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<document id="431F21C0E01BE88CBC8FF6D032F1B6B5" ID-CLB-Dataset="196616" ID-DOI="10.11646/zootaxa.5209.1.7" ID-GBIF-Dataset="8e12b2ee-eaaa-4bac-b2d9-757ac0e152fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="7322612" ID-ZooBank="A934D1CF-61B6-4B90-A405-E39A7551A321" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668502024656" checkinUser="plazi" docAuthor="Ng, Ngan Kee, Trivedi, Jigneshkumar &amp; Bhat, Mithila" docDate="2022" docId="03A50E3735699D7A64EEFF381EB9F986" docLanguage="en" docName="zootaxa.5209.1.7.pdf" docOrigin="Zootaxa 5209 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chhapgarus Ng &amp; Trivedi &amp; Bhat 2022, n. gen." docType="treatment" docVersion="9" lastPageNumber="130" masterDocId="FF9C764F356A9D796479FFAF1F50FF85" masterDocTitle="Redescription of Pseudograpsus intermedius Chhapgar, 1955 (Decapoda, Brachyura, Varunidae) from India" masterLastPageNumber="138" masterPageNumber="127" pageNumber="130" updateTime="1699411558422" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="22EA5E3983650925CE123E3B7F97D38B">Redescription of Pseudograpsus intermedius Chhapgar, 1955 (Decapoda, Brachyura, Varunidae) from India</mods:title>
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<mods:namePart id="AE985EEB09696860E069742B55E0C856">Ng, Ngan Kee</mods:namePart>
<mods:nameIdentifier id="5FDD90715850216D837D18E31DF95149" type="ORCID">0000-0003-1022-1600</mods:nameIdentifier>
<mods:affiliation id="0E5F3662F313716622D56E0959F9CC67">Department of Biological Sciences, Faculty of Science, National University of Singapore, 14, Science Drive 4, Singapore 117377, Republic of Singapore. ngankee @ nus. edu. sg; https: // orcid. org / 0000 - 0003 - 1022 - 1600</mods:affiliation>
<mods:nameIdentifier id="039AA690BBB836673A1D4D4C8F3322A8" type="email">ngankee@nus.edu.sg</mods:nameIdentifier>
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<mods:namePart id="830033DBE386C9E009E143CE6234A0DF">Trivedi, Jigneshkumar</mods:namePart>
<mods:affiliation id="F2C84D23346E2175149D5D94871CE239">Department of Life Sciences, Hemchandracharya North Gujarat University, Patan, Gujarat, India.</mods:affiliation>
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<mods:namePart id="02ABAAE46851B679F3989736DC957039">Bhat, Mithila</mods:namePart>
<mods:affiliation id="E7CDC8C20258B105036C899E24B407E0">School of Earth, Ocean and Atmospheric Sciences, Goa University, Taleigao, Goa, India.</mods:affiliation>
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<mods:title id="E8A39A4F5A4FCA21EADD49B3EE919236">Zootaxa</mods:title>
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<mods:date id="0E382CAA10ACFEED93A347B513E5C74C">2022</mods:date>
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<mods:number id="F360C468CDDAEA4257738BEDCFBBD9F0">2022-11-15</mods:number>
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<treatment id="03A50E3735699D7A64EEFF381EB9F986" ID-DOI="http://doi.org/10.5281/zenodo.7330391" ID-GBIF-Taxon="205139777" ID-Zenodo-Dep="7330391" LSID="urn:lsid:plazi:treatment:03A50E3735699D7A64EEFF381EB9F986" httpUri="http://treatment.plazi.org/id/03A50E3735699D7A64EEFF381EB9F986" lastPageNumber="130" pageId="3" pageNumber="130">
<subSubSection id="C316ECAA35699D7A64EEFF381ED6FF34" box="[151,390,151,178]" pageId="3" pageNumber="130" type="nomenclature">
<paragraph id="8BB3BF2135699D7A64EEFF381ED6FF34" blockId="3.[151,390,151,178]" box="[151,390,151,178]" pageId="3" pageNumber="130">
<heading id="D0FB084D35699D7A64EEFF381ED6FF34" bold="true" box="[151,390,151,178]" fontSize="11" level="1" pageId="3" pageNumber="130" reason="1">
<emphasis id="B978633335699D7A64EEFF381ED6FF34" bold="true" box="[151,390,151,178]" italics="true" pageId="3" pageNumber="130">
<taxonomicName id="4C0CC4A235699D7A64EEFF381E7DFF37" ID-CoL="6badc1d4-70ef-42c4-a9d5-eb73161a1ab6" authority="Ng &amp; Trivedi &amp; Bhat, 2022" authorityName="Ng &amp; Trivedi &amp; Bhat" authorityYear="2022" box="[151,301,151,178]" class="Malacostraca" family="Varunidae" genus="Chhapgarus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus" status="gen. nov.">Chhapgarus</taxonomicName>
<taxonomicNameLabel id="A24BDE4835699D7A654CFF381ED6FF34" box="[309,390,151,177]" pageId="3" pageNumber="130" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316ECAA35699D7A64EEFF761B6AFF77" box="[151,1082,216,243]" pageId="3" pageNumber="130" type="type_taxon">
<paragraph id="8BB3BF2135699D7A64EEFF761B6AFF77" blockId="3.[151,1437,216,1539]" box="[151,1082,216,243]" pageId="3" pageNumber="130">
<emphasis id="B978633335699D7A64EEFF761E62FF77" bold="true" box="[151,306,216,243]" pageId="3" pageNumber="130">
<typeStatus id="54B7018335699D7A64EEFF761F82FF76" box="[151,210,217,243]" pageId="3" pageNumber="130">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C0CC4A235699D7A6540FF771C7BFF76" ID-CoL="4NXPD" authority="Chhapgar, 1955" authorityName="Chhapgar" authorityYear="1955" box="[313,811,216,243]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B978633335699D7A6540FF771D3FFF77" box="[313,623,216,242]" italics="true" pageId="3" pageNumber="130">Pseudograpsus intermedius</emphasis>
<bibRefCitation id="EF9DC2D035699D7A660EFF771C7BFF76" author="Chhapgar, B. F." box="[631,811,216,243]" pageId="3" pageNumber="130" pagination="251 - 259" refId="ref6039" refString="Chhapgar, B. F. (1955) On two new species and a new variety of crabs (Decapoda: Brachyura) from Bombay State. Records of Indian Museum, 53, 251 - 259." type="journal article" year="1955">Chhapgar, 1955</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C316ECAA35699D7A64BEFF531C3DFDB3" pageId="3" pageNumber="130" type="description">
<paragraph id="8BB3BF2135699D7A64BEFF531C3DFDB3" blockId="3.[151,1437,216,1539]" pageId="3" pageNumber="130">
<emphasis id="B978633335699D7A64BEFF531E09FE93" bold="true" box="[199,345,252,278]" pageId="3" pageNumber="130">Description.</emphasis>
Carapace squarish, dorsal surface covered with short, brown setae; regions well defined, convex; frontal margin slightly convex, straight; anterolateral margin subcristate with three teeth including first orbital tooth; posterolateral margins not sharply demarcated from anterolateral margin, concave at branchial region, distinctly subparallel; orbits small, eyes completely filling orbit. Third maxillipeds short, stout; small, distinct rhomboidal gape when closed, palp short, with short setae; exopod with short flagellum. Epistome broad, flat, posterior margin entire. Inner surface of chela with granules, setose. Ambulatory legs long, slender with short setae, stout dactyli. Lateral margins of thoracic sternites 4–5 finely granulated; anterior sternal plates without any medial grooves. Male pleon narrowly triangular, with 7 free segments (6 somites plus telson). Female pleon ovate, very broad. G1 stout, gently curving outwards, terminal lobe elongate, slightly curved.
</paragraph>
</subSubSection>
<subSubSection id="C316ECAA35699D7A64BEFDEE1BC3FDFB" pageId="3" pageNumber="130" type="etymology">
<paragraph id="8BB3BF2135699D7A64BEFDEE1BC3FDFB" blockId="3.[151,1437,216,1539]" pageId="3" pageNumber="130">
<emphasis id="B978633335699D7A64BEFDEE1E18FDDE" bold="true" box="[199,328,577,603]" pageId="3" pageNumber="130">Etymology</emphasis>
. The new genus is named in honor of Dr. Boman Framji Chhapgar for his valuable contributions to the study of crustacean fauna of
<collectingRegion id="49C871C335699D7A667AFDCB1D07FDFB" box="[515,599,612,638]" country="India" name="Gujarat" pageId="3" pageNumber="130">Gujarat</collectingRegion>
and
<collectingRegion id="49C871C335699D7A66F4FDCB1C49FDFB" box="[653,793,612,638]" country="India" name="Maharashtra" pageId="3" pageNumber="130">Maharashtra</collectingRegion>
States,
<collectingCountry id="F31BFFB135699D7A6708FDCB1CFEFDFB" box="[881,942,612,638]" name="India" pageId="3" pageNumber="130">India</collectingCountry>
. Gender: masculine.
</paragraph>
</subSubSection>
<subSubSection id="C316ECAA35699D7A64BEFD261EB9F986" pageId="3" pageNumber="130" type="discussion">
<paragraph id="8BB3BF2135699D7A64BEFD261C1CFC63" blockId="3.[151,1437,216,1539]" pageId="3" pageNumber="130">
<emphasis id="B978633335699D7A64BEFD261E64FD26" bold="true" box="[199,308,649,675]" pageId="3" pageNumber="130">Remarks</emphasis>
. The description of the genus
<taxonomicName id="4C0CC4A235699D7A66F1FD271C63FD27" authorityName="H. Milne Edwards" authorityYear="1837" box="[648,819,648,674]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A66F1FD271C63FD27" box="[648,819,648,674]" italics="true" pageId="3" pageNumber="130">Pseudograpsus</emphasis>
</taxonomicName>
provided by H. Milne Edwards (1837) is brief that has been updated by
<bibRefCitation id="EF9DC2D035699D7A652CFD021D5AFD42" author="Ng, N. K. &amp; Jeng, M. S. &amp; Ng, P. K. L." box="[341,522,684,711]" pageId="3" pageNumber="130" pagination="759 - 776" refId="ref6387" refString="Ng, N. K., Jeng, M. S. &amp; Ng, P. K. L. (2002) On the taxonomy of Pseudograpsus setosus (Fabricius, 1778) (Decapoda, Brachyura, Grapsidae). Crustaceana, 75 (6), 759 - 776. https: // doi. org / 10.1163 / 156854002760289746" type="journal article" year="2002">
Ng
<emphasis id="B978633335699D7A6507FD021EFFFD43" box="[382,431,684,710]" italics="true" pageId="3" pageNumber="130">et al</emphasis>
. (2002)
</bibRefCitation>
. In his description, H. Milne Edwards (1837: 81) named two species as belonging to the genus,
<taxonomicName id="4C0CC4A235699D7A6541FD7F1EB3FD6F" authorityName="H. Milne Edwards" authorityYear="1837" box="[312,483,720,746]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A6541FD7F1EB3FD6F" box="[312,483,720,746]" italics="true" pageId="3" pageNumber="130">Pseudograpsus</emphasis>
</taxonomicName>
viz.
<taxonomicName id="4C0CC4A235699D7A665BFD7E1CD3FD6E" authority="(Latreille, 1817)" baseAuthorityName="Latreille" baseAuthorityYear="1817" box="[546,899,720,747]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="pencilliger">
<emphasis id="B978633335699D7A665BFD7E1DEAFD6F" box="[546,698,720,746]" italics="true" pageId="3" pageNumber="130">P. pencilliger</emphasis>
(Latreille, 1817)
</taxonomicName>
, and
<taxonomicName id="4C0CC4A235699D7A67BFFD7E1A3BFD6E" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[966,1387,720,747]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B978633335699D7A67BFFD7E1B6EFD6F" box="[966,1086,720,746]" italics="true" pageId="3" pageNumber="130">P. pallipes</emphasis>
H. Milne Edwards, 1837
</taxonomicName>
. No
<typeStatus id="54B7018335699D7A64EEFD5A1F97FC8A" box="[151,199,757,783]" pageId="3" pageNumber="130">type</typeStatus>
species was designated. Holthuis (1977) was the first to validly select
<taxonomicName id="4C0CC4A235699D7A6795FD5A1A29FC8A" authority="Latreille, 1817" authorityName="Latreille" authorityYear="1817" box="[1004,1401,756,783]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="penicilliger">
<emphasis id="B978633335699D7A6795FD5A1B98FC8B" box="[1004,1224,756,783]" italics="true" pageId="3" pageNumber="130">Cancer penicilliger</emphasis>
Latreille, 1817
</taxonomicName>
, as the
<typeStatus id="54B7018335699D7A64BBFCB61FA2FCB6" box="[194,242,793,819]" pageId="3" pageNumber="130">type</typeStatus>
species (see also Ng &amp; Nakasone 1993: 2).
<taxonomicName id="4C0CC4A235699D7A668BFCB61C9EFCB7" authorityName="Latreille" authorityYear="1817" box="[754,974,792,819]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="penicilliger">
<emphasis id="B978633335699D7A668BFCB61C9EFCB7" box="[754,974,792,819]" italics="true" pageId="3" pageNumber="130">Cancer penicilliger</emphasis>
</taxonomicName>
was later synonymized under the senior name,
<taxonomicName id="4C0CC4A235699D7A649CFC931EBBFCD3" baseAuthorityName="Fabricius" baseAuthorityYear="1778" box="[229,491,828,854]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="B978633335699D7A649CFC931EBBFCD3" box="[229,491,828,854]" italics="true" pageId="3" pageNumber="130">Pseudograpsus setosus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9DC2D035699D7A6655FC921D8AFCD2" author="Ng, N. K. &amp; Jeng, M. S. &amp; Ng, P. K. L." box="[556,730,828,855]" pageId="3" pageNumber="130" pagination="759 - 776" refId="ref6387" refString="Ng, N. K., Jeng, M. S. &amp; Ng, P. K. L. (2002) On the taxonomy of Pseudograpsus setosus (Fabricius, 1778) (Decapoda, Brachyura, Grapsidae). Crustaceana, 75 (6), 759 - 776. https: // doi. org / 10.1163 / 156854002760289746" type="journal article" year="2002">
Ng
<emphasis id="B978633335699D7A6621FC921DC4FCD3" box="[600,660,828,854]" italics="true" pageId="3" pageNumber="130">et al.</emphasis>
2002
</bibRefCitation>
). The genus
<taxonomicName id="4C0CC4A235699D7A6709FC931B4BFCD3" authorityName="H. Milne Edwards" authorityYear="1837" box="[880,1051,828,854]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A6709FC931B4BFCD3" box="[880,1051,828,854]" italics="true" pageId="3" pageNumber="130">Pseudograpsus</emphasis>
</taxonomicName>
has a long history of association with
<taxonomicName id="4C0CC4A235699D7A64A8FCCE1E36FCFF" authorityName="Dana" authorityYear="1851" box="[209,358,865,890]" class="Malacostraca" family="Varunidae" genus="Hemigrapsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A64A8FCCE1E36FCFF" box="[209,358,865,890]" italics="true" pageId="3" pageNumber="130">Hemigrapsus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0CC4A235699D7A65D9FCCF1D7EFCFF" authorityName="De Haan, 1833 [in De Haan, 1833-" authorityYear="1850" box="[416,558,864,890]" class="Malacostraca" family="Varunidae" genus="Brachynotus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A65D9FCCF1D7EFCFF" box="[416,558,864,890]" italics="true" pageId="3" pageNumber="130">Brachynotus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9DC2D035699D7A6612FCCF1DA0FCFE" author="Tesch, J. J." box="[619,752,864,891]" pageId="3" pageNumber="130" pagination="1 - 148" refId="ref6706" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. I. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga Expedition Monographie, 39 (c 2), 1 - 148. https: // doi. org / 10.5962 / bhl. title. 10267" type="journal article" year="1918">Tesch 1918</bibRefCitation>
; McLay &amp; Schubart 2004;
<bibRefCitation id="EF9DC2D035699D7A6050FCCE1B82FCFE" author="Ng, N. K. &amp; Jeng, M. S. &amp; Ng, P. K. L." box="[1065,1234,864,891]" pageId="3" pageNumber="130" pagination="759 - 776" refId="ref6387" refString="Ng, N. K., Jeng, M. S. &amp; Ng, P. K. L. (2002) On the taxonomy of Pseudograpsus setosus (Fabricius, 1778) (Decapoda, Brachyura, Grapsidae). Crustaceana, 75 (6), 759 - 776. https: // doi. org / 10.1163 / 156854002760289746" type="journal article" year="2002">
Ng
<emphasis id="B978633335699D7A602AFCCE1BDDFCFF" box="[1107,1165,864,890]" italics="true" pageId="3" pageNumber="130">et al.</emphasis>
2002
</bibRefCitation>
). This association is now being studied in-depth (in preparation). Balss (1934) synonymized
<taxonomicName id="4C0CC4A235699D7A67A2FC2A1A65FC1A" authority="Kossmann, 1894" authorityName="Kossmann" authorityYear="1894" box="[987,1333,900,927]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B978633335699D7A67A2FC2A1B3FFC1B" box="[987,1135,900,926]" italics="true" pageId="3" pageNumber="130">P. erythraeus</emphasis>
Kossmann, 1894
</taxonomicName>
, under
<taxonomicName id="4C0CC4A235699D7A61F1FC2A1E54FC47" authorityName="A. Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B978633335699D7A61F1FC2A1E54FC47" italics="true" pageId="3" pageNumber="130">P. elongatus</emphasis>
</taxonomicName>
. Ng &amp; Nakasone (1993) transferred
<taxonomicName id="4C0CC4A235699D7A66E2FC061C56FC47" authorityName="Tesch" authorityYear="1918" box="[667,774,936,962]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="laniger">
<emphasis id="B978633335699D7A66E2FC061C56FC47" box="[667,774,936,962]" italics="true" pageId="3" pageNumber="130">P. laniger</emphasis>
</taxonomicName>
to their new genus,
<taxonomicName id="4C0CC4A235699D7A679FFC061A30FC46" authority="Ng &amp; Nakasone, 1993" authorityName="Ng &amp; Nakasone" authorityYear="1993" box="[998,1376,936,963]" class="Malacostraca" family="Varunidae" genus="Scutumara" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A679FFC061B0FFC46" box="[998,1119,937,963]" italics="true" pageId="3" pageNumber="130">Scutumara</emphasis>
Ng &amp; Nakasone, 1993
</taxonomicName>
, with a comment that the genus
<taxonomicName id="4C0CC4A235699D7A65C5FC631D37FC63" authorityName="H. Milne Edwards" authorityYear="1837" box="[444,615,972,998]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A65C5FC631D37FC63" box="[444,615,972,998]" italics="true" pageId="3" pageNumber="130">Pseudograpsus</emphasis>
</taxonomicName>
needs to be revised.
</paragraph>
<paragraph id="8BB3BF2135699D7A64BEFC5F1FFCFACA" blockId="3.[151,1437,216,1539]" pageId="3" pageNumber="130">
The genus,
<taxonomicName id="4C0CC4A235699D7A653FFC5F1EA1FB8F" authorityName="H. Milne Edwards" authorityYear="1837" box="[326,497,1008,1034]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A653FFC5F1EA1FB8F" box="[326,497,1008,1034]" italics="true" pageId="3" pageNumber="130">Pseudograpsus</emphasis>
</taxonomicName>
, currently comprises of five species viz.
<taxonomicName id="4C0CC4A235699D7A67CDFC5E1BB4FB8E" authority="(Fabricius, 1778)" baseAuthorityName="Fabricius" baseAuthorityYear="1778" box="[948,1252,1008,1035]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="B978633335699D7A67CDFC5E1B4FFB8F" box="[948,1055,1009,1034]" italics="true" pageId="3" pageNumber="130">P. setosus</emphasis>
(Fabricius, 1778)
</taxonomicName>
(=
<taxonomicName id="4C0CC4A235699D7A6170FC5E1E05FBAA" authority="(Latreille, 1817)" baseAuthorityName="Latreille" baseAuthorityYear="1817" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="pencilliger">
<emphasis id="B978633335699D7A6170FC5E1ACCFB8F" box="[1289,1436,1008,1034]" italics="true" pageId="3" pageNumber="130">P. pencilliger</emphasis>
(Latreille, 1817)
</taxonomicName>
,
<typeStatus id="54B7018335699D7A651CFBBA1EC5FBAA" box="[357,405,1045,1071]" pageId="3" pageNumber="130">type</typeStatus>
species);
<taxonomicName id="4C0CC4A235699D7A6674FBBA1CFAFBAB" authority="A. Milne-Edwards, 1868" authorityName="A. Milne-Edwards" authorityYear="1868" box="[525,938,1044,1071]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B978633335699D7A6674FBBA1DD1FBAB" box="[525,641,1045,1070]" italics="true" pageId="3" pageNumber="130">P. crassus</emphasis>
A. Milne-Edwards, 1868
</taxonomicName>
,
<taxonomicName id="4C0CC4A235699D7A67C5FBBA1B8AFBAA" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[956,1242,1044,1071]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="albus">
<emphasis id="B978633335699D7A67C5FBBA1B49FBAB" box="[956,1049,1044,1070]" italics="true" pageId="3" pageNumber="130">P. albus</emphasis>
Stimpson, 1858
</taxonomicName>
,
<taxonomicName id="4C0CC4A235699D7A6090FBBA1EDEFBD6" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B978633335699D7A6090FBBA1A26FBAB" box="[1257,1398,1044,1070]" italics="true" pageId="3" pageNumber="130">P. elongatus</emphasis>
A. Milne-Edwards, 1873
</taxonomicName>
, and
<taxonomicName id="4C0CC4A235699D7A65A9FB961C63FBD6" authority="Chhapgar, 1955" authorityName="Chhapgar" authorityYear="1955" box="[464,819,1080,1107]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B978633335699D7A65A9FB961D22FBD7" box="[464,626,1080,1106]" italics="true" pageId="3" pageNumber="130">P. intermedius</emphasis>
<bibRefCitation id="EF9DC2D035699D7A6605FB971C63FBD6" author="Chhapgar, B. F." box="[636,819,1080,1107]" pageId="3" pageNumber="130" pagination="251 - 259" refId="ref6039" refString="Chhapgar, B. F. (1955) On two new species and a new variety of crabs (Decapoda: Brachyura) from Bombay State. Records of Indian Museum, 53, 251 - 259." type="journal article" year="1955">Chhapgar, 1955</bibRefCitation>
</taxonomicName>
. Out of these five species,
<taxonomicName id="4C0CC4A235699D7A6017FB961A40FBD7" authorityName="Chhapgar" authorityYear="1955" box="[1134,1296,1080,1106]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B978633335699D7A6017FB961A40FBD7" box="[1134,1296,1080,1106]" italics="true" pageId="3" pageNumber="130">P. intermedius</emphasis>
</taxonomicName>
varies from other species of
<taxonomicName id="4C0CC4A235699D7A6537FBF31EA9FBF3" authorityName="H. Milne Edwards" authorityYear="1837" box="[334,505,1116,1142]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B978633335699D7A6537FBF31EA9FBF3" box="[334,505,1116,1142]" italics="true" pageId="3" pageNumber="130">Pseudograpsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BDE4835699D7A6679FBF21D74FBF2" box="[512,548,1117,1143]" pageId="3" pageNumber="130" sensu="stricto">s.s.</taxonomicNameLabel>
in having setae on the carapace, chela having soft setae on outer surface, broad and stout ambulatory legs and differences in morphology of reproductive structures, hence
<taxonomicName id="4C0CC4A235699D7A60CFFB2F1A10FB1F" authority="Ng &amp; Trivedi &amp; Bhat, 2022" authorityName="Ng &amp; Trivedi &amp; Bhat" authorityYear="2022" box="[1206,1344,1152,1178]" class="Malacostraca" family="Varunidae" genus="Chhapgarus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B978633335699D7A60CFFB2F1A10FB1F" box="[1206,1344,1152,1178]" italics="true" pageId="3" pageNumber="130">Chhapgarus</emphasis>
</taxonomicName>
<emphasis id="B978633335699D7A6132FB2E1ACCFB1E" bold="true" box="[1355,1436,1153,1179]" pageId="3" pageNumber="130">
<taxonomicNameLabel id="A24BDE4835699D7A6132FB2E1ACCFB1E" box="[1355,1436,1153,1179]" pageId="3" pageNumber="130" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
is established for
<taxonomicName id="4C0CC4A235699D7A651AFB0B1C0DFB3A" authority="Chhapgar, 1955" authorityName="Chhapgar" authorityYear="1955" box="[355,861,1188,1215]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B978633335699D7A651AFB0B1DCCFB3B" box="[355,668,1188,1214]" italics="true" pageId="3" pageNumber="130">Pseudograpsus intermedius</emphasis>
<bibRefCitation id="EF9DC2D035699D7A66DFFB0B1C0DFB3A" author="Chhapgar, B. F." box="[678,861,1188,1215]" pageId="3" pageNumber="130" pagination="251 - 259" refId="ref6039" refString="Chhapgar, B. F. (1955) On two new species and a new variety of crabs (Decapoda: Brachyura) from Bombay State. Records of Indian Museum, 53, 251 - 259." type="journal article" year="1955">Chhapgar, 1955</bibRefCitation>
</taxonomicName>
. Interestingly,
<taxonomicName id="4C0CC4A235699D7A607FFB0A1BFEFB3B" authorityName="Ng &amp; Trivedi &amp; Bhat" authorityYear="2022" box="[1030,1198,1188,1215]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="intermedius" status="comb. nov.">
<emphasis id="B978633335699D7A607FFB0A1BFEFB3B" box="[1030,1198,1188,1215]" italics="true" pageId="3" pageNumber="130">C. intermedius</emphasis>
</taxonomicName>
<emphasis id="B978633335699D7A60CEFB0A1A6AFB3A" bold="true" box="[1207,1338,1189,1215]" pageId="3" pageNumber="130">
<taxonomicNameLabel id="A24BDE4835699D7A60CEFB0A1A6AFB3A" box="[1207,1338,1189,1215]" pageId="3" pageNumber="130" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
inhabits the mid-intertidal zone in mangrove habitats (
<bibRefCitation id="EF9DC2D035699D7A66DDFB671C07FB66" author="Chhapgar, B. F." box="[676,855,1224,1251]" pageId="3" pageNumber="130" pagination="251 - 259" refId="ref6039" refString="Chhapgar, B. F. (1955) On two new species and a new variety of crabs (Decapoda: Brachyura) from Bombay State. Records of Indian Museum, 53, 251 - 259." type="journal article" year="1955">Chhapgar 1955</bibRefCitation>
), while species of
<taxonomicName id="4C0CC4A235699D7A6056FB671B8AFB67" authorityName="H. Milne Edwards" authorityYear="1837" box="[1071,1242,1224,1250]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B978633335699D7A6056FB671B8AFB67" box="[1071,1242,1224,1250]" italics="true" pageId="3" pageNumber="130">Pseudograpsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BDE4835699D7A609DFB661A57FB66" box="[1252,1287,1225,1251]" pageId="3" pageNumber="130" sensu="stricto">s.s.</taxonomicNameLabel>
are found in freshwater streams and/or under pebbles/coral sand habitats along beaches (
<bibRefCitation id="EF9DC2D035699D7A6079FB431BD8FA82" author="Tesch, J. J." box="[1024,1160,1260,1287]" pageId="3" pageNumber="130" pagination="1 - 148" refId="ref6706" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. I. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga Expedition Monographie, 39 (c 2), 1 - 148. https: // doi. org / 10.5962 / bhl. title. 10267" type="journal article" year="1918">Tesch 1918</bibRefCitation>
;
<bibRefCitation id="EF9DC2D035699D7A60EEFB431A48FA82" author="Balss, H." box="[1175,1304,1260,1287]" pageId="3" pageNumber="130" pagination="94 - 166" refId="ref5931" refString="Balss, H. (1922) Ostasiatische Decapoden. IV. Die Brachyrhynchen. Archiv Naturgesche (A), 88 (11), 94 - 166, pls. 1 - 2." type="journal article" year="1922">Balss 1922</bibRefCitation>
; H.T. Shih, pers. comm). The morphological differences between
<taxonomicName id="4C0CC4A235699D7A6687FABF1CD8FAAF" authority="Ng &amp; Trivedi &amp; Bhat, 2022" authorityName="Ng &amp; Trivedi &amp; Bhat" authorityYear="2022" box="[766,904,1296,1322]" class="Malacostraca" family="Varunidae" genus="Chhapgarus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B978633335699D7A6687FABF1CD8FAAF" box="[766,904,1296,1322]" italics="true" pageId="3" pageNumber="130">Chhapgarus</emphasis>
</taxonomicName>
<emphasis id="B978633335699D7A67E8FABE1CB1FAAE" bold="true" box="[913,993,1297,1323]" pageId="3" pageNumber="130">
<taxonomicNameLabel id="A24BDE4835699D7A67E8FABE1CB1FAAE" box="[913,993,1297,1323]" pageId="3" pageNumber="130" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0CC4A235699D7A6062FABF1B96FAAF" authorityName="H. Milne Edwards" authorityYear="1837" box="[1051,1222,1296,1322]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A6062FABF1B96FAAF" box="[1051,1222,1296,1322]" italics="true" pageId="3" pageNumber="130">Pseudograpsus</emphasis>
</taxonomicName>
are given in
<tableCitation id="C68E8A9A35699D7A6127FABF1FF6FACA" captionStart="TABLE 1" captionStartId="3.[151,239,1581,1605]" captionTargetPageId="3" captionText="TABLE 1. Morphological comparison of Chhapgarus n. gen. and Pseudograpsus H. Milne Edwards, 1837." httpUri="http://table.plazi.org/id/DF73EFA935699D7A64EEF9821A47F9C0" pageId="3" pageNumber="130" tableUuid="DF73EFA935699D7A64EEF9821A47F9C0">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BB3BF2135699D7A64BEFAF71EB9F986" blockId="3.[151,1437,216,1539]" pageId="3" pageNumber="130">
<taxonomicName id="4C0CC4A235699D7A64BEFAF71E01FAF7" authority="Ng &amp; Trivedi &amp; Bhat, 2022" authorityName="Ng &amp; Trivedi &amp; Bhat" authorityYear="2022" box="[199,337,1368,1394]" class="Malacostraca" family="Varunidae" genus="Chhapgarus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B978633335699D7A64BEFAF71E01FAF7" box="[199,337,1368,1394]" italics="true" pageId="3" pageNumber="130">Chhapgarus</emphasis>
</taxonomicName>
<emphasis id="B978633335699D7A6523FAF61EFAFAF6" bold="true" box="[346,426,1369,1395]" pageId="3" pageNumber="130">
<taxonomicNameLabel id="A24BDE4835699D7A6523FAF61EFAFAF6" box="[346,426,1369,1395]" pageId="3" pageNumber="130" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
, also shows similarity to
<taxonomicName id="4C0CC4A235699D7A66A8FAF61CF4FAF6" authority="White, 1847" authorityName="White" authorityYear="1847" box="[721,932,1368,1395]" class="Malacostraca" family="Varunidae" genus="Utica" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A66A8FAF61C5FFAF7" box="[721,783,1369,1394]" italics="true" pageId="3" pageNumber="130">Utica</emphasis>
White,
<quantity id="4CF412C435699D7A6715FAF61CF4FAF6" box="[876,932,1369,1395]" metricMagnitude="1" metricUnit="m" metricValue="4.69138" pageId="3" pageNumber="130" unit="in" value="1847.0">1847</quantity>
</taxonomicName>
in having setae on the carapace surface and outer surface of chela, but differs from the latter in the following characters: carapace squarish (versus distinctly quadrangular in
<taxonomicName id="4C0CC4A235699D7A652DFA0E1EC2FA3F" authorityName="White" authorityYear="1847" box="[340,402,1441,1466]" class="Malacostraca" family="Varunidae" genus="Utica" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A652DFA0E1EC2FA3F" box="[340,402,1441,1466]" italics="true" pageId="3" pageNumber="130">Utica</emphasis>
</taxonomicName>
); gastric region without “V” shaped ridge (versus prominent “V” shaped ridge on gastric region in
<taxonomicName id="4C0CC4A235699D7A6578FA6A1E6FFA5B" authorityName="White" authorityYear="1847" box="[257,319,1477,1502]" class="Malacostraca" family="Varunidae" genus="Utica" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A6578FA6A1E6FFA5B" box="[257,319,1477,1502]" italics="true" pageId="3" pageNumber="130">Utica</emphasis>
</taxonomicName>
); branchial region without horizontal ridge (versus horizontal setose granulated ridge present across the branchial region in
<taxonomicName id="4C0CC4A235699D7A65E2FA461E89F987" authorityName="White" authorityYear="1847" box="[411,473,1513,1538]" class="Malacostraca" family="Varunidae" genus="Utica" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B978633335699D7A65E2FA461E89F987" box="[411,473,1513,1538]" italics="true" pageId="3" pageNumber="130">Utica</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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