388 lines
38 KiB
XML
388 lines
38 KiB
XML
<document id="65E0A548640F70AD22A7651BB4E708CE" ID-DOI="10.11646/zootaxa.3702.4.8" ID-GBIF-Dataset="7d6bc0a4-2a4a-4b40-ac25-cd6e11ef1952" ID-ISSN="1175-5326" ID-Zenodo-Dep="221634" ID-ZooBank="D1C5C309-6AA4-4FF0-A8C7-6D1164E6CBEB" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458306068066" checkinUser="plazi" docAuthor="Fate, Caitlin, Lapeyrie, Jean & Nel, Andre" docDate="2013" docId="039687A2FF96FF93FF46FD98225AFDA9" docLanguage="en" docName="zt03702p400.pdf.imf" docOrigin="Zootaxa 3702 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Salagoulestes wesleyi Fate, Lapeyrie & Nel, 2013, sp. n." docType="treatment" docVersion="7" lastPageNumber="400" masterDocId="FFAFFFDAFF97FF90FFD1FFEC2111FF85" masterDocTitle="A new Permagrionidae from the Middle Permian of the South of France (Odonatoptera: Protozygoptera)" masterLastPageNumber="400" masterPageNumber="397" pageNumber="398" updateTime="1698185389629" updateUser="plazi">
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<mods:titleInfo id="D4AE7979BF9D659B616961FCA13B9F65">
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<mods:title id="2B3001927702D7D524B5F22F80D72E13">A new Permagrionidae from the Middle Permian of the South of France (Odonatoptera: Protozygoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="FD1A785B388EFF8F224E971756CA0E0E" type="personal">
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<mods:role id="E37FD753D9F24725B3793E00A8786BBE">
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<mods:roleTerm id="BF2CFE7CB61F6318B4301C7F78056BDB">Author</mods:roleTerm>
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<mods:namePart id="0F3951254FD961ADD0924E8BC3569476">Fate, Caitlin</mods:namePart>
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</mods:name>
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<mods:name id="4DB39C855CF2023DA0893425F4FB53F1" type="personal">
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<mods:roleTerm id="819084EBA4D7E79BB69C8EB02CC463EE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="51D43FC53398543F3523D3FBBBFF9FC4">Lapeyrie, Jean</mods:namePart>
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</mods:name>
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<mods:name id="D5591D73C11554F47032AC5194B05FEF" type="personal">
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<mods:roleTerm id="C6442D76E01E56193CE562A1BA22AD45">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="09E88917C089708AA908934BBED54E32">Nel, Andre</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="977661E007BD293B810CD07DB41C14C3">text</mods:typeOfResource>
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<mods:title id="0FD2279F1AC9966B2BBB59C682B35C66">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="30CBE02494C6C9C252FDFE0AD1359275">
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<mods:date id="DA5556515DBA75CB1B652A9485349EB1">2013</mods:date>
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<mods:detail id="1EE488C5AA58B9C1FD2474EB50727FB7" type="volume">
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<mods:number id="14C4221604332F3B588CC1A88DDB783C">3702</mods:number>
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</mods:detail>
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<mods:detail id="C00883C8F84EF74C8E4A0B901D813F5A" type="issue">
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<mods:number id="53CBCCC9C323E55BD0C2C50909E942E1">4</mods:number>
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<mods:classification id="AE9B50A2322FE73590BB0B2E5915916A">journal article</mods:classification>
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<mods:identifier id="2F5C886AE62F0DBA26BA692AE186C3D5" type="DOI">10.11646/zootaxa.3702.4.8</mods:identifier>
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<mods:identifier id="1BAE49A95B1EDD517DDCA8E80644B5C4" type="GBIF-Dataset">7d6bc0a4-2a4a-4b40-ac25-cd6e11ef1952</mods:identifier>
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<mods:identifier id="0B476BB26E87A6F940355215FD567492" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="92FB4BEDB702DC021310DB1CB69F5F49" type="Zenodo-Dep">221634</mods:identifier>
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<treatment id="039687A2FF96FF93FF46FD98225AFDA9" ID-DOI="http://doi.org/10.5281/zenodo.6147820" ID-GBIF-Taxon="117939530" ID-Zenodo-Dep="6147820" LSID="urn:lsid:plazi:treatment:039687A2FF96FF93FF46FD98225AFDA9" httpUri="http://treatment.plazi.org/id/039687A2FF96FF93FF46FD98225AFDA9" lastPageId="3" lastPageNumber="400" pageId="1" pageNumber="398">
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<subSubSection id="C325653FFF96FF91FF46FD982014FD2A" pageId="1" pageNumber="398" type="nomenclature">
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<paragraph id="8B8036B4FF96FF91FF46FD982014FD2A" blockId="1.[151,452,628,687]" pageId="1" pageNumber="398">
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<emphasis id="B94BEAA6FF96FF91FF46FD9820D5FD09" bold="true" box="[151,452,628,652]" pageId="1" pageNumber="398">
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<taxonomicName id="4C3F4D37FF96FF91FF46FD982093FD09" box="[151,386,628,652]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="wesleyi" status="sp. nov.">
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<emphasis id="B94BEAA6FF96FF91FF46FD982093FD09" bold="true" box="[151,386,628,652]" italics="true" pageId="1" pageNumber="398">Salagoulestes wesleyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27857DDFF96FF91FE59FD9920D5FD09" box="[392,452,629,652]" pageId="1" pageNumber="398" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
(
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<figureCitation id="13042A31FF96FF91FF4FFD7421EDFD2A" box="[158,252,664,687]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,701,723]" captionStartId-1="2.[151,250,1369,1391]" captionStartId-2="2.[151,250,1938,1960]" captionTargetBox-0="[227,1360,193,668]" captionTargetBox-1="[224,1358,803,1342]" captionTargetBox-2="[232,1309,1526,1841]" captionTargetId-0="figure@2.[227,1361,193,669]" captionTargetId-1="figure@2.[224,1358,803,1343]" captionTargetId-2="figure@2.[220,1354,1471,1923]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="2" captionText-0="FIGURE 1. Salagoulestes wesleyi gen. n., sp. n., holotype LdLAP 584 A (imprint), Photograph of wing. Scale bar represents 1 mm." captionText-1="FIGURE 2. Salagoulestes wesleyi gen. n., sp. n., holotype LdLAP 584 B (counterimprint), Photograph of wing. Scale bar represents 2 mm." captionText-2="FIGURE 3. Salagoulestes wesleyi gen. n., sp. n., holotype LdLAP 584 A – B, reconstruction of wing. Scale bar represents 2 mm." httpUri-0="https://zenodo.org/record/221635/files/figure.png" httpUri-1="https://zenodo.org/record/221636/files/figure.png" httpUri-2="https://zenodo.org/record/221637/files/figure.png" pageId="1" pageNumber="398">Figs 1–3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C325653FFF96FF91FF46FD34253FFD6A" box="[151,1070,728,751]" pageId="1" pageNumber="398" type="materials_examined">
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<paragraph id="8B8036B4FF96FF91FF46FD34253FFD6A" blockId="1.[151,1437,728,2033]" box="[151,1070,728,751]" pageId="1" pageNumber="398">
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<emphasis id="B94BEAA6FF96FF91FF46FD3421EEFD6A" bold="true" box="[151,255,728,751]" pageId="1" pageNumber="398">
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<typeStatus id="54848816FF96FF91FF46FD3421EEFD6A" box="[151,255,728,751]" pageId="1" pageNumber="398" type="holotype">Holotype</typeStatus>
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</emphasis>
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. Specimen LdLAP 584 A–B, Lapeyrie coll., Musée Fleury. Lodève,
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<collectingCountry id="F3287624FF96FF91FC0FFD34253BFD6A" box="[990,1066,728,751]" name="France" pageId="1" pageNumber="398">France</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C325653FFF96FF91FF14FD16234CFC94" box="[197,605,762,785]" pageId="1" pageNumber="398" type="diagnosis">
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<paragraph id="8B8036B4FF96FF91FF14FD16234CFC94" blockId="1.[151,1437,728,2033]" box="[197,605,762,785]" pageId="1" pageNumber="398">
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<emphasis id="B94BEAA6FF96FF91FF14FD162023FC94" bold="true" box="[197,306,762,785]" pageId="1" pageNumber="398">Diagnosis</emphasis>
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. The same as for the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C325653FFF96FF91FF14FCF120D9FCFC" pageId="1" pageNumber="398" type="etymology">
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<paragraph id="8B8036B4FF96FF91FF14FCF1226DFCB1" blockId="1.[151,1437,728,2033]" box="[197,892,797,820]" pageId="1" pageNumber="398">
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<emphasis id="B94BEAA6FF96FF91FF14FCF1202FFCB1" bold="true" box="[197,318,797,820]" pageId="1" pageNumber="398">Etymology</emphasis>
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. Named after Dale Wesley Fate, father of first author.
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</paragraph>
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<paragraph id="8B8036B4FF96FF91FF14FCAC20D9FCFC" blockId="1.[151,1437,728,2033]" pageId="1" pageNumber="398">
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<emphasis id="B94BEAA6FF96FF91FF14FCAC209AFCD2" bold="true" box="[197,395,832,855]" pageId="1" pageNumber="398">Age and outcrop</emphasis>
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. Middle Permian, Guadalupian, Mérifons Member, Salagou Formation, outcrop “La Prade”, Lodève, Languedoc,
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<collectingCountry id="F3287624FF96FF91FEA9FC8E20D4FCFC" box="[376,453,866,889]" name="France" pageId="1" pageNumber="398">France</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C325653FFF96FF91FF14FC692595FA9C" pageId="1" pageNumber="398" type="description">
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<paragraph id="8B8036B4FF96FF91FF14FC692595FA9C" blockId="1.[151,1437,728,2033]" pageId="1" pageNumber="398">
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<emphasis id="B94BEAA6FF96FF91FF14FC69205BFC19" bold="true" box="[197,330,901,924]" pageId="1" pageNumber="398">Description</emphasis>
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. Imprint and counterimprint of a wing with apical part, base of anal vein, and subcostal area missing;
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<quantity id="4CC79B51FF96FF91FF46FC4421E4FC3A" box="[151,245,936,959]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="398" unit="mm" value="12.8">12.8 mm</quantity>
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long,
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<quantity id="4CC79B51FF96FF91FEE6FC442099FC3A" box="[311,392,936,959]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="398" unit="mm" value="2.7">2.7 mm</quantity>
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wide; distance from base to nodus
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<quantity id="4CC79B51FF96FF91FD2DFC442241FC3A" box="[764,848,936,959]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="398" unit="mm" value="5.1">5.1 mm</quantity>
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, to arculus
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<quantity id="4CC79B51FF96FF91FC19FC44250DFC3A" box="[968,1052,936,959]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="398" unit="mm" value="2.6">2.6 mm</quantity>
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; from nodus to pterostigma
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<quantity id="4CC79B51FF96FF91FA95FC442489FC3A" box="[1348,1432,936,959]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="1" pageNumber="398" unit="mm" value="6.6">6.6 mm</quantity>
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||
,
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<collectingCountry id="F3287624FF96FF91FF46FC2621A9FC64" box="[151,184,970,993]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
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||
separating from MA
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||
<quantity id="4CC79B51FF96FF91FE73FC2620E5FC64" box="[418,500,970,993]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="398" unit="mm" value="0.9">0.9 mm</quantity>
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||
from distal base of arculus; MAa and MAb separating
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||
<quantity id="4CC79B51FF96FF91FB9BFC26258CFC64" box="[1098,1181,970,993]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="398" unit="mm" value="0.4">0.4 mm</quantity>
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from base of
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||
<collectingCountry id="F3287624FF96FF91FAE5FC262449FC64" box="[1332,1368,970,993]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
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||
; MAb
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||
<quantity id="4CC79B51FF96FF91FF46FC0121FBFB81" box="[151,234,1005,1028]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="398" unit="mm" value="0.3">0.3 mm</quantity>
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long; MP reaching CuA
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<quantity id="4CC79B51FF96FF91FDD1FC012345FB81" box="[512,596,1005,1028]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="398" unit="mm" value="1.3">1.3 mm</quantity>
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from base of CuA, CuA separating again from MP
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||
<quantity id="4CC79B51FF96FF91FB5EFC0125F2FB81" box="[1167,1251,1005,1028]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="398" unit="mm" value="3.3">3.3 mm</quantity>
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||
from wing base, slightly distal of separation of
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<collectingCountry id="F3287624FF96FF91FE08FBFC20EBFBA2" box="[473,506,1040,1063]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
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||
from MA; CuP not preserved; wing petiolated with posterior margin not preserved; one complete antenodal vein preserved (Ax1?),
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<quantity id="4CC79B51FF96FF91FDBBFBDE23AAFBCC" box="[618,699,1074,1097]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="398" unit="mm" value="1.2">1.2 mm</quantity>
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basal to arculus; Cr and subnodus aligned and strongly oblique; four postnodals; four postsubnodals, not aligned with postnodals; Asn present; base of IR1
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<quantity id="4CC79B51FF96FF91FBE5FBB92597FBE9" box="[1076,1158,1109,1132]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="398" unit="mm" value="8.7">8.7 mm</quantity>
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||
distal of separation of
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<collectingCountry id="F3287624FF96FF91FAAAFBB9248DFBE9" box="[1403,1436,1109,1132]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
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and MA, base of
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||
<collectingCountry id="F3287624FF96FF91FE9CFB94207EFB0A" box="[333,367,1144,1167]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
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||
2
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||
<quantity id="4CC79B51FF96FF91FE50FB9420C5FB0A" box="[385,468,1144,1167]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="1" pageNumber="398" unit="mm" value="7.6">7.6 mm</quantity>
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, IR2
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<quantity id="4CC79B51FF96FF91FDDDFB942371FB0A" box="[524,608,1144,1167]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="398" unit="mm" value="5.9">5.9 mm</quantity>
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||
,
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<collectingCountry id="F3287624FF96FF91FDBBFB94239DFB0A" box="[618,652,1144,1167]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
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||
3/4
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||
<quantity id="4CC79B51FF96FF91FD62FB942217FB0A" box="[691,774,1144,1167]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="398" unit="mm" value="2.9">2.9 mm</quantity>
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, thus all distal of subnodus;
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<collectingCountry id="F3287624FF96FF91FBE0FB942542FB0A" box="[1073,1107,1144,1167]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
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2 apparently branching on IR2, with its real base appearing as a rather weakly oblique vein between it and
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<collectingCountry id="F3287624FF96FF91FC7EFB7622C7FB34" box="[943,982,1178,1201]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
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1; no secondary longitudinal veins between main veins in preserved parts; longitudinal veins, except CuA, at their distal part, are straight; area between MP and CuA distally reduced; cubital area with one row of cells and straight cross-veins; Pterostigma elongated, probably sclerotized, darker than the wing membrane, ca.
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<quantity id="4CC79B51FF96FF91FDCDFAEE237CFA9C" box="[540,621,1282,1305]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="398" unit="mm" value="1.7">1.7 mm</quantity>
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||
long,
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<quantity id="4CC79B51FF96FF91FD60FAEE221EFA9C" box="[689,783,1282,1305]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="398" unit="mm" value="0.25">0.25 mm</quantity>
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||
wide, with one crossvein below it.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C325653FFF96FF93FF14FAC9225AFDA9" lastPageId="3" lastPageNumber="400" pageId="1" pageNumber="398" type="discussion">
|
||
<paragraph id="8B8036B4FF96FF91FF14FAC92012F989" blockId="1.[151,1437,728,2033]" pageId="1" pageNumber="398">
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||
<emphasis id="B94BEAA6FF96FF91FF14FAC9202CFAB9" bold="true" box="[197,317,1317,1340]" pageId="1" pageNumber="398">Discussion</emphasis>
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.
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<taxonomicName id="4C3F4D37FF96FF91FE98FAC82321FAB9" box="[329,560,1316,1340]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="wesleyi" status="gen. nov.">
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<emphasis id="B94BEAA6FF96FF91FE98FAC82321FAB9" box="[329,560,1316,1340]" italics="true" pageId="1" pageNumber="398">Salagoulestes wesleyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27857DDFF96FF91FDE6FAC92390FAB9" box="[567,641,1317,1340]" pageId="1" pageNumber="398" rank="genus">
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||
<emphasis id="B94BEAA6FF96FF91FDE6FAC92390FAB9" bold="true" box="[567,641,1317,1340]" pageId="1" pageNumber="398">gen. n.</emphasis>
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</taxonomicNameLabel>
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,
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<emphasis id="B94BEAA6FF96FF91FD5CFAC923D9FAB9" bold="true" box="[653,712,1317,1340]" pageId="1" pageNumber="398">sp. n.</emphasis>
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can be considered as a member of the Protozygoptera + Panodonata because it has a true pterostigma, CuA captured by MP, and reduction of CuP to a short vein (not visible here but certainly reduced). It has two obvious characters present in Protozygoptera, viz. the point of separation between CuA and MP not aligned with the posterior branch of MA (MAb) and the nodal crossvein Cr and subnodus oblique and well aligned, but as plesiomorphies, these characters are not sufficient to attribute
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<taxonomicName id="4C3F4D37FF96FF91FC0FFA432563FA42" box="[990,1138,1455,1479]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B94BEAA6FF96FF91FC0FFA432563FA42" box="[990,1138,1455,1479]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
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</taxonomicName>
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to this group. Nevertheless
|
||
<taxonomicName id="4C3F4D37FF96FF91FF46FA3D203CFA6C" box="[151,301,1489,1513]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF96FF91FF46FA3D203CFA6C" box="[151,301,1489,1513]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
shares with the
|
||
<taxonomicName id="4C3F4D37FF96FF91FE33FA3E239FFA6C" box="[482,654,1490,1513]" class="Insecta" family="Permagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="family">Permagrionidae</taxonomicName>
|
||
the apomorphic presence of an ASn vein between RA and
|
||
<collectingCountry id="F3287624FF96FF91FACCFA3E242FFA6C" box="[1309,1342,1490,1513]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
|
||
basal of subnodus.
|
||
</paragraph>
|
||
<paragraph id="8B8036B4FF96FF91FF16F9F4200FF8E2" blockId="1.[151,1437,728,2033]" pageId="1" pageNumber="398">
|
||
Within this family,
|
||
<taxonomicName id="4C3F4D37FF96FF91FE4AF9FB233FF9AA" box="[411,558,1559,1583]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF96FF91FE4AF9FB233FF9AA" box="[411,558,1559,1583]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<emphasis id="B94BEAA6FF96FF91FD11F9F42250F9AA" box="[704,833,1560,1583]" italics="true" pageId="1" pageNumber="398">Permagrion</emphasis>
|
||
<taxonomicName id="4C3F4D37FF96FF91FC9BF9FB256BF9AA" authority="Tillyard, 1928" authorityName="Tillyard" authorityYear="1928" box="[842,1146,1559,1583]" pageId="1" pageNumber="398" rank="species" species="falklandicum">
|
||
<emphasis id="B94BEAA6FF96FF91FC9BF9FB22C4F9AA" box="[842,981,1559,1583]" italics="true" pageId="1" pageNumber="398">falklandicum</emphasis>
|
||
Tillyard, 1928
|
||
</taxonomicName>
|
||
, in the CuA much shorter, ending on posterior wing margin at level of base of IR2, instead of base of IR1.
|
||
<taxonomicName id="4C3F4D37FF96FF91FBFDF9D525AEF9D4" box="[1068,1215,1593,1617]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF96FF91FBFDF9D525AEF9D4" box="[1068,1215,1593,1617]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
differs
|
||
<emphasis id="B94BEAA6FF96FF91FACBF9D521F9F9F1" italics="true" pageId="1" pageNumber="398">Permolestes gracilis</emphasis>
|
||
Martynov, 1932,
|
||
<emphasis id="B94BEAA6FF96FF91FE74F9B023A2F9F1" box="[421,691,1628,1652]" italics="true" pageId="1" pageNumber="398">
|
||
Permolestes
|
||
<taxonomicName id="4C3F4D37FF96FF91FDFFF9B023A2F9F1" authority="Nel et al., 2012" authorityName="Nel et al." authorityYear="2012" box="[558,691,1628,1652]" pageId="1" pageNumber="398" rank="species" species="sheimogorai">sheimogorai</taxonomicName>
|
||
</emphasis>
|
||
Nel
|
||
<emphasis id="B94BEAA6FF96FF91FD37F9B1220DF9F1" box="[742,796,1628,1652]" italics="true" pageId="1" pageNumber="398">et al.</emphasis>
|
||
, 2012, and
|
||
<emphasis id="B94BEAA6FF96FF91FC49F9B025B9F9F1" box="[920,1192,1628,1652]" italics="true" pageId="1" pageNumber="398">
|
||
Permolestes
|
||
<taxonomicName id="4C3F4D37FF96FF91FBF1F9B125B9F9F1" authority="Nel et al., 2012" authorityName="Nel et al." authorityYear="2012" box="[1056,1192,1629,1652]" pageId="1" pageNumber="398" rank="species" species="soyanaiensis">soyanaiensis</taxonomicName>
|
||
</emphasis>
|
||
Nel
|
||
<emphasis id="B94BEAA6FF96FF91FB0DF9B12400F9F1" box="[1244,1297,1628,1652]" italics="true" pageId="1" pageNumber="398">et al.</emphasis>
|
||
, 2012, in the presence of three rows of cells or more in area between CuA and posterior wing margin.
|
||
<taxonomicName id="4C3F4D37FF96FF91FB4EF96C2458F912" box="[1183,1353,1663,1687]" pageId="1" pageNumber="398" rank="species" species="falklandicum">
|
||
<emphasis id="B94BEAA6FF96FF91FB4EF96C2458F912" box="[1183,1353,1663,1687]" italics="true" pageId="1" pageNumber="398">P. falklandicum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F4D37FF96FF91FA59F96C200EF93C" pageId="1" pageNumber="398" rank="species" species="soyanaiensis">
|
||
<emphasis id="B94BEAA6FF96FF91FA59F96C2484F912" box="[1416,1429,1664,1687]" italics="true" pageId="1" pageNumber="398">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAA6FF96FF91FF46F94E200EF93C" box="[151,287,1698,1721]" italics="true" pageId="1" pageNumber="398">soyanaiensis</emphasis>
|
||
</taxonomicName>
|
||
also have shorter pterostigmas (character state unknown in
|
||
<taxonomicName id="4C3F4D37FF96FF91FC3BF94E2583F93C" box="[1002,1170,1697,1721]" pageId="1" pageNumber="398" rank="species" species="sheimogorai">
|
||
<emphasis id="B94BEAA6FF96FF91FC3BF94E2583F93C" box="[1002,1170,1697,1721]" italics="true" pageId="1" pageNumber="398">P. sheimogorai</emphasis>
|
||
</taxonomicName>
|
||
), unlike
|
||
<taxonomicName id="4C3F4D37FF96FF91FAD2F94D2487F93C" box="[1283,1430,1697,1721]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF96FF91FAD2F94D2487F93C" box="[1283,1430,1697,1721]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B94BEAA6FF96FF91FF46F9282065F959" box="[151,372,1732,1756]" italics="true" pageId="1" pageNumber="398">Scytolestes stigmalis</emphasis>
|
||
Martynov, 1937, has also three rows of cells in cubital area, and it seems to have a shorter petiole relatively to the distance between arculus and nodus, than in
|
||
<taxonomicName id="4C3F4D37FF96FF91FCEAF90B22DFF97A" box="[827,974,1767,1791]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF96FF91FCEAF90B22DFF97A" box="[827,974,1767,1791]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
. The three species of
|
||
<emphasis id="B94BEAA6FF96FF91FB15F90B2435F97A" box="[1220,1316,1767,1791]" italics="true" pageId="1" pageNumber="398">Epilestes</emphasis>
|
||
Martynov, 1937 (
|
||
<emphasis id="B94BEAA6FF96FF91FF0EF8E6206BF8A4" box="[223,378,1801,1825]" italics="true" pageId="1" pageNumber="398">E. kargalensis</emphasis>
|
||
Martynov, 1932,
|
||
<emphasis id="B94BEAA6FF96FF91FD90F8E623E5F8A4" box="[577,756,1802,1825]" italics="true" pageId="1" pageNumber="398">E. angustapterix</emphasis>
|
||
Nel
|
||
<emphasis id="B94BEAA6FF96FF91FCFFF8E62276F8A4" box="[814,871,1801,1825]" italics="true" pageId="1" pageNumber="398">et al.</emphasis>
|
||
, 2012, and
|
||
<emphasis id="B94BEAA6FF96FF91FC3CF8E62549F8A4" box="[1005,1112,1801,1825]" italics="true" pageId="1" pageNumber="398">E. gallica</emphasis>
|
||
Nel
|
||
<emphasis id="B94BEAA6FF96FF91FB42F8E625DAF8A4" box="[1171,1227,1801,1825]" italics="true" pageId="1" pageNumber="398">et al.</emphasis>
|
||
, 1999) differ from
|
||
<taxonomicName id="4C3F4D37FF96FF91FF46F8C0203BF8C1" box="[151,298,1836,1860]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF96FF91FF46F8C0203BF8C1" box="[151,298,1836,1860]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
in the presence of two-three rows of cells in cubital area and a longer CuA reaching the level of the base of
|
||
<collectingCountry id="F3287624FF96FF91FF46F8BC21A8F8E2" box="[151,185,1872,1895]" name="Poland" pageId="1" pageNumber="398">RP</collectingCountry>
|
||
2 at least.
|
||
</paragraph>
|
||
<paragraph id="8B8036B4FF96FF91FF16F89D227FF874" blockId="1.[151,1437,728,2033]" pageId="1" pageNumber="398">
|
||
<emphasis id="B94BEAA6FF96FF91FF16F89D20C2F80C" box="[199,467,1905,1929]" italics="true" pageId="1" pageNumber="398">Solikamptilon pectinatus</emphasis>
|
||
Nel
|
||
<emphasis id="B94BEAA6FF96FF91FDC1F89E235BF80C" box="[528,586,1905,1929]" italics="true" pageId="1" pageNumber="398">et al.</emphasis>
|
||
, 2012, differs from
|
||
<taxonomicName id="4C3F4D37FF96FF91FCE0F89D22D5F80C" box="[817,964,1905,1929]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF96FF91FCE0F89D22D5F80C" box="[817,964,1905,1929]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
in the presence of sigmoidal veinlets and transversely elongate cells in cubital area, even if they share a zigzagged CuA.
|
||
<emphasis id="B94BEAA6FF96FF91FBDCF878243EF829" box="[1037,1327,1940,1964]" italics="true" pageId="1" pageNumber="398">Solikamptilon remuliforme</emphasis>
|
||
Zalessky, 1948, has a very long CuA ending at the level of the base of IR1, and sigmoidal veinlets in cubital area. Both have fewer postnodal and postsubnodal cells and crossveins than
|
||
<taxonomicName id="4C3F4D37FF96FF91FD05F8352276F874" box="[724,871,2009,2033]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF96FF91FD05F8352276F874" box="[724,871,2009,2033]" italics="true" pageId="1" pageNumber="398">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF40663CFF95FF92FF46FD5121C8FD74" httpUri="https://zenodo.org/record/221635/files/figure.png" pageId="2" pageNumber="399" targetBox="[227,1360,193,668]" targetPageId="2">
|
||
<paragraph id="8B8036B4FF95FF92FF46FD5121C8FD74" blockId="2.[151,1436,700,753]" pageId="2" pageNumber="399">
|
||
<emphasis id="B94BEAA6FF95FF92FF46FD512005FD57" bold="true" box="[151,276,701,723]" pageId="2" pageNumber="399">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C3F4D37FF95FF92FECBFD5120E6FD57" box="[282,503,701,722]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="wesleyi" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF95FF92FECBFD5120E6FD57" box="[282,503,701,722]" italics="true" pageId="2" pageNumber="399">Salagoulestes wesleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27857DDFF95FF92FE2CFD512354FD57" box="[509,581,701,722]" pageId="2" pageNumber="399" rank="genus">
|
||
<emphasis id="B94BEAA6FF95FF92FE2CFD512354FD57" bold="true" box="[509,581,701,722]" pageId="2" pageNumber="399">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B94BEAA6FF95FF92FD83FD51239AFD57" bold="true" box="[594,651,701,722]" pageId="2" pageNumber="399">sp. n.</emphasis>
|
||
, holotype LdLAP 584 A (imprint), Photograph of wing. Scale bar represents 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF40663CFF95FF92FF46FAB52054FA08" httpUri="https://zenodo.org/record/221636/files/figure.png" pageId="2" pageNumber="399" targetBox="[224,1358,803,1342]" targetPageId="2">
|
||
<paragraph id="8B8036B4FF95FF92FF46FAB52054FA08" blockId="2.[151,1436,1369,1421]" pageId="2" pageNumber="399">
|
||
<emphasis id="B94BEAA6FF95FF92FF46FAB52004FAEB" bold="true" box="[151,277,1369,1391]" pageId="2" pageNumber="399">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C3F4D37FF95FF92FECFFAB520ECFAEB" box="[286,509,1369,1390]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="wesleyi" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF95FF92FECFFAB520ECFAEB" box="[286,509,1369,1390]" italics="true" pageId="2" pageNumber="399">Salagoulestes wesleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27857DDFF95FF92FDD4FAB5235EFAEB" box="[517,591,1369,1390]" pageId="2" pageNumber="399" rank="genus">
|
||
<emphasis id="B94BEAA6FF95FF92FDD4FAB5235EFAEB" bold="true" box="[517,591,1369,1390]" pageId="2" pageNumber="399">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B94BEAA6FF95FF92FD8FFAB52388FAEB" bold="true" box="[606,665,1369,1390]" pageId="2" pageNumber="399">sp. n.</emphasis>
|
||
, holotype LdLAP 584 B (counterimprint), Photograph of wing. Scale bar represents 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF40663CFF95FF92FF46F87E21D7F843" httpUri="https://zenodo.org/record/221637/files/figure.png" pageId="2" pageNumber="399" targetBox="[232,1309,1526,1841]" targetPageId="2">
|
||
<paragraph id="8B8036B4FF95FF92FF46F87E21D7F843" blockId="2.[151,1435,1938,1990]" pageId="2" pageNumber="399">
|
||
<emphasis id="B94BEAA6FF95FF92FF46F87E2005F822" bold="true" box="[151,276,1938,1960]" pageId="2" pageNumber="399">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C3F4D37FF95FF92FECDF87E20EBF822" box="[284,506,1938,1959]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="wesleyi" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF95FF92FECDF87E20EBF822" box="[284,506,1938,1959]" italics="true" pageId="2" pageNumber="399">Salagoulestes wesleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27857DDFF95FF92FDD3F87E235AF822" box="[514,587,1938,1959]" pageId="2" pageNumber="399" rank="genus">
|
||
<emphasis id="B94BEAA6FF95FF92FDD3F87E235AF822" bold="true" box="[514,587,1938,1959]" pageId="2" pageNumber="399">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B94BEAA6FF95FF92FD88F87E2385F822" bold="true" box="[601,660,1938,1959]" pageId="2" pageNumber="399">sp. n.</emphasis>
|
||
, holotype LdLAP 584 A–B, reconstruction of wing. Scale bar represents 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8036B4FF94FF93FF16FF7B25EBFF71" blockId="3.[151,1437,151,556]" pageId="3" pageNumber="400">
|
||
<taxonomicName id="4C3F4D37FF94FF93FF16FF7B204BFF2A" box="[199,346,151,175]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF94FF93FF16FF7B204BFF2A" box="[199,346,151,175]" italics="true" pageId="3" pageNumber="400">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
has a
|
||
<collectingCountry id="F3287624FF94FF93FE4CFF7420AFFF2A" box="[413,446,152,175]" name="Poland" pageId="3" pageNumber="400">RP</collectingCountry>
|
||
2 that apparently emerges from IR2 as in
|
||
<emphasis id="B94BEAA6FF94FF93FCABFF7B22FEFF2A" box="[890,1007,151,175]" italics="true" pageId="3" pageNumber="400">Scytolestes</emphasis>
|
||
and
|
||
<emphasis id="B94BEAA6FF94FF93FBF2FF7425B5FF2A" box="[1059,1188,152,175]" italics="true" pageId="3" pageNumber="400">Permagrion</emphasis>
|
||
, but not as in
|
||
<emphasis id="B94BEAA6FF94FF93FAEDFF7B248DFF2A" box="[1340,1436,151,175]" italics="true" pageId="3" pageNumber="400">Epilestes</emphasis>
|
||
and
|
||
<emphasis id="B94BEAA6FF94FF93FF14FF552048FF54" box="[197,345,185,209]" italics="true" pageId="3" pageNumber="400">Solikamptilon</emphasis>
|
||
. This character state is a potential synapomorphy of the three genera
|
||
<taxonomicName id="4C3F4D37FF94FF93FB81FF5525F3FF54" box="[1104,1250,185,209]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF94FF93FB81FF5525F3FF54" box="[1104,1250,185,209]" italics="true" pageId="3" pageNumber="400">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B94BEAA6FF94FF93FB23FF552476FF54" box="[1266,1383,185,209]" italics="true" pageId="3" pageNumber="400">Scytolestes</emphasis>
|
||
, and
|
||
<emphasis id="B94BEAA6FF94FF93FF46FF312009FF71" box="[151,280,221,244]" italics="true" pageId="3" pageNumber="400">Permagrion</emphasis>
|
||
.
|
||
<emphasis id="B94BEAA6FF94FF93FEF5FF3020B7FF71" box="[292,422,220,244]" italics="true" pageId="3" pageNumber="400">Permolestes</emphasis>
|
||
is in an intermediate situation with base of
|
||
<collectingCountry id="F3287624FF94FF93FCA8FF31228BFF71" box="[889,922,221,244]" name="Poland" pageId="3" pageNumber="400">RP</collectingCountry>
|
||
2 midway between
|
||
<collectingCountry id="F3287624FF94FF93FBB8FF31259FFF71" box="[1129,1166,221,244]" name="Poland" pageId="3" pageNumber="400">RP</collectingCountry>
|
||
1 and IR2.
|
||
</paragraph>
|
||
<paragraph id="8B8036B4FF94FF93FF14FF13255EFED9" blockId="3.[151,1437,151,556]" pageId="3" pageNumber="400">
|
||
<taxonomicName id="4C3F4D37FF94FF93FF14FF132049FE92" box="[197,344,255,279]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF94FF93FF14FF132049FE92" box="[197,344,255,279]" italics="true" pageId="3" pageNumber="400">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3F4D37FF94FF93FE35FF13224BFE92" authority="Nel et al., 1999" authorityName="Nel et al." authorityYear="1999" box="[484,858,255,279]" class="Insecta" genus="Lodevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="longialata">
|
||
<emphasis id="B94BEAA6FF94FF93FE35FF1323BEFE92" box="[484,687,255,279]" italics="true" pageId="3" pageNumber="400">Lodevia longialata</emphasis>
|
||
Nel
|
||
<emphasis id="B94BEAA6FF94FF93FD32FEEC2209FE92" box="[739,792,255,279]" italics="true" pageId="3" pageNumber="400">et al.</emphasis>
|
||
, 1999
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54848816FF94FF93FCB8FEEC2289FE92" box="[873,920,256,279]" pageId="3" pageNumber="400">type</typeStatus>
|
||
species of the Lodeviidae Nel
|
||
<emphasis id="B94BEAA6FF94FF93FB32FEEC2409FE92" box="[1251,1304,255,279]" italics="true" pageId="3" pageNumber="400">et al.</emphasis>
|
||
, 2012, from Lodève Basin), in the nodal Cr well aligned with subnodus, CuA more zigzagged and ending on posterior wing margin well basal of level of base of
|
||
<collectingCountry id="F3287624FF94FF93FE01FEA920E2FED9" box="[464,499,325,348]" name="Poland" pageId="3" pageNumber="400">RP</collectingCountry>
|
||
2, and many fewer cells and crossveins (Nel
|
||
<emphasis id="B94BEAA6FF94FF93FC1FFEA92515FED9" box="[974,1028,324,348]" italics="true" pageId="3" pageNumber="400">et al.</emphasis>
|
||
1999).
|
||
</paragraph>
|
||
<paragraph id="8B8036B4FF94FF93FF14FE84225AFDA9" blockId="3.[151,1437,151,556]" pageId="3" pageNumber="400">
|
||
<collectingCountry id="F3287624FF94FF93FF14FE8421F4FEFA" box="[197,229,360,383]" name="American Samoa" pageId="3" pageNumber="400">As</collectingCountry>
|
||
the second representative of the
|
||
<taxonomicName id="4C3F4D37FF94FF93FDA2FE84223FFEFA" box="[627,814,360,383]" class="Insecta" family="Permagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="400" phylum="Arthropoda" rank="family">Permagrionidae</taxonomicName>
|
||
and third Protozygoptera in the Salagou Formation,
|
||
<taxonomicName id="4C3F4D37FF94FF93FF46FE65203BFE24" box="[151,298,393,417]" class="Insecta" family="Permagrionidae" genus="Salagoulestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94BEAA6FF94FF93FF46FE65203BFE24" box="[151,298,393,417]" italics="true" pageId="3" pageNumber="400">Salagoulestes</emphasis>
|
||
</taxonomicName>
|
||
confirms the hypothesis that this group was relatively diverse in this palaeoenvironment. More generally the Odonatoptera are quite well represented in this place with nine species of
|
||
<taxonomicName id="4C3F4D37FF94FF93FBD1FE412479FE41" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[1024,1384,429,452]" class="Insecta" family="Meganeuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protodonata" pageId="3" pageNumber="400" phylum="Arthropoda" rank="family">Meganeuridae Handlirsch, 1906</taxonomicName>
|
||
, one Lapeyriidae Nel
|
||
<emphasis id="B94BEAA6FF94FF93FE9BFE3C2091FE62" box="[330,384,463,487]" italics="true" pageId="3" pageNumber="400">et al.</emphasis>
|
||
1999, and even the oldest Panodonata,
|
||
<emphasis id="B94BEAA6FF94FF93FCF7FE232518FE62" box="[806,1033,463,487]" italics="true" pageId="3" pageNumber="400">Saxonagrion minutus</emphasis>
|
||
Nel
|
||
<emphasis id="B94BEAA6FF94FF93FBECFE3C2563FE62" box="[1085,1138,463,487]" italics="true" pageId="3" pageNumber="400">et al.</emphasis>
|
||
, 1999 (Nel
|
||
<emphasis id="B94BEAA6FF94FF93FB23FE3C2439FE62" box="[1266,1320,463,487]" italics="true" pageId="3" pageNumber="400">et al.</emphasis>
|
||
1999a,b,c, 2008, 2009). The absence in this place of the Permian clade Protanisoptera, well recorded in North and South
|
||
<collectingCountry id="F3287624FF94FF93FAE8FE1E2489FD8C" box="[1337,1432,498,521]" name="United States of America" pageId="3" pageNumber="400">America</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3287624FF94FF93FF46FDF921EAFDA9" box="[151,251,533,556]" name="Australia" pageId="3" pageNumber="400">Australia</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F3287624FF94FF93FEE2FDF9206BFDA9" box="[307,378,533,556]" name="Russia" pageId="3" pageNumber="400">Russia</collectingCountry>
|
||
(Huguet
|
||
<emphasis id="B94BEAA6FF94FF93FE0EFDF92304FDA9" box="[479,533,532,556]" italics="true" pageId="3" pageNumber="400">et al.</emphasis>
|
||
2002), remains unexplained.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |