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<mods:title id="9F00A1D7D2453E0EF35F616799B64028">Three new species of the family Cirolanidae Dana, 1852 (Isopoda, Cymothoida) from the shallow Brazilian bathyal</mods:title>
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<mods:affiliation id="D7C99FD7607AF1493F5E248EC9E1EB6D">Museu de Oceanografia Petrônio Alves Coelho, Universidade Federal de Pernambuco, Laboratório de Carcinologia. Av. Arquitetura, s / n, Cidade Universitária, CEP 50740 - 550, Recife, Pernambuco, Brazil. &amp; ricardopaiva _ 5 @ hotmail. com; https: // orcid. org / 0000 - 0003 - 4881 - 0646</mods:affiliation>
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<mods:affiliation id="16A82832D7C773D5AA2B0B5E5513C71A">Museu de Oceanografia Petrônio Alves Coelho, Universidade Federal de Pernambuco, Laboratório de Carcinologia. Av. Arquitetura, s / n, Cidade Universitária, CEP 50740 - 550, Recife, Pernambuco, Brazil. &amp; jesser. fidelis @ ufpe. br, https: // orcid. org / 0000 - 0001 - 5248 - 2134</mods:affiliation>
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Genus
<taxonomicName id="4C024D0AFFBDF7113CC2FC12FDE1FC48" ID-CoL="7NXBH" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[238,544,968,995]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF7113CC2FC12FE45FC49" bold="true" box="[238,388,968,995]" italics="true" pageId="1" pageNumber="2">Dolicholana</emphasis>
<bibRefCitation id="EF934B78FFBDF7113DA0FC13FDE1FC48" author="Bruce, N. L." box="[396,544,968,995]" pageId="1" pageNumber="2" pagination="1 - 239" refId="ref9720" refString="Bruce, N. L. (1986) Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, 6 (Supplement), 1 - 239. https: // doi. org / 10.3853 / j. 0812 - 7387.6.1986.98" type="journal article" year="1986">Bruce, 1986</bibRefCitation>
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<treatmentCitation id="0AA31098FFBDF7113CBBFBD7FE0FFB8F" author="Bruce, N. L." box="[151,462,1037,1061]" page="122" pageId="1" pageNumber="2" year="1986">
<taxonomicName id="4C024D0AFFBDF7113CBBFBD7FE0FFB8F" ID-CoL="7NXBH" authority="Bruce, 1986: 122" authorityName="Bruce" authorityPageNumber="122" authorityYear="1986" box="[151,462,1037,1061]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF7113CBBFBD7FED7FB8F" box="[151,278,1037,1061]" italics="true" pageId="1" pageNumber="2">Dolicholana</emphasis>
<bibRefCitation id="EF934B78FFBDF7113D30FBD7FE0FFB8F" author="Bruce, N. L." box="[284,462,1037,1061]" pageId="1" pageNumber="2" pagination="1 - 239" refId="ref9720" refString="Bruce, N. L. (1986) Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, 6 (Supplement), 1 - 239. https: // doi. org / 10.3853 / j. 0812 - 7387.6.1986.98" type="journal article" year="1986">Bruce, 1986: 122</bibRefCitation>
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;
<treatmentCitation id="0AA31098FFBDF7113DF4FBD7FD6DFB8F" author="Wetzer, R. &amp; Delaney, P. M. &amp; Brusca, R. C." box="[472,684,1037,1061]" page="2" pageId="1" pageNumber="2" year="1987">
<bibRefCitation id="EF934B78FFBDF7113DF4FBD7FD6DFB8F" author="Wetzer, R. &amp; Delaney, P. M. &amp; Brusca, R. C." box="[472,684,1037,1061]" pageId="1" pageNumber="2" pagination="1 - 10" refId="ref11351" refString="Wetzer, R., Delaney, P. M. &amp; Brusca, R. C. (1987) Politolana wickstenae new species, a new cirolanid isopod from the Gulf of Mexico, and a review of the &quot; Conilera genus group &quot; of Bruce (1986). Contributions in Science, Natural History Museum of Los Angeles County, 392, 1 - 10. https: // doi. org / 10.5962 / p. 226825" type="journal article" year="1987">
Wetzer
<emphasis id="B976EA9BFFBDF7113E0AFBD4FD98FB8F" box="[550,601,1037,1061]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
1987: 2
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(key),
<treatmentCitation id="0AA31098FFBDF7113ED8FBD7FC6BFB8F" author="Keable, S. J." box="[756,938,1037,1061]" page="396" pageId="1" pageNumber="2" year="1999">
<bibRefCitation id="EF934B78FFBDF7113ED8FBD7FC6BFB8F" author="Keable, S. J." box="[756,938,1037,1061]" pageId="1" pageNumber="2" pagination="395 - 414" refId="ref10279" refString="Keable, S. J. (1999) Description of a new species of Dolicholana Bruce, 1986 (Crustacea: Isopoda: Cirolanidae) and a redescription of Dolicholana porcellana (Barnard, 1936) comb. nov. Journal of Natural History, 33, 395 - 414. https: // doi. org / 10.1080 / 002229399300317" type="journal article" year="1999">Keable 1999: 396</bibRefCitation>
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,
<treatmentCitation id="0AA31098FFBDF7113F99FBD7FB58FB8F" author="Bruce, N. L. &amp; Lew Ton, H. M. &amp; Poore, G. C. B." box="[949,1177,1037,1061]" page="145" pageId="1" pageNumber="2" year="2002">
<bibRefCitation id="EF934B78FFBDF7113F99FBD7FB58FB8F" author="Bruce, N. L. &amp; Lew Ton, H. M. &amp; Poore, G. C. B." box="[949,1177,1037,1061]" pageId="1" pageNumber="2" pagination="138 - 157" refId="ref9869" refString="Bruce, N. L., Lew Ton, H. M. &amp; Poore, G. C. B. (2002) Cirolanidae Dana, 1852. In: Poore, G. C. B. (Ed.), Crustacea: Malacostraca: Syncarida and Peracarida: Isopoda, Tanaidacea, Mictacea, Thermosbaenacea, Spelaeogriphacea, Zoological Catalogue of Australia. CSIRO Publishing, Melbourne, pp. 138 - 157." type="book chapter" year="2002">
Bruce
<emphasis id="B976EA9BFFBDF7113FDBFBD4FBEAFB8F" box="[1015,1067,1037,1061]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
2002: 145
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.
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<emphasis id="B976EA9BFFBDF7113CBBFB89FEEAFBC7" bold="true" box="[151,299,1107,1133]" pageId="1" pageNumber="2">
<typeStatus id="54B9882BFFBDF7113CBBFB89FF13FBC7" box="[151,210,1107,1133]" pageId="1" pageNumber="2">Type</typeStatus>
species
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.
<taxonomicName id="4C024D0AFFBDF7113D15FB89FCE6FBC7" authority="H. Milne-Edwards, 1840" authorityName="H. Milne-Edwards" authorityYear="1840" box="[313,807,1107,1134]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B976EA9BFFBDF7113D15FB89FDC7FBC7" box="[313,518,1107,1133]" italics="true" pageId="1" pageNumber="2">Cirolana elongata</emphasis>
H. Milne-Edwards, 1840
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; by original designation.
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<paragraph id="8BBD3689FFBDF7113CEBFBADFB72FAC0" blockId="1.[151,1437,1107,1710]" pageId="1" pageNumber="2">
<emphasis id="B976EA9BFFBDF7113CEBFBADFE28FB3B" bold="true" box="[199,489,1143,1169]" pageId="1" pageNumber="2">Remarks on the genus.</emphasis>
The genus
<taxonomicName id="4C024D0AFFBDF7113E50FBADFC65FB3B" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[636,932,1143,1170]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF7113E50FBADFCC9FB3B" box="[636,776,1143,1169]" italics="true" pageId="1" pageNumber="2">Dolicholana</emphasis>
<bibRefCitation id="EF934B78FFBDF7113F39FBA2FC65FB3B" author="Bruce, N. L." box="[789,932,1143,1170]" pageId="1" pageNumber="2" pagination="1 - 239" refId="ref9720" refString="Bruce, N. L. (1986) Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, 6 (Supplement), 1 - 239. https: // doi. org / 10.3853 / j. 0812 - 7387.6.1986.98" type="journal article" year="1986">Bruce, 1986</bibRefCitation>
</taxonomicName>
includes four species:
<taxonomicName id="4C024D0AFFBDF7113890FBADFE16FB1C" authority="Paiva &amp; Souza-Filho, 2015" authorityName="Paiva &amp; Souza-Filho" authorityYear="2015" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="brucei">
<emphasis id="B976EA9BFFBDF7113890FBADFA5CFB3B" box="[1212,1437,1143,1169]" italics="true" pageId="1" pageNumber="2">Dolicholana brucei</emphasis>
<bibRefCitation id="EF934B78FFBDF7113CBBFB41FE16FB1C" author="Paiva, R. J. C. &amp; Souza-Filho, J. F." box="[151,471,1179,1206]" pageId="1" pageNumber="2" pagination="276 - 288" refId="ref10860" refString="Paiva, R. J. C. &amp; Souza-Filho, J. F. (2015) A new species of Dolicholana Bruce, 1986 (Isopoda, Cymothoidea, Cirolanidae), the first record of the genus from the Atlantic Ocean. Zootaxa, 4039 (2), 276 - 288. https: // doi. org / 10.11646 / zootaxa. 4039.2.4" type="journal article" year="2015">Paiva &amp; Souza-Filho, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C024D0AFFBDF7113DCAFB41FC39FB1F" authority="(Milne Edwards, 1840)" baseAuthorityName="Milne Edwards" baseAuthorityYear="1840" box="[486,1016,1179,1205]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B976EA9BFFBDF7113DCAFB41FD1EFB1F" box="[486,735,1179,1205]" italics="true" pageId="1" pageNumber="2">Dolicholana elongata</emphasis>
(
<bibRefCitation id="EF934B78FFBDF7113EDDFB41FC30FB1F" author="Milne Edwards, H." box="[753,1009,1179,1205]" pageId="1" pageNumber="2" refId="ref10764" refString="Milne Edwards, H. (1840) Histoire naturelle des Crustacees, comprenant l'anatomie, la physiologie et la classification de ces animaux. Librairie encyclopedique de Roret, Paris, 638 pp." type="book" year="1840">Milne Edwards, 1840</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C024D0AFFBDF7113824FB41FA59FB1F" authority="Keable, 1999" authorityName="Keable" authorityYear="1999" box="[1032,1432,1179,1205]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B976EA9BFFBDF7113824FB41FB30FB1F" box="[1032,1265,1179,1205]" italics="true" pageId="1" pageNumber="2">Dolicholana enigma</emphasis>
<bibRefCitation id="EF934B78FFBDF71138D0FB41FA59FB1F" author="Keable, S. J." box="[1276,1432,1179,1205]" pageId="1" pageNumber="2" pagination="395 - 414" refId="ref10279" refString="Keable, S. J. (1999) Description of a new species of Dolicholana Bruce, 1986 (Crustacea: Isopoda: Cirolanidae) and a redescription of Dolicholana porcellana (Barnard, 1936) comb. nov. Journal of Natural History, 33, 395 - 414. https: // doi. org / 10.1080 / 002229399300317" type="journal article" year="1999">Keable, 1999</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C024D0AFFBDF7113CEBFB65FD50FB73" authority="(Barnard, 1936)" baseAuthorityName="Barnard" baseAuthorityYear="1936" box="[199,657,1215,1241]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="porcellana">
<emphasis id="B976EA9BFFBDF7113CEBFB65FE13FB73" box="[199,466,1215,1241]" italics="true" pageId="1" pageNumber="2">Dolicholana porcellana</emphasis>
(
<bibRefCitation id="EF934B78FFBDF7113DCEFB65FD49FB73" author="Barnard, K. H." box="[482,648,1215,1241]" pageId="1" pageNumber="2" pagination="147 - 191" refId="ref9228" refString="Barnard, K. H. (1936) Isopods collected by the R. I. M. S. Investigator. Records of the Indian Museum, 38, 147 - 191. https: // doi. org / 10.26515 / rzsi / v 38 / i 2 / 1936 / 162336" type="journal article" year="1936">Barnard, 1936</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C024D0AFFBDF7113EB0FB65FCE9FB73" authorityName="Bruce" authorityYear="1986" box="[668,808,1215,1241]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF7113EB0FB65FCE9FB73" box="[668,808,1215,1241]" italics="true" pageId="1" pageNumber="2">Dolicholana</emphasis>
</taxonomicName>
resembles both
<taxonomicName id="4C024D0AFFBDF7113FCCFB65FB2EFB73" authority="Bruce, 1981" authorityName="Bruce" authorityYear="1981" box="[992,1263,1215,1242]" class="Malacostraca" family="Cirolanidae" genus="Natatolana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF7113FCCFB65FB9FFB73" box="[992,1118,1215,1241]" italics="true" pageId="1" pageNumber="2">Natatolana</emphasis>
<bibRefCitation id="EF934B78FFBDF7113849FB1AFB2EFB73" author="Bruce, N. L." box="[1125,1263,1215,1242]" pageId="1" pageNumber="2" pagination="945 - 966" refId="ref9650" refString="Bruce, N. L. (1981) Cirolanidae (Crustacea: Isopoda) of Australia: diagnoses of Cirolana Leach, Metacirolana Nierstrasz, Neocirolana Hale, Anopsilana Paulian &amp; Debouteville, and three new genera - Natatolana, Politolana and Cartetolana. Australian Journal of Marine and Freshwater Research, 6, 945 - 966. https: // doi. org / 10.1071 / MF 9810945" type="journal article" year="1981">Bruce, 1981</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D0AFFBDF711390AFB65FEE0FB57" authority="Bruce, 1981" authorityName="Bruce" authorityYear="1981" class="Malacostraca" family="Cirolanidae" genus="Politolana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF711390AFB65FA5DFB73" box="[1318,1436,1215,1241]" italics="true" pageId="1" pageNumber="2">Politolana</emphasis>
<bibRefCitation id="EF934B78FFBDF7113CBBFB3EFEE0FB57" author="Bruce, N. L." box="[151,289,1251,1278]" pageId="1" pageNumber="2" pagination="945 - 966" refId="ref9650" refString="Bruce, N. L. (1981) Cirolanidae (Crustacea: Isopoda) of Australia: diagnoses of Cirolana Leach, Metacirolana Nierstrasz, Neocirolana Hale, Anopsilana Paulian &amp; Debouteville, and three new genera - Natatolana, Politolana and Cartetolana. Australian Journal of Marine and Freshwater Research, 6, 945 - 966. https: // doi. org / 10.1071 / MF 9810945" type="journal article" year="1981">Bruce, 1981</bibRefCitation>
</taxonomicName>
, and in order to distinguish these genera,
<bibRefCitation id="EF934B78FFBDF7113EC1FB3EFC40FB57" author="Bruce, N. L." box="[749,897,1251,1278]" pageId="1" pageNumber="2" pagination="1 - 239" refId="ref9720" refString="Bruce, N. L. (1986) Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, 6 (Supplement), 1 - 239. https: // doi. org / 10.3853 / j. 0812 - 7387.6.1986.98" type="journal article" year="1986">Bruce (1986)</bibRefCitation>
emphasized the projection of the frontal lamina, the morphology of the peduncle of antenna 1, and the absence of plumose setae on the endopod of pleopods
<quantity id="4CFA9B6CFFBDF7113979FADDFA5DFA8B" box="[1365,1436,1287,1314]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="1" pageNumber="2" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">35 in</quantity>
<taxonomicName id="4C024D0AFFBDF7113CBBFAF1FEE2FAEF" authorityName="Bruce" authorityYear="1986" box="[151,291,1323,1349]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF7113CBBFAF1FEE2FAEF" box="[151,291,1323,1349]" italics="true" pageId="1" pageNumber="2">Dolicholana</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF934B78FFBDF7113D1EFAF1FE15FAEF" author="Keable, S. J." box="[306,468,1323,1349]" pageId="1" pageNumber="2" pagination="395 - 414" refId="ref10279" refString="Keable, S. J. (1999) Description of a new species of Dolicholana Bruce, 1986 (Crustacea: Isopoda: Cirolanidae) and a redescription of Dolicholana porcellana (Barnard, 1936) comb. nov. Journal of Natural History, 33, 395 - 414. https: // doi. org / 10.1080 / 002229399300317" type="journal article" year="1999">Keable (1999)</bibRefCitation>
emended the diagnosis of the genus and added a further character, the formation of a 90° angle between the frontal lamina and the clypeus, which is unique in the cirolanid genera.
</paragraph>
<paragraph id="8BBD3689FFBDF7113CEBFAA9FD8EF907" blockId="1.[151,1437,1107,1710]" pageId="1" pageNumber="2">
<bibRefCitation id="EF934B78FFBDF7113CEBFAA9FEABFA27" author="Keable, S. J." box="[199,362,1395,1421]" pageId="1" pageNumber="2" pagination="395 - 414" refId="ref10279" refString="Keable, S. J. (1999) Description of a new species of Dolicholana Bruce, 1986 (Crustacea: Isopoda: Cirolanidae) and a redescription of Dolicholana porcellana (Barnard, 1936) comb. nov. Journal of Natural History, 33, 395 - 414. https: // doi. org / 10.1080 / 002229399300317" type="journal article" year="1999">Keable (1999)</bibRefCitation>
described some additional diagnostic characters of the genus
<taxonomicName id="4C024D0AFFBDF711381FFAA9FB7EFA27" authorityName="Bruce" authorityYear="1986" box="[1075,1215,1395,1421]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF711381FFAA9FB7EFA27" box="[1075,1215,1395,1421]" italics="true" pageId="1" pageNumber="2">Dolicholana</emphasis>
</taxonomicName>
and, in the case of the pleotelson, he stated that the dorsal surface is smooth with an anterodorsal depression, with no tubercles, pits or ridges, the margins are convex, meeting at an acute or rounded apex, with robust setae, and abundant plumose setae restricted to the posterolateral margins, but no marginal teeth-like serrations. In the adult males, the appendix masculina of pleopod 2 arises basally, although in juvenile
<taxonomicName id="4C024D0AFFBDF7113F32F9DEFC51F9B7" authorityName="Keable" authorityYear="1999" box="[798,912,1540,1565]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B976EA9BFFBDF7113F32F9DEFC51F9B7" box="[798,912,1540,1565]" italics="true" pageId="1" pageNumber="2">D. enigma</emphasis>
</taxonomicName>
, it arises medially. In the new species described here, however, the distal margin of the pleotelson has no robust setae, and the adult appendix masculina arises medially (
<figureCitation id="13392A0CFFBDF7113D24F991FEA1F9CF" box="[264,352,1611,1637]" captionStart="FIGURE 6" captionStartId="8.[151,250,1876,1900]" captionTargetBox="[155,1432,529,1848]" captionTargetId="figure-206@8.[151,1436,525,1852]" captionTargetPageId="8" captionText="FIGURE 6. Dolicholana ivanyae spec. nov., Holotype, male, 15 mm (MNRJ 20208). AE = Pleopods 15; F = Uropod. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8182014" httpUri="https://zenodo.org/record/8182014/files/figure.png" pageId="1" pageNumber="2">Fig. 6B</figureCitation>
). Despite these differences, we decided to allocate the new species to the genus
<taxonomicName id="4C024D0AFFBDF71138FBF991FAA2F9CF" authorityName="Bruce" authorityYear="1986" box="[1239,1379,1611,1637]" class="Malacostraca" family="Cirolanidae" genus="Dolicholana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B976EA9BFFBDF71138FBF991FAA2F9CF" box="[1239,1379,1611,1637]" italics="true" pageId="1" pageNumber="2">Dolicholana</emphasis>
</taxonomicName>
until a more ample phylogenetic review of the
<taxonomicName id="4C024D0AFFBDF7113E59F9B5FD39F923" authorityName="Dana" authorityYear="1852" box="[629,760,1647,1673]" class="Malacostraca" family="Cirolanidae" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Cirolanidae</taxonomicName>
can clarify the relationships between this and other related genera, as suggested by
<bibRefCitation id="EF934B78FFBDF7113D8BF949FD8BF907" author="Keable, S. J." box="[423,586,1683,1709]" pageId="1" pageNumber="2" pagination="395 - 414" refId="ref10279" refString="Keable, S. J. (1999) Description of a new species of Dolicholana Bruce, 1986 (Crustacea: Isopoda: Cirolanidae) and a redescription of Dolicholana porcellana (Barnard, 1936) comb. nov. Journal of Natural History, 33, 395 - 414. https: // doi. org / 10.1080 / 002229399300317" type="journal article" year="1999">Keable (1999)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>