180 lines
22 KiB
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180 lines
22 KiB
XML
<document id="749E0745BD65DAF8CBDE6716278BDADC" ID-DOI="10.11646/zootaxa.1074.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5050779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625108368558" checkinUser="felipe" docAuthor="Ponder, W. F., Clark, S. A., Eberhard, S. & Studdert, J. B." docDate="2005" docId="03AB87BA2D25A225E41FF9027FEAFA65" docLanguage="en" docName="zt01074p066.pdf" docOrigin="Zootaxa 1074 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1074.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Nanocochlea Ponder and Clark 1993" docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="FF92FFC22D02A20CE517FFCC7E3DFF83" masterDocTitle="A radiation of hydrobiid snails in the caves and streams at Precipitous Bluff southwest Tasmania, Australia (Mollusca: Caenogastropoda: Rissooidea: Hydrobiidae s. l.)," masterLastPageNumber="66" masterPageNumber="1" pageNumber="40" updateTime="1698981223375" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="43AFD9C19F802F335C8A5395DA492B55">A radiation of hydrobiid snails in the caves and streams at Precipitous Bluff southwest Tasmania, Australia (Mollusca: Caenogastropoda: Rissooidea: Hydrobiidae s. l.),</mods:title>
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<mods:name id="FAB394C5683C08B29011F9EEA9C514AE" type="personal">
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<mods:roleTerm id="E2EE7D9FC7BADEDFD4EFE09DD5D0ECEE">Author</mods:roleTerm>
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<mods:namePart id="D3F779CD1893C985FFDD9E3A8D829B01">Ponder, W. F.</mods:namePart>
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<mods:affiliation id="CD7D1EED74AA503B16D7235214F321F3">Australian Museum, 6 College Street, Sydney, NSW, 2010, Australia. Email: winstonp @ austmus. gov. au Previously Australian Museum, now Department of Biological Sciences, Biodiversity and Systematics. University of Alabama, Box 870345, Tuscaloosa, AL 35487, United States of America Department of Conservation and Land Management, Science Division, PO Box 51, Wanneroo, WA 6065, Australia</mods:affiliation>
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<mods:roleTerm id="A258D5DD4BB7D2E1BD371C4E904658D8">Author</mods:roleTerm>
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<mods:namePart id="AAE945A3F0503E0940D5D6BD80F056D8">Clark, S. A.</mods:namePart>
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<mods:namePart id="D9F5F9A629A7ED34A0631DCFA50AAAFD">Eberhard, S.</mods:namePart>
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</mods:role>
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<mods:namePart id="FE67DB69F9450B7CBC414E600E14B1CD">Studdert, J. B.</mods:namePart>
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<mods:affiliation id="66365C087829DF9941095E6DFFC81AAE">Australian Museum, 6 College Street, Sydney, NSW, 2010, Australia. Email: winstonp @ austmus. gov. au Previously Australian Museum, now Department of Biological Sciences, Biodiversity and Systematics. University of Alabama, Box 870345, Tuscaloosa, AL 35487, United States of America Department of Conservation and Land Management, Science Division, PO Box 51, Wanneroo, WA 6065, Australia</mods:affiliation>
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<mods:typeOfResource id="05AF22BC55C0AFAC7300FB577A2AF5EB">text</mods:typeOfResource>
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<mods:title id="7C46E4AD00AC45A1DC7571703A606DC5">Zootaxa</mods:title>
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<mods:part id="8D28F51A0F5A47218B839103C6E6C380">
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<mods:date id="E5A15B1D4E5DF803189D51A3CC0C92B0">2005</mods:date>
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<mods:number id="84792D57B573A9831E283809044F0256">2005-11-01</mods:number>
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<mods:number id="A7E27B6190A059F1AABB89F4F79F1942">1074</mods:number>
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<mods:classification id="5AA5708CF6DE64BEF499514080979FCD">journal article</mods:classification>
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<mods:identifier id="35F65F4A5757E1A50D4233E9FC374757" type="DOI">10.11646/zootaxa.1074.1.1</mods:identifier>
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<treatment id="03AB87BA2D25A225E41FF9027FEAFA65" ID-DOI="http://doi.org/10.5281/zenodo.5053153" ID-Zenodo-Dep="5053153" LSID="urn:lsid:plazi:treatment:03AB87BA2D25A225E41FF9027FEAFA65" httpUri="http://treatment.plazi.org/id/03AB87BA2D25A225E41FF9027FEAFA65" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40">
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<subSubSection id="C31865272D25A22BE41FF9027A34F96B" box="[264,1033,1742,1768]" pageId="39" pageNumber="40" type="nomenclature">
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<paragraph id="8BBD36AC2D25A22BE41FF9027A34F96B" blockId="39.[264,1033,1742,1768]" box="[264,1033,1742,1768]" pageId="39" pageNumber="40">
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<heading id="D0F581C02D25A22BE41FF9027A34F96B" bold="true" box="[264,1033,1742,1768]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
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<emphasis id="B976EABE2D25A22BE41FF9027A34F96B" bold="true" box="[264,1033,1742,1768]" pageId="39" pageNumber="40">
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Genus
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<taxonomicName id="4C024D2F2D25A22BE448F9027CD5F96B" authority="Ponder and Clark" authorityName="Ponder and Clark" authorityYear="1993" box="[351,744,1742,1768]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
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<emphasis id="B976EABE2D25A22BE448F9027FC3F96B" bold="true" box="[351,510,1742,1768]" italics="true" pageId="39" pageNumber="40">Nanocochlea</emphasis>
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Ponder and Clark
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</taxonomicName>
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(in Ponder
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<emphasis id="B976EABE2D25A22BE66BF9027D92F96B" bold="true" box="[892,943,1742,1768]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
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.), 1993
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31865272D25A224E41FF8D67CE2FD45" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="materials_examined">
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<paragraph id="8BBD36AC2D25A22BE41FF8D67F9DF8DF" blockId="39.[264,1323,1818,1884]" pageId="39" pageNumber="40">
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<typeStatus id="54B9880E2D25A22BE41FF8D67F7FF8B7" box="[264,322,1818,1844]" pageId="39" pageNumber="40">Type</typeStatus>
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species:
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<taxonomicName id="4C024D2F2D25A22BE4A4F8D67D97F8B7" authority="Ponder and Clark" authorityName="Ponder and Clark" box="[435,938,1818,1844]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="monticola">
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<emphasis id="B976EABE2D25A22BE4A4F8D67CF0F8B0" box="[435,717,1818,1843]" italics="true" pageId="39" pageNumber="40">Nanocochlea monticola</emphasis>
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Ponder and Clark
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</taxonomicName>
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<emphasis id="B976EABE2D25A22BE6A3F8D67DF6F8B0" box="[948,971,1818,1843]" italics="true" pageId="39" pageNumber="40">in</emphasis>
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<bibRefCitation id="EF934B5D2D25A22BE6C2F8D67A86F8B7" author="Ponder, W. F. & Clark, G. A. & Miller, A. & Toluzzi, A." box="[981,1211,1818,1844]" pageId="39" pageNumber="40" pagination="501 - 750" refId="ref28895" refString="Ponder, W. F., Clark, G. A., Miller, A. & Toluzzi, A. (1993) On a major radiation of freshwater snails in Tasmania and eastern Victoria - a preliminary overview of the Beddomeia group (Mollusca: Gastropoda: Hydrobiidae). Invertebrate Taxonomy, 7, 501 - 750." type="journal article" year="1993">
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Ponder
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<emphasis id="B976EABE2D25A22BE125F8D67A55F8B0" box="[1074,1128,1818,1843]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
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., 1993
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</bibRefCitation>
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(original designation).
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</paragraph>
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<paragraph id="8BBD36AC2D2AA224E41FFED07FAFFEB6" blockId="40.[264,1324,284,1870]" box="[264,402,284,309]" pageId="40" pageNumber="41">
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<emphasis id="B976EABE2D2AA224E41FFED07FAFFEB6" box="[264,402,284,309]" italics="true" pageId="40" pageNumber="41">Description</emphasis>
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</paragraph>
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<paragraph id="8BBD36AC2D2AA224E42FFE8F7CE2FD45" blockId="40.[264,1324,284,1870]" pageId="40" pageNumber="41">
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<emphasis id="B976EABE2D2AA224E42FFE8F7F4BFEDE" bold="true" box="[312,374,323,349]" pageId="40" pageNumber="41">Shell</emphasis>
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. Minute to small in size (adults range from
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<quantity id="4CFA9B492D2AA224E661FE887A25FEDD" box="[886,1048,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="40" pageNumber="41" unit="mm" value="2.05" valueMax="2.5" valueMin="1.6">1.6 to 2.5 mm</quantity>
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in length), pupiform to elongate pupiform or elongateconic. Protoconch of about 1.4–1.5 whorls, typically sculptured with uniform pitting; separation of protoconch from teleoconch distinct, with varixlike border. Teleoconch with spire much longer than length of aperture. Aperture ovate, slightly angled posteriorly, inner lip attached to parietal wall or partially to completely detached, although not markedly so, usually lower part not much separated from base; outer lip orthocline to opisthocline, weakly thickened, simple or with very slight reflection. Periphery rounded, base simple, imperforate in adults, rarely narrowly umbilicate in juveniles. All species described below are semiopaque to opaque, white with a thin pale yellowish periostracum.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31865272D2AA225E42FFD187FEAFA65" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
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<paragraph id="8BBD36AC2D2AA224E42FFD187AE9FCBD" blockId="40.[264,1324,284,1870]" pageId="40" pageNumber="41">
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<emphasis id="B976EABE2D2AA224E42FFD187FF3FD6D" bold="true" box="[312,462,724,750]" pageId="40" pageNumber="41">Operculum.</emphasis>
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Ovate, paucispiral, flat, yellowish; columellar edge slightly convex, outer edge strongly convex; outer surface simple, paucispiral, nucleus markedly eccentric; inner surface usually smooth with white smear or (in one species) large, broad peg.
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</paragraph>
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<paragraph id="8BBD36AC2D2AA224E42FFC807A18FA9D" blockId="40.[264,1324,284,1870]" pageId="40" pageNumber="41">
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<emphasis id="B976EABE2D2AA224E42FFC807FA7FCE5" bold="true" box="[312,410,844,870]" pageId="40" pageNumber="41">Radula.</emphasis>
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Central teeth large, cutting edge broad, indented mid dorsally, with 4–5 cusps on either side of narrow sharp median cusp; narrow, unthickened lateral projections emerge at about 45º from mid laterally; base with short, narrow, rounded basal projection not extending beyond lateral projections; two basal cusps emerge from ventral face of tooth on either side of basal projection, innermost longest. Lateral teeth with short cutting edge, about 4–5 small cusps on inner side of narrow median cusp and 4–6 small cusps on outer side; neck prominent, near vertical; lateral flange more than twice as long as cutting edge; prominent Ushaped projection on base below cutting edge; inner edge short, inner side of base excavated. Inner marginal teeth with 21–38 tiny cusps on rather wide cutting edge (ratio of cutting edge to shaft on inner marginal teeth about ¼); sides approximately parallel, outer edge thickened. Outer marginal teeth narrow, thickened on inner edge, distal end with 17–29 minute cusps; ratio of cutting edge to shaft about ¼.
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</paragraph>
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<paragraph id="8BBD36AC2D2AA224E42FFAE07A5DFA15" blockId="40.[264,1324,284,1870]" pageId="40" pageNumber="41">
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<emphasis id="B976EABE2D2AA224E42FFAE07F81FAC5" bold="true" box="[312,444,1324,1350]" pageId="40" pageNumber="41">Headfoot.</emphasis>
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Simple, unpigmented, with long cephalic tentacles, unpigmented to lightly pigmented eyes present in weak bulges at outer bases of tentacles; snout of moderate length, tapering, weakly bilobed distally. Foot short, rounded posteriorly.
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</paragraph>
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<paragraph id="8BBD36AC2D2AA224E42FFA687C13F955" blockId="40.[264,1324,284,1870]" pageId="40" pageNumber="41">
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<emphasis id="B976EABE2D2AA224E42FFA687C74FA3D" bold="true" box="[312,585,1444,1470]" pageId="40" pageNumber="41">Nongenital anatomy.</emphasis>
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Pallial cavity elongate, osphradium large and oval, towards posterior end of ctenidium; ctenidium absent or with small (reduced from normal triangular
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<typeStatus id="54B9880E2D2AA224E495FA387F8BF98D" box="[386,438,1524,1550]" pageId="40" pageNumber="41">type</typeStatus>
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) ctenidial filaments; efferent vein very short to long (ctenidium occupying nearly entire length of pallial cavity to about ½). Hypobranchial gland variably developed. Kidney and pericardium usually about ½ in pallial roof; renal gland orientated longitudinally. Stomach usually with anterior chamber larger than posterior; long style sac moderately long; no posterior caecum. Rectum with long Sshaped coil; overlying but not indenting pallial oviduct.
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</paragraph>
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<paragraph id="8BBD36AC2D2AA225E42FF9287DB4FE05" blockId="40.[264,1324,284,1870]" lastBlockId="41.[264,1323,284,990]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
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<emphasis id="B976EABE2D2AA224E42FF9287CB9F97D" bold="true" box="[312,644,1764,1790]" pageId="40" pageNumber="41">Male reproductive system.</emphasis>
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Prostate gland about ½ within pallial cavity; compressed to oval in section, with very thin ventral wall; pallial vas deferens opens at about ⅓–½ length of pallial portion. Penis located on right side of head well behind base of right tentacle; lacking but has an inconspicuous to moderately developed swelling in the distal portion just behind a papillalike distal end; distal portion long and tapering; medial part simple and parallel sided; basal part wide to moderate.
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</paragraph>
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<paragraph id="8BBD36AC2D2BA225E42FFE5F7D02FC5D" blockId="41.[264,1323,284,990]" pageId="41" pageNumber="42">
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<emphasis id="B976EABE2D2BA225E42FFE5F7C95FE2E" bold="true" box="[312,680,403,429]" pageId="41" pageNumber="42">Female reproductive system.</emphasis>
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Ovary simple sac; coiled oviduct smooth, firm, not embedded in connective tissue, initially inverted Ushape, usually straight distal to seminal receptacle, usually reaching posterior end of bursa (sometimes a little more or less) then sharply bent to run anteriorly; Seminal receptacle ovoid to pyriform, with short duct, opening to oviduct, located opposite middle to ventral part of left side of bursa copulatrix. Bursa copulatrix medium to rather large, globular to pyriform, posterior to albumen gland except for small overlap on right side, either behind posterior pallial wall or extending to it; bursal duct arrises from anterior or ventroanterior wall, simple to undulating, joins oviduct dorsally at posterior pallial wall or just in front of it. Common duct straight. Albumen gland with ½ or more within pallial roof, shorter to longer than capsule gland. Capsule gland compressedoval in section, divided into three glandular zones, short anterior and posterior translucent white zones and long yellowish to orange middle zone; anterior end blunt to steeply tapering; genital opening short, terminal to slightly anterior of capsule gland. Ventral channel extended into weakly to moderately developed vestibule anteriorly. No brood pouch.
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</paragraph>
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<paragraph id="8BBD36AC2D2BA225E41FFBD87C18FBAE" blockId="41.[264,1324,1044,1150]" box="[264,549,1044,1069]" pageId="41" pageNumber="42">
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<emphasis id="B976EABE2D2BA225E41FFBD87C18FBAE" box="[264,549,1044,1069]" italics="true" pageId="41" pageNumber="42">Distribution and habitat</emphasis>
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</paragraph>
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<paragraph id="8BBD36AC2D2BA225E42FFBF07FD1FBFD" blockId="41.[264,1324,1044,1150]" pageId="41" pageNumber="42">
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Members of the genus are known from mountain lakes, seepages, streams and caves in southern
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<collectingRegion id="49C6F84E2D2BA225E463FBA87FD5FBFD" box="[372,488,1124,1150]" country="Australia" name="Tasmania" pageId="41" pageNumber="42">Tasmania</collectingRegion>
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.
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</paragraph>
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<paragraph id="8BBD36AC2D2BA225E41FFB787F53FB4E" blockId="41.[264,1323,1204,1510]" box="[264,366,1204,1229]" pageId="41" pageNumber="42">
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<emphasis id="B976EABE2D2BA225E41FFB787F53FB4E" box="[264,366,1204,1229]" italics="true" pageId="41" pageNumber="42">Remarks</emphasis>
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</paragraph>
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<paragraph id="8BBD36AC2D2BA225E42FFB107FEAFA65" blockId="41.[264,1323,1204,1510]" pageId="41" pageNumber="42">
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<taxonomicName id="4C024D2F2D2BA225E42FFB107FEFFB76" authorityName="Ponder and Clark" authorityYear="1993" box="[312,466,1244,1269]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis id="B976EABE2D2BA225E42FFB107FEFFB76" box="[312,466,1244,1269]" italics="true" pageId="41" pageNumber="42">Nanocochlea</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C024D2F2D2BA225E779FB107D2FFB76" authorityName="Ponder" authorityYear="1992" box="[622,786,1244,1269]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis id="B976EABE2D2BA225E779FB107D2FFB76" box="[622,786,1244,1269]" italics="true" pageId="41" pageNumber="42">Pseudotricula</emphasis>
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</taxonomicName>
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in its small, tallspired pupiform to elongateconic shell with a thin, simple, orthocline to opisthocline, usually nonreflected outer lip. There are no obvious anatomical differences between the two genera and there is a rather minor radular difference—the basal tongue of the central teeth is narrower in
|
||
<taxonomicName id="4C024D2F2D2BA225E41FFAB07F9EFA16" authorityName="Ponder and Clark" authorityYear="1993" box="[264,419,1404,1429]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D2BA225E41FFAB07F9EFA16" box="[264,419,1404,1429]" italics="true" pageId="41" pageNumber="42">Nanocochlea</emphasis>
|
||
</taxonomicName>
|
||
. The outer shaft on the lateral teeth is longer in species of
|
||
<taxonomicName id="4C024D2F2D2BA225E141FAB07ACDFA16" authorityName="Ponder and Clark" authorityYear="1993" box="[1110,1264,1404,1429]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D2BA225E141FAB07ACDFA16" box="[1110,1264,1404,1429]" italics="true" pageId="41" pageNumber="42">Nanocochlea</emphasis>
|
||
</taxonomicName>
|
||
than in typical species of
|
||
<taxonomicName id="4C024D2F2D2BA225E714FA687C9AFA3E" authorityName="Ponder" authorityYear="1992" box="[515,679,1444,1469]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D2BA225E714FA687C9AFA3E" box="[515,679,1444,1469]" italics="true" pageId="41" pageNumber="42">Pseudotricula</emphasis>
|
||
</taxonomicName>
|
||
but
|
||
<taxonomicName id="4C024D2F2D2BA225E7F7FA687D70FA3E" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[736,845,1444,1469]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="conica">
|
||
<emphasis id="B976EABE2D2BA225E7F7FA687D70FA3E" box="[736,845,1444,1469]" italics="true" pageId="41" pageNumber="42">P. conica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C024D2F2D2BA225E649FA687A25FA3E" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[862,1048,1444,1469]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="arthurclarkei">
|
||
<emphasis id="B976EABE2D2BA225E649FA687A25FA3E" box="[862,1048,1444,1469]" italics="true" pageId="41" pageNumber="42">P. arthurclarkei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D2BA225E141FA687ACCFA3E" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[1110,1265,1444,1469]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D2BA225E141FA687ACCFA3E" box="[1110,1265,1444,1469]" italics="true" pageId="41" pageNumber="42">P. progenitor</emphasis>
|
||
</taxonomicName>
|
||
also have a long shaft.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |