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<document id="4AB3D5492DCDC2C83EB91E6406EFCD0C" ID-DOI="10.5281/zenodo.186062" ID-GBIF-Dataset="52e8d2c4-47e1-4886-a141-7f728a5bb382" ID-ISSN="1175-5326" ID-Zenodo-Dep="186062" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460065944461" checkinUser="plazi" docAuthor="Plant, Adrian R." docDate="2009" docId="03AB8789FFC1FFCE5FC77891645EFC42" docLanguage="en" docName="zt02020p050.pdf" docOrigin="Zootaxa 2020" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Achelipoda tumida Plant, 2009, sp. n." docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="FF92FFF1FFC8FFC45F507C516739FFAB" masterDocTitle="Revision of the east Asian genus Achelipoda Yang, Zhang &amp; Zhang, 2007 (Diptera: Empididae: Hemerodromiinae) including designation of a neotype for Achelipoda pictipennis (Bezzi, 1912) and descriptions of six new species" masterLastPageNumber="50" masterPageNumber="37" pageNumber="46" updateTime="1698234646946" updateUser="plazi">
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<mods:title id="80F7B29CC719C166B26CE3DC4FA65494">Revision of the east Asian genus Achelipoda Yang, Zhang &amp; Zhang, 2007 (Diptera: Empididae: Hemerodromiinae) including designation of a neotype for Achelipoda pictipennis (Bezzi, 1912) and descriptions of six new species</mods:title>
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<mods:namePart id="A8DBF7205B5FE9F7DB8A6806B2D1D0F0">Plant, Adrian R.</mods:namePart>
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<mods:date id="C5AF225CC0B327FAFAE13A13C8B1F321">2009</mods:date>
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<treatment id="03AB8789FFC1FFCE5FC77891645EFC42" ID-DOI="http://doi.org/10.5281/zenodo.6214488" ID-GBIF-Taxon="124874416" ID-Zenodo-Dep="6214488" LSID="urn:lsid:plazi:treatment:03AB8789FFC1FFCE5FC77891645EFC42" httpUri="http://treatment.plazi.org/id/03AB8789FFC1FFCE5FC77891645EFC42" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46">
<subSubSection id="C3186514FFC1FFCD5FC7789166DBFB71" box="[151,482,1216,1242]" pageId="9" pageNumber="46" type="nomenclature">
<paragraph id="8BBD369FFFC1FFCD5FC7789166DBFB71" blockId="9.[151,482,1216,1277]" box="[151,482,1216,1242]" pageId="9" pageNumber="46">
<heading id="D0F581F3FFC1FFCD5FC7789166DBFB71" bold="true" box="[151,482,1216,1242]" fontSize="11" level="1" pageId="9" pageNumber="46" reason="1">
<emphasis id="B976EA8DFFC1FFCD5FC7789166DBFB71" bold="true" box="[151,482,1216,1242]" pageId="9" pageNumber="46">
6.
<taxonomicName id="4C024D1CFFC1FFCD5FE4789166A0FB71" ID-CoL="9HZFJ" box="[180,409,1216,1242]" class="Insecta" family="Empididae" genus="Achelipoda" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="tumida" status="sp. nov.">
<emphasis id="B976EA8DFFC1FFCD5FE4789166A0FB71" bold="true" box="[180,409,1216,1242]" italics="true" pageId="9" pageNumber="46">Achelipoda tumida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557F6FFC1FFCD5EF0789166DBFB71" box="[416,482,1216,1242]" pageId="9" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3186514FFC1FFCD5FC778B2660DFB56" box="[151,308,1251,1277]" pageId="9" pageNumber="46" type="description">
<paragraph id="8BBD369FFFC1FFCD5FC778B2660DFB56" blockId="9.[151,482,1216,1277]" box="[151,308,1251,1277]" pageId="9" pageNumber="46">
[
<figureCitation id="13392A1AFFC1FFCD5FCE78B26637FB56" box="[158,270,1251,1277]" captionStart="FIGURES 5 8" captionStartId="9.[151,270,1073,1097]" captionTargetBox="[167,1341,197,1036]" captionTargetId="figure@9.[151,1436,179,1067]" captionTargetPageId="9" captionText="FIGURES 5 8. Genitalia of Achelipoda spp. 5 6. 3 genitalia, lateral view. 5. Achelipoda tumida sp. n. 6. Achelipoda vietnamensis sp. n. 7 8. Achelipoda tumida sp. n. Ƥ genitalia. 7. lateral view. 8. ventral view." httpUri="https://zenodo.org/record/186064/files/figure.png" pageId="9" pageNumber="46">Figs. 5, 7</figureCitation>
9]
</paragraph>
</subSubSection>
<subSubSection id="C3186514FFC1FFCD5FC779616587FA15" pageId="9" pageNumber="46" type="materials_examined">
<paragraph id="8BBD369FFFC1FFCD5FC779616587FA15" blockId="9.[151,1437,1328,2023]" pageId="9" pageNumber="46">
<emphasis id="B976EA8DFFC1FFCD5FC77961666AFAE1" bold="true" box="[151,339,1328,1354]" pageId="9" pageNumber="46">
<typeStatus id="54B9883DFFC1FFCD5FC7796167ECFAE1" box="[151,213,1328,1354]" pageId="9" pageNumber="46">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B9883DFFC1FFCD5E0D796166F6FAE1" box="[349,463,1328,1354]" pageId="9" pageNumber="46" type="holotype">Holotype</typeStatus>
3:
<emphasis id="B976EA8DFFC1FFCD5D50796165BDFAE1" bold="true" box="[512,644,1328,1354]" pageId="9" pageNumber="46">
<collectingCountry id="F315760FFFC1FFCD5D50796165BDFAE1" box="[512,644,1328,1354]" name="Singapore" pageId="9" pageNumber="46">Singapore</collectingCountry>
</emphasis>
, Nee Soon, swamp forest, Malaise trap,
<date id="FFBC105FFFC1FFCD5B2B796163C2FAE1" box="[1147,1275,1328,1354]" pageId="9" pageNumber="46" value="2002-12-04">4.xii.2002</date>
, P. Grootaert [
<collectionCode id="ED13AE5AFFC1FFCD5FCE790667C7FADA" box="[158,254,1367,1393]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="9" pageNumber="46">IRSNB</collectionCode>
].
<typeStatus id="54B9883DFFC1FFCD5E41790666B1FADA" box="[273,392,1367,1393]" pageId="9" pageNumber="46" type="paratype">Paratypes</typeStatus>
: 3Ƥ, same data as
<typeStatus id="54B9883DFFC1FFCD5D36790665E9FADA" box="[614,720,1367,1393]" pageId="9" pageNumber="46" type="holotype">holotype</typeStatus>
,
<date id="FFBC105FFFC1FFCD5D8D7906646EFADA" box="[733,855,1367,1393]" pageId="9" pageNumber="46" value="2002-12-04">4.xii.2002</date>
,
<date id="FFBC105FFFC1FFCD5C34790664E1FADA" box="[868,984,1367,1393]" pageId="9" pageNumber="46" value="2006-03-03">3.iii.2006</date>
,
<date id="FFBC105FFFC1FFCD5CB579066359FADA" box="[997,1120,1367,1393]" pageId="9" pageNumber="46" value="2003-12-03">3.xii.2003</date>
; 9Ƥ, Sime Forest, Malaise trap
<date id="FFBC105FFFC1FFCD5F81792F666AFA33" box="[209,339,1406,1432]" pageId="9" pageNumber="46" value="2006-02-17">17.ii.2006</date>
,
<date id="FFBC105FFFC1FFCD5E31792F66E6FA33" box="[353,479,1406,1432]" pageId="9" pageNumber="46" value="2005-05-19">19.v.2005</date>
,
<date id="FFBC105FFFC1FFCD5EBD792F6562FA33" box="[493,603,1406,1432]" pageId="9" pageNumber="46" value="2005-05-06">6.v.2005</date>
,
<date id="FFBC105FFFC1FFCD5D39792F643DFA33" box="[617,772,1406,1432]" pageId="9" pageNumber="46" value="2005-08-19">19.viii.2005</date>
,
<date id="FFBC105FFFC1FFCD5C41792F64B3FA33" box="[785,906,1406,1432]" pageId="9" pageNumber="46" value="2005-09-07">7.ix.2005</date>
,
<date id="FFBC105FFFC1FFCD5CC9792F6318FA33" box="[921,1057,1406,1432]" pageId="9" pageNumber="46" value="2005-09-16">16.ix.2005</date>
,
<date id="FFBC105FFFC1FFCD5B60792F6380FA33" box="[1072,1209,1406,1432]" pageId="9" pageNumber="46" value="2005-11-18">18.xi.2005</date>
,
<date id="FFBC105FFFC1FFCD5B97792F627FFA33" box="[1223,1350,1406,1432]" pageId="9" pageNumber="46" value="2005-12-03">3.xii.2005</date>
[
<collectionCode id="ED13AE5AFFC1FFCD5A06792F62A1FA33" box="[1366,1432,1406,1432]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="46">ZRC</collectionCode>
,
<collectionCode id="ED13AE5AFFC1FFCD5FC779F567C9FA15" box="[151,240,1444,1470]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="9" pageNumber="46">IRSNB</collectionCode>
,
<collectionCode id="ED13AE5AFFC1FFCD5FAC79F56658FA15" box="[252,353,1444,1470]" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="9" pageNumber="46">NMWC</collectionCode>
].
<typeStatus id="54B9883DFFC1FFCD5E2379F566D9FA15" box="[371,480,1444,1470]" pageId="9" pageNumber="46" type="holotype">Holotype</typeStatus>
deposited in
<collectionCode id="ED13AE5AFFC1FFCD5D2C79F56580FA15" box="[636,697,1444,1470]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="46">ZRC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3186514FFC1FFCD5F95799A6442F9A7" pageId="9" pageNumber="46" type="etymology">
<paragraph id="8BBD369FFFC1FFCD5F95799A6442F9A7" blockId="9.[151,1437,1328,2023]" pageId="9" pageNumber="46">
<emphasis id="B976EA8DFFC1FFCD5F95799A6663FA4E" bold="true" box="[197,346,1483,1509]" pageId="9" pageNumber="46">Etymology.</emphasis>
The specific epithet derives from tumidus (Latin) meaning swollen or inflated in reference to the swollen hind tibia in the female of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3186514FFC1FFCD5F957A4966B6F92B" pageId="9" pageNumber="46" type="diagnosis">
<paragraph id="8BBD369FFFC1FFCD5F957A4966B6F92B" blockId="9.[151,1437,1328,2023]" pageId="9" pageNumber="46">
<emphasis id="B976EA8DFFC1FFCD5F957A49667EF999" bold="true" box="[197,327,1560,1586]" pageId="9" pageNumber="46">Diagnosis.</emphasis>
A yellowish species with wings faintly infuscated but apically clear in female. Female with hind tibia inflated and with a distinct posteroapical spur. Male distinguished from
<taxonomicName id="4C024D1CFFC1FFCD5B207A6E63C6F9F3" box="[1136,1279,1599,1624]" class="Insecta" family="Empididae" genus="Achelipoda" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="kanaklua">
<emphasis id="B976EA8DFFC1FFCD5B207A6E63C6F9F3" box="[1136,1279,1599,1624]" italics="true" pageId="9" pageNumber="46">A. kanaklua</emphasis>
</taxonomicName>
primarily by form of the genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C3186514FFC1FFCE5F957ADD633CFC37" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46" type="description">
<paragraph id="8BBD369FFFC1FFCD5F957ADD6499F95F" blockId="9.[151,1437,1328,2023]" pageId="9" pageNumber="46">
<emphasis id="B976EA8DFFC1FFCD5F957ADD6697F90D" bold="true" box="[197,430,1676,1702]" pageId="9" pageNumber="46">Description. Male:</emphasis>
length
<quantity id="4CFA9B7AFFC1FFCD5D577ADD655CF90D" box="[519,613,1676,1702]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="46" unit="mm" value="2.5">2.5 mm</quantity>
. The
<typeStatus id="54B9883DFFC1FFCD5DF67ADD6434F90D" box="[678,781,1676,1702]" pageId="9" pageNumber="46" type="holotype">holotype</typeStatus>
male appears to have become partially bleached during preservation and colour may be darker in life than in following description. It is also somewhat damaged with antennal stylus broken and most thoracic and head setae detached.
</paragraph>
<paragraph id="8BBD369FFFC1FFCD5F957B50639CF8B1" blockId="9.[151,1437,1328,2023]" box="[197,1189,1792,1818]" pageId="9" pageNumber="46">
<emphasis id="B976EA8DFFC1FFCD5F957B506637F8B1" box="[197,270,1793,1818]" italics="true" pageId="9" pageNumber="46">Head:</emphasis>
dark brown with antenna and mouthparts (including palpus) brownish yellow.
</paragraph>
<paragraph id="8BBD369FFFC1FFCD5F957B76624DF8EA" blockId="9.[151,1437,1328,2023]" box="[197,1396,1831,1861]" pageId="9" pageNumber="46">
<emphasis id="B976EA8DFFC1FFCD5F957B76661DF8EB" box="[197,292,1831,1856]" italics="true" pageId="9" pageNumber="46">Thorax:</emphasis>
orange-brown with only a vaguely darker patch on katepisternum ventrally behind base of C1.
</paragraph>
<paragraph id="8BBD369FFFC1FFCD5F957B00627EF815" blockId="9.[151,1437,1328,2023]" pageId="9" pageNumber="46">
<emphasis id="B976EA8DFFC1FFCD5F957B00663FF8C1" box="[197,262,1873,1898]" italics="true" pageId="9" pageNumber="46">Legs:</emphasis>
yellow, faintly darker on apical tarsomeres. C1 0.9X as long as thorax, hardly narrowed distally. F1 as long as C1, inflated ventrally with usual double row of spines and denticles beneath; basal spines of
<emphasis id="B976EA8DFFC1FFCD5A1E7B2B6250F838" box="[1358,1385,1914,1939]" italics="true" pageId="9" pageNumber="46">av</emphasis>
and
<emphasis id="B976EA8DFFC1FFCD5FC77BF5678AF816" box="[151,179,1956,1981]" italics="true" pageId="9" pageNumber="46">pv</emphasis>
series almost as long as F1 is deep, yellow; basal spine distinct. C3 not inflated, similar in size as C2.
</paragraph>
<paragraph id="8BBD369FFFC1FFCE5F957B9F66ACFE8D" blockId="9.[151,1437,1328,2023]" lastBlockId="10.[151,1437,152,1001]" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46">
<emphasis id="B976EA8DFFC1FFCD5F957B9F6671F84C" box="[197,328,1998,2023]" italics="true" pageId="9" pageNumber="46">Abdomem:</emphasis>
brownish yellow, bare apart from a few small setae on sternite 7 ventrally. Hypandrium + epandrium (
<figureCitation id="13392A1AFFC2FFCE5E787CC96649FF19" box="[296,368,152,178]" captionStart="FIGURES 5 8" captionStartId="9.[151,270,1073,1097]" captionTargetBox="[167,1341,197,1036]" captionTargetId="figure@9.[151,1436,179,1067]" captionTargetPageId="9" captionText="FIGURES 5 8. Genitalia of Achelipoda spp. 5 6. 3 genitalia, lateral view. 5. Achelipoda tumida sp. n. 6. Achelipoda vietnamensis sp. n. 7 8. Achelipoda tumida sp. n. Ƥ genitalia. 7. lateral view. 8. ventral view." httpUri="https://zenodo.org/record/186064/files/figure.png" pageId="10" pageNumber="47">Fig. 5</figureCitation>
) narrowed posteriorly, yellow. Anterior lobe of cercus flattened, brownish, with fan of strong yellowish setae and sharply pointed, black anteriorly direct process (
<figureCitation id="13392A1AFFC2FFCE5B787CEF634BFF73" box="[1064,1138,190,216]" captionStart="FIGURES 5 8" captionStartId="9.[151,270,1073,1097]" captionTargetBox="[167,1341,197,1036]" captionTargetId="figure@9.[151,1436,179,1067]" captionTargetPageId="9" captionText="FIGURES 5 8. Genitalia of Achelipoda spp. 5 6. 3 genitalia, lateral view. 5. Achelipoda tumida sp. n. 6. Achelipoda vietnamensis sp. n. 7 8. Achelipoda tumida sp. n. Ƥ genitalia. 7. lateral view. 8. ventral view." httpUri="https://zenodo.org/record/186064/files/figure.png" pageId="10" pageNumber="47">Fig. 5</figureCitation>
); median lobe yellowish with two distinct setae. Phallus yellow, long, very evenly curved anteriorly. Subepandrial process yellow, bent at right angle distally.
</paragraph>
<paragraph id="8BBD369FFFC2FFCE5F957D63642AFE31" blockId="10.[151,1437,152,1001]" pageId="10" pageNumber="47">
<emphasis id="B976EA8DFFC2FFCE5F957D636636FEE0" box="[197,271,306,331]" italics="true" pageId="10" pageNumber="47">Wing:</emphasis>
veins yellowish brown, membrane faintly tinged yellowish, without banding pattern and not obviously clear apically. Wings with crossveins closing cells bm and br almost strictly aligned, cell br equal to in length to cell bm; vein CuA2 reaching wing margin.
</paragraph>
<paragraph id="8BBD369FFFC2FFCE5F957DF86443FE68" blockId="10.[151,1437,152,1001]" box="[197,890,425,451]" pageId="10" pageNumber="47">
<emphasis id="B976EA8DFFC2FFCE5F957DF86686FE68" bold="true" box="[197,447,425,451]" pageId="10" pageNumber="47">Description. Female</emphasis>
: similar to male, differing as follows.
</paragraph>
<paragraph id="8BBD369FFFC2FFCE5F957D816646FD9C" blockId="10.[151,1437,152,1001]" pageId="10" pageNumber="47">
<emphasis id="B976EA8DFFC2FFCE5F957D816628FE42" box="[197,273,464,489]" italics="true" pageId="10" pageNumber="47">Head:</emphasis>
black with antenna and mouthparts brownish. Ocellar, vertical and postocular setae rather well developed, blackish; lower occipital pile pale and fine. Antenna with postpedicel 33.5X as long as wide, stylus 1.5X as long.
</paragraph>
<paragraph id="8BBD369FFFC2FFCE5F957E156301FD2C" blockId="10.[151,1437,152,1001]" pageId="10" pageNumber="47">
<emphasis id="B976EA8DFFC2FFCE5F957E15661DFDF6" box="[197,292,580,605]" italics="true" pageId="10" pageNumber="47">Thorax:</emphasis>
orange-brown; dark patch behind C1 darker than in male but not strongly defined; dorsocentrals rather stronger than in
<taxonomicName id="4C024D1CFFC2FFCE5EF27E3F657AFD2C" box="[418,579,622,647]" class="Insecta" family="Empididae" genus="Achelipoda" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B976EA8DFFC2FFCE5EF27E3F657AFD2C" box="[418,579,622,647]" italics="true" pageId="10" pageNumber="47">A. pictipennis</emphasis>
</taxonomicName>
but
<emphasis id="B976EA8DFFC2FFCE5D277E3F65A8FD2C" box="[631,657,622,647]" italics="true" pageId="10" pageNumber="47">sa</emphasis>
clearly strongest seta on the thorax.
</paragraph>
<paragraph id="8BBD369FFFC2FFCE5F957EC56332FD55" blockId="10.[151,1437,152,1001]" pageId="10" pageNumber="47">
<emphasis id="B976EA8DFFC2FFCE5F957EC5663EFD06" box="[197,263,660,685]" italics="true" pageId="10" pageNumber="47">Legs:</emphasis>
Femoral formula approximately
<date id="FFBC105FFFC2FFCE5DC57EC565D6FD05" box="[661,751,660,686]" pageId="10" pageNumber="47" value="1918-05-21">5/21/18</date>
/5 +1 with basal spine and 23 spines of
<emphasis id="B976EA8DFFC2FFCE5B8C7EC563C1FD06" box="[1244,1272,660,685]" italics="true" pageId="10" pageNumber="47">av</emphasis>
and
<emphasis id="B976EA8DFFC2FFCE5A647EC56269FD06" box="[1332,1360,660,685]" italics="true" pageId="10" pageNumber="47">pv</emphasis>
series basally as long as limb is deep. T3 as narrow as T2 basally but considerably inflated on distal 0.5 where distinctly wider than T2 (Fig. 9), bearing a distinct black posteroapical spur.
</paragraph>
<paragraph id="8BBD369FFFC2FFCE5F957F5F6604FCDE" blockId="10.[151,1437,152,1001]" pageId="10" pageNumber="47">
<emphasis id="B976EA8DFFC2FFCE5F957F5F6604FC8C" box="[197,317,782,807]" italics="true" pageId="10" pageNumber="47">Abdomen:</emphasis>
almost bare, pale brown with terminal segments bearing a few longer hairs laterally and cercus darker still with longer hairs. Sternite 8 viewed from below with conspicuous U-shaped darker brown sclerotization.
</paragraph>
<paragraph id="8BBD369FFFC2FFCE5F957FD3633CFC37" blockId="10.[151,1437,152,1001]" box="[197,1029,898,924]" pageId="10" pageNumber="47">
<emphasis id="B976EA8DFFC2FFCE5F957FD36633FC30" box="[197,266,898,923]" italics="true" pageId="10" pageNumber="47">Wing:</emphasis>
similar to male but membrane indistinctly clear at extreme apex.
</paragraph>
</subSubSection>
<subSubSection id="C3186514FFC2FFCE5F957FF9645EFC42" pageId="10" pageNumber="47" type="discussion">
<paragraph id="8BBD369FFFC2FFCE5F957FF9645EFC42" blockId="10.[151,1437,152,1001]" pageId="10" pageNumber="47">
<emphasis id="B976EA8DFFC2FFCE5F957FF96671FC69" bold="true" box="[197,328,936,962]" pageId="10" pageNumber="47">Comment.</emphasis>
<taxonomicName id="4C024D1CFFC2FFCE5E1E7FF86514FC69" box="[334,557,937,962]" class="Insecta" family="Empididae" genus="Achelipoda" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="B976EA8DFFC2FFCE5E1E7FF86514FC69" box="[334,557,937,962]" italics="true" pageId="10" pageNumber="47">Achelipoda tumida</emphasis>
</taxonomicName>
is recorded only from moist forest biotopes in
<collectingCountry id="F315760FFFC2FFCE5B1D7FF863F1FC68" box="[1101,1224,937,963]" name="Singapore" pageId="10" pageNumber="47">Singapore</collectingCountry>
. Adults have been collected between February and September and in December.
</paragraph>
</subSubSection>
</treatment>
</document>