD2A32AB766FAF35DF1FAB8F83B" bold="true" box="[1329,1342,1938,1962]" pageId="3" pageNumber="429">6</emphasis>
|
||
, habitus, total ventral view.
|
||
<emphasis id="B976EAD2A32AB766FE905DD5FED9F85F" bold="true" box="[338,351,1974,1998]" pageId="3" pageNumber="429">7</emphasis>
|
||
, habitus, total lateral view.
|
||
<emphasis id="B976EAD2A32AB766FDB65DD5FD07F85F" bold="true" box="[628,641,1974,1998]" pageId="3" pageNumber="429">8</emphasis>
|
||
, mesosoma, head and antenna, lateral view.
|
||
<emphasis id="B976EAD2A32AB766FBFD5DD5FBCAF85F" bold="true" box="[1087,1100,1974,1998]" pageId="3" pageNumber="429">9</emphasis>
|
||
, head, mesosoma and metasoma, lateral view.
|
||
<emphasis id="B976EAD2A32AB766FEDB5DB9FEB4F863" bold="true" box="[281,306,2010,2034]" pageId="3" pageNumber="429">10</emphasis>
|
||
, fore leg part., meso- and metalegs, lateral view. Scale bar 1 mm (Figs 6, 8–10 not to scale).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C318654BA32DB761FF055AF4FE1DFF68" pageId="4" pageNumber="430" type="etymology">
|
||
<paragraph id="8BBD36C0A32DB761FF055AF4FE1DFF68" blockId="4.[151,1436,151,321]" pageId="4" pageNumber="430">
|
||
<emphasis id="B976EAD2A32DB761FF055AF4FECEFF20" bold="true" box="[199,328,151,177]" pageId="4" pageNumber="430">Etymology</emphasis>
|
||
. The new species is named for Cecilia Waichert (Complexo Biopráticas, Universidade Vila Velha, Vila Velha,
|
||
<collectingCountry id="F3157650A32DB761FED95AD8FEE2FF44" box="[283,356,187,213]" name="Brazil" pageId="4" pageNumber="430">Brazil</collectingCountry>
|
||
), one of the most active pompilid researchers that made a great contribution to the taxonomy of extant and extinct taxa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318654BA32DB761FF055B60FD15FED0" pageId="4" pageNumber="430" type="discussion">
|
||
<paragraph id="8BBD36C0A32DB761FF055B60FD15FED0" blockId="4.[151,1436,151,321]" pageId="4" pageNumber="430">
|
||
<emphasis id="B976EAD2A32DB761FF055B60FEAFFE8C" bold="true" box="[199,297,259,285]" pageId="4" pageNumber="430">Remark</emphasis>
|
||
. Size of the piece and syninclusions clearly indicates that
|
||
<typeStatus id="54B98862A32DB761FC7B5B60FB9DFE8C" box="[953,1051,259,285]" pageId="4" pageNumber="430">holotype</typeStatus>
|
||
was caught in the resin trap at the base of the trunk (see
|
||
<bibRefCitation id="EF934B31A32DB761FE4C5B44FD01FED0" author="Perkovsky, E. E. & Rasnitsyn, A. P. & Vlaskin, A. P. & Rasnitsyn, S. P." box="[398,647,295,321]" pageId="4" pageNumber="430" pagination="293 - 301" refId="ref4279" refString="Perkovsky, E. E., Rasnitsyn, A. P., Vlaskin, A. P. & Rasnitsyn, S. P. (2012) Contribution to the study of the structure of amber forest communities based on analysis of syninclusions in the Rovno amber (Late Eocene of Ukraine). Paleontological Journal, 46 (3), 293 - 301. https: // doi. org / 10.1134 / S 0031030112030136" type="journal article" year="2012">
|
||
Perkovsky
|
||
<emphasis id="B976EAD2A32DB761FDC95B44FDBAFED0" box="[523,572,295,321]" italics="true" pageId="4" pageNumber="430">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |