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<document id="2C4EC29569CAE378E0A2EF02182029EF" ID-CLB-Dataset="22094" ID-DOI="10.5852/ejt.2015.162" ID-GBIF-Dataset="e7c55171-03d5-4714-a0c2-b7e70ab80fce" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829511" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1504785881774" checkinUser="jeremy" docAuthor="Huber, Bernhard A., Nuñeza, Olga M. &amp; Ung, Charles Leh Moi" docDate="2015" docId="0390E827601DFFB09597FD7F46C0FD81" docLanguage="en" docName="ejt-2015-162.pdf.imf" docOrigin="European Journal of Taxonomy 162" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aetana kinabalu Huber 2005" docType="treatment" docVersion="6" lastPageNumber="38" masterDocId="FFA9905F603FFF959719FF814549FFA4" masterDocTitle="Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="35" updateTime="1698468106960" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6856D4494011359DD391D2DA83E1A987">Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="97D57F2A661220297405E198B6926FDB">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="05B42B2C7CE8526A77D82962D2BF3F7D">Alexander Koenig Research Museum of Zoology, Adenauerallee 160, 53113 Bonn, Germany. &amp; urn: lsid: zoobank. org: author: 33607 F 65 - 19 BF- 4 DC 9 - 94 FD- 4 BB 88 CED 455 F</mods:affiliation>
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<mods:namePart id="2F08FA60B5C0002CEC639E55211B62BB">Nuñeza, Olga M.</mods:namePart>
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<mods:affiliation id="4641F827D6C238276E8D157D5A2BA7A4">Department of Biological Sciences, Mindanao State University - Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines. &amp; Email: olgamnuneza @ yahoo. com &amp; urn: lsid: zoobank. org: author: 6 D 14 FCCC-F 148 - 4293 - 9 F 23 - 0 EB 5 A 4119305</mods:affiliation>
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<mods:namePart id="4AF6046472A9DDD59C1A19AE94A8AD8B">Ung, Charles Leh Moi</mods:namePart>
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<mods:affiliation id="CCECECD3E622E5BC3D1A2243EDAEDA2C">Sarawak Museum, Jalan Tun Abang Haji Openg, 93566 Kuching, Sarawak, Malaysia. &amp; Email: charllmu @ sarawak. gov. my &amp; urn: lsid: zoobank. org: author: 8738 F 336 - D 1 CC- 4 AD 3 - 97 E 4 - 82 BC 1729 A 123</mods:affiliation>
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<taxonomicName id="4C3922B2601DFFB79597FD7F46B8FCBC" ID-CoL="65BCS" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[654,1009,765,792]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523601DFFB79597FD7F4611FCBC" bold="true" box="[654,856,765,792]" italics="true" pageId="34" pageNumber="35">Aetana kinabalu</emphasis>
Huber, 2005
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<paragraph id="8B865931601DFFB7959AFCA046B5FC98" blockId="34.[643,1020,765,828]" box="[643,1020,801,828]" pageId="34" pageNumber="35">
<figureCitation id="130245B4601DFFB7959AFCA04651FC9F" box="[643,792,801,827]" captionStart="Figs 99104" captionStartId="35.[227,279,1930,1956]" captionTargetBox="[227,1437,731,1888]" captionTargetId="figure@35.[227,1437,731,1888]" captionTargetPageId="35" captionText="Figs 99104. Live specimens. Aetana kinabalu group. — 99103. A. kinabalu Huber, 2005. ♂ and ♀ with eggsac from Gunung Mulu, Sarawak (99, 100); ♀ with eggsac and ♂ from Crocker Range, Sabah (101, 102); and ♂ from Mt. Kinabalu, Sabah (103). — 104. A. gaya Huber, sp. nov., ♂ from Gaya Island, Sabah." figureDoi="http://doi.org/10.5281/zenodo.3829555" httpUri="https://zenodo.org/record/3829555/files/figure.png" pageId="34" pageNumber="35">Figs 99103</figureCitation>
,
<figureCitation id="130245B4601DFFB7943AFCA046C0FC9F" box="[803,905,801,827]" captionStart="Figs 111112" captionStartId="37.[227,279,1997,2023]" captionTargetBox="[239,1430,1527,1927]" captionTargetId="figure@37.[227,1437,1519,1939]" captionTargetPageId="37" captionText="Figs 111112. Aetana kinabalu Huber, 2005. Cleared female genitalia, ventral and dorsal views (arrow in Fig. 112 points at membranous pocket). ip = internal sclerotized pocket. Scale line: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829560" httpUri="https://zenodo.org/record/3829560/files/figure.png" pageId="34" pageNumber="35">111112</figureCitation>
,
<figureCitation id="130245B4601DFFB7948CFCA046B5FC98" box="[917,1020,801,828]" captionStart="Figs 163177" captionStartId="51.[227,279,1968,1994]" captionTargetBox="[227,1437,454,1927]" captionTargetId="figure@51.[227,1437,454,1927]" captionTargetPageId="51" captionText="Figs 163177. Aetana kinabalu group. Female genitalia; untreated in ventral view, cleared in ventral and dorsal views. — 163165. A. kinabalu Huber, 2005. — 166168. A. lambir Huber, sp. nov. — 169 171. A. gaya Huber, sp. nov. — 172174. A. poring Huber, sp. nov. — 175177. A. indah Huber, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829578" httpUri="https://zenodo.org/record/3829578/files/figure.png" pageId="34" pageNumber="35">163165</figureCitation>
</paragraph>
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<taxonomicName id="4C3922B2601DFFB797FAFCE247EFFCD9" ID-CoL="65BCS" authority="Huber, 2005 a: 75 - 76" authorityName="Huber" authorityPageNumber="75 - 76" authorityYear="2005" box="[227,678,866,893]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523601DFFB797FAFCE244EDFCD8" box="[227,420,866,893]" italics="true" pageId="34" pageNumber="35">Aetana kinabalu</emphasis>
<bibRefCitation id="EFA824C0601DFFB796B2FCE347EFFCD9" author="Huber B. A." box="[427,678,866,893]" pageId="34" pageNumber="35" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber, 2005a: 7576</bibRefCitation>
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, figs 108109, 121123 (
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).
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<heading id="D0CEEE5D601DFFB797FAFC254457FC1A" bold="true" box="[227,286,932,958]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B94D8523601DFFB797FAFC254457FC1A" bold="true" box="[227,286,932,958]" pageId="34" pageNumber="35">Note</emphasis>
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The original description was based on two males. Here we present data on new material from the
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locality, a description of the female, and an emended diagnosis to account for the newly described congeners.
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<emphasis id="B94D8523601DFFB797FAFBDB4415FBD0" bold="true" box="[227,348,1114,1140]" pageId="34" pageNumber="35">Diagnosis</emphasis>
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Distinguished from closest known relative (
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<emphasis id="B94D8523601DFFB795EFFB08462EFB07" box="[758,871,1161,1187]" italics="true" pageId="34" pageNumber="35">A. lambir</emphasis>
Huber
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,
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) by shape of prolatero-ventral apophysis of male palpal femur (pointed tip and subdistal branch of approximately same length;
<emphasis id="B94D8523601DFFB79299FB2D40D4FB62" box="[1408,1437,1196,1222]" italics="true" pageId="34" pageNumber="35">cf.</emphasis>
<bibRefCitation id="EFA824C0601DFFB797FAFB4E44CAFB4E" author="Huber B. A." box="[227,387,1231,1258]" pageId="34" pageNumber="35" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber 2005a</bibRefCitation>
: fig. 121); also by presence of prolateral apophysis on femur, by details of procursus (shapes of sclerites on complex distal part;
<emphasis id="B94D8523601DFFB795CEFB7347A5FAA8" box="[727,748,1266,1292]" italics="true" pageId="34" pageNumber="35">cf</emphasis>
.
<bibRefCitation id="EFA824C0601DFFB795E3FB7346DFFAA9" author="Huber B. A." box="[762,918,1266,1293]" pageId="34" pageNumber="35" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber 2005a</bibRefCitation>
: figs 121122), and by female genitalia (pair of internal pockets; position of pore plates;
<figureCitation id="130245B4601DFFB795FBFA9446CDFA94" box="[738,900,1301,1328]" captionStart="Figs 111112" captionStartId="37.[227,279,1997,2023]" captionTargetBox="[239,1430,1527,1927]" captionTargetId="figure@37.[227,1437,1519,1939]" captionTargetPageId="37" captionText="Figs 111112. Aetana kinabalu Huber, 2005. Cleared female genitalia, ventral and dorsal views (arrow in Fig. 112 points at membranous pocket). ip = internal sclerotized pocket. Scale line: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829560" httpUri="https://zenodo.org/record/3829560/files/figure.png" pageId="34" pageNumber="35">Figs 111112</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA601DFFB797FAFAD64414F86A" pageId="34" pageNumber="35" type="materials_examined">
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<emphasis id="B94D8523601DFFB797FAFAD64745FAD5" bold="true" box="[227,524,1367,1393]" pageId="34" pageNumber="35">New material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B865931601DFFB797FAFA074009F932" blockId="34.[227,1437,1413,1687]" pageId="34" pageNumber="35">
MALAYSIA-BORNEO,
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:
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,
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,
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, Mt. Kinabalu, forest along Silau Silau Trail (
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° N, 116.537
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),
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a.s.l., domed webs among vegetation,
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(B.A. Huber, S.B. Huber), ZFMK (Ar 13961);
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,
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, in pure ethanol, same data, ZFMK (Bor 211).
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, Kinabalu N.P.,
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a.s.l.,
<date id="FF877FF1601DFFB7958DFA6E4674F9AD" box="[660,829,1519,1545]" pageId="34" pageNumber="35" value="1998-04-02" valueMax="1998-04-08" valueMin="1998-04-02">28 Apr. 1998</date>
(C.L. Deeleman-Reinhold, P. Zborowski), RMNH.
<specimenCount id="9D3F92B8601DFFB797E1F9934475F989" box="[248,316,1554,1581]" pageId="34" pageNumber="35" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D3F92B8601DFFB79650F99344C2F989" box="[329,395,1554,1581]" pageId="34" pageNumber="35" type="female">5 ♀♀</specimenCount>
, 3 juvs, Crocker Range between Kota Kinabalu and Tambuan, S-slope, forest along river (
<geoCoordinate id="EE0D3FF6601DFFB797F2F9B44411F9F4" box="[235,344,1589,1616]" degrees="5.783" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="5.783">5.783° N</geoCoordinate>
, 116.338
<geoCoordinate id="EE0D3FF6601DFFB796C8F9B4471DF9F4" box="[465,596,1589,1616]" degrees="116.340" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="55" value="116.34">116.340° E</geoCoordinate>
),
<quantity id="4CC1F4D4601DFFB79571F9B74640F9F4" box="[616,777,1590,1616]" metricMagnitude="3" metricUnit="m" metricValue="1.455" metricValueMax="1.48" metricValueMin="1.43" pageId="34" pageNumber="35" unit="m" value="1455.0" valueMax="1480.0" valueMin="1430.0">14301480 m</quantity>
a.s.l., domed webs among vegetation,
<date id="FF877FF1601DFFB793D1F9B7401CF9F4" box="[1224,1365,1590,1616]" pageId="34" pageNumber="35" value="2014-08-03">3 Aug. 2014</date>
(B.A. Huber, S.B. Huber), ZFMK (Ar 13962); 4 juvs, in pure ethanol, same data, ZFMK (Bor 171).
<specimenCount id="9D3F92B8601DFFB7924BF9D840DEF9D7" box="[1362,1431,1625,1651]" pageId="34" pageNumber="35" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D3F92B8601DFFB797FAF9FD446FF932" box="[227,294,1660,1686]" pageId="34" pageNumber="35" type="female">3 ♀♀</specimenCount>
, in very poor condition, Tawau (
<geoCoordinate id="EE0D3FF6601DFFB795BDF9FD465AF932" box="[676,787,1660,1686]" degrees="4.406" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="4.406">4.406° N</geoCoordinate>
,
<geoCoordinate id="EE0D3FF6601DFFB79406F9FD46ECF932" box="[799,933,1660,1686]" degrees="117.892" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="55" value="117.892">117.892° E</geoCoordinate>
),
<date id="FF877FF1601DFFB794A2F9FD410FF932" box="[955,1094,1660,1686]" pageId="34" pageNumber="35" value="2009-09-06">6 Sep. 2009</date>
(A. Floren), RMNH.
</paragraph>
<paragraph id="8B865931601DFFB797FAF93C4414F86A" blockId="34.[227,1436,1725,1998]" pageId="34" pageNumber="35">
MALAYSIA-BORNEO,
<collectingCountry id="F32E19A1601DFFB79509F93C4733F973" box="[528,634,1725,1751]" name="Malaysia" pageId="34" pageNumber="35">Sarawak</collectingCountry>
:
<specimenCount id="9D3F92B8601DFFB79591F93C4799F97C" box="[648,720,1725,1752]" pageId="34" pageNumber="35" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D3F92B8601DFFB795F8F93C4661F97C" box="[737,808,1725,1752]" pageId="34" pageNumber="35" type="female">6 ♀♀</specimenCount>
, 3 juvs, Gunung Mulu N.P., forest near Deer Cave (
<geoCoordinate id="EE0D3FF6601DFFB797F2F9604413F95F" box="[235,346,1761,1787]" degrees="4.027" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="4.027">4.027° N</geoCoordinate>
,
<geoCoordinate id="EE0D3FF6601DFFB7967FF96044A5F95F" box="[358,492,1761,1787]" degrees="114.818" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="55" value="114.818">114.818° E</geoCoordinate>
),
<quantity id="4CC1F4D4601DFFB7951BF9614774F95F" box="[514,573,1760,1787]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="34" pageNumber="35" unit="m" value="60.0">60 m</quantity>
a.s.l.,
<date id="FF877FF1601DFFB79593F9604607F95F" box="[650,846,1760,1787]" pageId="34" pageNumber="35" value="2014-07-23" valueMax="2014-07-24" valueMin="2014-07-23">2324 July 2014</date>
(B.A. Huber, S.B. Huber), ZFMK (Ar 13963-64); same data,
<specimenCount id="9D3F92B8601DFFB7967EF88544E2F8BA" box="[359,427,1796,1822]" pageId="34" pageNumber="35" type="female">3 ♀♀</specimenCount>
, 5 juvs, in pure ethanol, ZFMK (Bor 182).
<specimenCount id="9D3F92B8601DFFB794DDF8824140F8BA" box="[964,1033,1795,1822]" pageId="34" pageNumber="35" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D3F92B8601DFFB7930FF8854113F8BA" box="[1046,1114,1796,1822]" pageId="34" pageNumber="35" type="female">2 ♀♀</specimenCount>
, 4 juvs, Bario, forest along river W of town (
<geoCoordinate id="EE0D3FF6601DFFB796A8F8A64769F8E5" box="[433,544,1831,1857]" degrees="3.736" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="3.736">3.736° N</geoCoordinate>
, 115.437
<geoCoordinate id="EE0D3FF6601DFFB79585F8A64669F8E5" box="[668,800,1831,1858]" degrees="115.443" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="55" value="115.443">115.443° E</geoCoordinate>
),
<quantity id="4CC1F4D4601DFFB7942EF8A64690F8E5" box="[823,985,1831,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="34" pageNumber="35" unit="m" value="1200.0" valueMax="1250.0" valueMin="1150.0">11501250 m</quantity>
a.s.l., domed webs among vegetation,
<date id="FF877FF1601DFFB797FAF8CB4437F8C0" box="[227,382,1866,1892]" pageId="34" pageNumber="35" value="2014-07-30">30 July 2014</date>
(B.A. Huber, S.B. Huber), ZFMK (Ar 13965).
<specimenCount id="9D3F92B8601DFFB794D6F8CB46B5F8C1" box="[975,1020,1866,1893]" pageId="34" pageNumber="35" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D3F92B8601DFFB79314F8CB411BF8C0" box="[1037,1106,1866,1892]" pageId="34" pageNumber="35" type="generic">1 juv.</specimenCount>
, Bario, forest along river N of town (
<geoCoordinate id="EE0D3FF6601DFFB79648F8EC44DCF82C" box="[337,405,1901,1928]" degrees="3.765" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="3.765">3.765</geoCoordinate>
<geoCoordinate id="EE0D3FF6601DFFB7968CF8EC44A1F82C" box="[405,488,1901,1928]" degrees="3.771" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="55" value="-3.771">
<geoCoordinate id="EE0D3FF6601DFFB796BCF8EC44A1F82C" box="[421,488,1901,1928]" degrees="3.771" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="3.771">3.771</geoCoordinate>
</geoCoordinate>
° N, 115.444
<geoCoordinate id="EE0D3FF6601DFFB79589F8EC465FF82C" box="[656,790,1901,1928]" degrees="115.448" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="55" value="115.448">115.448° E</geoCoordinate>
),
<quantity id="4CC1F4D4601DFFB79437F8EC4698F82C" box="[814,977,1901,1928]" metricMagnitude="3" metricUnit="m" metricValue="1.21" metricValueMax="1.25" metricValueMin="1.17" pageId="34" pageNumber="35" unit="m" value="1210.0" valueMax="1250.0" valueMin="1170.0">11701250 m</quantity>
a.s.l., domed webs among vegetation,
<date id="FF877FF1601DFFB797FAF8104435F80F" box="[227,380,1936,1963]" pageId="34" pageNumber="35" value="2014-07-29">29 July 2014</date>
(B.A. Huber, S.B. Huber), ZFMK (Ar 13966); 9 juvs, in pure ethanol, same data, ZFMK (Bor 233).
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA601CFFB197FAFEB04485FD81" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="8B865931601CFFB697FAFEB047A8FEE8" blockId="35.[227,737,305,332]" box="[227,737,305,332]" pageId="35" pageNumber="36">
<emphasis id="B94D8523601CFFB697FAFEB0443DFEEF" bold="true" box="[227,372,305,331]" pageId="35" pageNumber="36">Description</emphasis>
(amendments to
<bibRefCitation id="EFA824C0601CFFB69526FEB04790FEE8" author="Huber B. A." box="[575,729,305,332]" pageId="35" pageNumber="36" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber 2005a</bibRefCitation>
)
</paragraph>
<paragraph id="8B865931601CFFB69609FEE14419FEDE" blockId="35.[227,1437,352,483]" box="[272,336,352,378]" pageId="35" pageNumber="36">
<emphasis id="B94D8523601CFFB69609FEE14419FEDE" bold="true" box="[272,336,352,378]" pageId="35" pageNumber="36">Male</emphasis>
</paragraph>
<paragraph id="8B865931601CFFB697FAFE024438FE47" blockId="35.[227,1437,352,483]" pageId="35" pageNumber="36">
Thoracic furrow absent (
<emphasis id="B94D8523601CFFB6951BFE054706FE39" box="[514,591,388,413]" italics="true" pageId="35" pageNumber="36">contra</emphasis>
<bibRefCitation id="EFA824C0601CFFB6954DFE0247B9FE3A" author="Huber B. A." box="[596,752,387,414]" pageId="35" pageNumber="36" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber 2005a</bibRefCitation>
); clypeus slightly protruding (more than in female). Tibia 2 slightly longer than tibia 4 (e.g., 5.1/4.9; 4.9/4.8); curved hairs on metatarsi 2; tibia
<quantity id="4CC1F4D4601CFFB693B3FE27419FFE64" box="[1194,1238,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="1.0">1 in</quantity>
<specimenCount id="9D3F92B8601CFFB693C5FE264073FE64" box="[1244,1338,422,449]" pageId="35" pageNumber="36" type="male">7 males</specimenCount>
: 7.68.8 (mean: 8.3).
</paragraph>
<paragraph id="8B865931601CFFB69609FD8A4422FD81" blockId="35.[227,1437,523,620]" box="[272,363,523,549]" pageId="35" pageNumber="36">
<emphasis id="B94D8523601CFFB69609FD8A4422FD81" bold="true" box="[272,363,523,549]" pageId="35" pageNumber="36">Female</emphasis>
</paragraph>
<paragraph id="8B865931601CFFB697FAFDAF40D5FDCF" blockId="35.[227,1437,523,620]" pageId="35" pageNumber="36">
Eye triads much closer together than in male (distance PME-PME ~
<quantity id="4CC1F4D4601CFFB69318FDAF4117FDEC" box="[1025,1118,558,585]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" pageId="35" pageNumber="36" unit="mm" value="235.0">235 µm</quantity>
<emphasis id="B94D8523601CFFB6937CFDAE41CCFDEC" box="[1125,1157,559,584]" italics="true" pageId="35" pageNumber="36">vs.</emphasis>
<quantity id="4CC1F4D4601CFFB69395FDAF41A2FDEC" box="[1164,1259,558,585]" metricMagnitude="-1" metricUnit="m" metricValue="3.75" pageId="35" pageNumber="36" unit="mm" value="375.0">375 µm</quantity>
), not on stalks. Tibia
<quantity id="4CC1F4D4601CFFB69631FDD0441FFDCF" box="[296,342,593,619]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="1.0">1 in</quantity>
<specimenCount id="9D3F92B8601CFFB69647FDD044ADFDCF" box="[350,484,593,620]" pageId="35" pageNumber="36" type="female">15 females</specimenCount>
: 5.97.3 (mean 6.4). Epigynum simple plate, darker laterally anteriorly; anterior
</paragraph>
<caption id="DF4609B9601CFFB697FAF80B447BF7AA" ID-DOI="http://doi.org/10.5281/zenodo.3829555" ID-Zenodo-Dep="3829555" httpUri="https://zenodo.org/record/3829555/files/figure.png" pageId="35" pageNumber="36" startId="35.[227,279,1930,1956]" targetBox="[227,1437,731,1888]" targetPageId="35">
<paragraph id="8B865931601CFFB697FAF80B447BF7AA" blockId="35.[227,1437,1930,2062]" pageId="35" pageNumber="36">
<emphasis id="B94D8523601CFFB697FAF80B4436F800" bold="true" box="[227,383,1930,1956]" pageId="35" pageNumber="36">Figs 99104.</emphasis>
Live specimens.
<taxonomicName id="4C3922B2601CFFB69548F80A465DF800" authorityName="Huber" authorityYear="2005" box="[593,788,1930,1957]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523601CFFB69548F80A465DF800" box="[593,788,1930,1957]" italics="true" pageId="35" pageNumber="36">Aetana kinabalu</emphasis>
</taxonomicName>
group. —
<emphasis id="B94D8523601CFFB6948EF80B46A6F800" bold="true" box="[919,1007,1930,1956]" pageId="35" pageNumber="36">99103</emphasis>
.
<taxonomicName id="4C3922B2601CFFB694E6F80A406EF801" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[1023,1319,1930,1957]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523601CFFB694E6F80A41C0F800" box="[1023,1161,1930,1957]" italics="true" pageId="35" pageNumber="36">A. kinabalu</emphasis>
Huber, 2005
</taxonomicName>
. ♂ and ♀ with eggsac from Gunung Mulu, Sarawak (99, 100); ♀ with eggsac and ♂ from Crocker Range, Sabah (101, 102); and ♂ from Mt. Kinabalu, Sabah (103). —
<emphasis id="B94D8523601CFFB69442F85046CEF84F" bold="true" box="[859,903,2001,2027]" pageId="35" pageNumber="36">104</emphasis>
.
<taxonomicName id="4C3922B2601CFFB6948CF8504177F84F" authority="Huber" authorityName="Huber" authorityYear="2015" box="[917,1086,2001,2027]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="gaya" status="sp. nov.">
<emphasis id="B94D8523601CFFB6948CF85046A4F84F" box="[917,1005,2001,2027]" italics="true" pageId="35" pageNumber="36">A. gaya</emphasis>
Huber
</taxonomicName>
,
<taxonomicNameLabel id="A27E3858601CFFB69351F85041EDF84F" box="[1096,1188,2001,2027]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ from Gaya Island, Sabah.
</paragraph>
</caption>
<paragraph id="8B865931601BFFB197FAFEB040C5FECA" blockId="36.[227,1436,305,367]" pageId="36" pageNumber="37">
internal arc and internal sclerotized pockets visible through cuticle (
<figureCitation id="130245B4601BFFB194F2FEB04119FEEF" box="[1003,1104,305,331]" captionStart="Figs 111112" captionStartId="37.[227,279,1997,2023]" captionTargetBox="[239,1430,1527,1927]" captionTargetId="figure@37.[227,1437,1519,1939]" captionTargetPageId="37" captionText="Figs 111112. Aetana kinabalu Huber, 2005. Cleared female genitalia, ventral and dorsal views (arrow in Fig. 112 points at membranous pocket). ip = internal sclerotized pocket. Scale line: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829560" httpUri="https://zenodo.org/record/3829560/files/figure.png" pageId="36" pageNumber="37">Figs 111</figureCitation>
,
<figureCitation id="130245B4601BFFB19343FEB0418CFEE8" box="[1114,1221,305,332]" captionStart="Figs 163177" captionStartId="51.[227,279,1968,1994]" captionTargetBox="[227,1437,454,1927]" captionTargetId="figure@51.[227,1437,454,1927]" captionTargetPageId="51" captionText="Figs 163177. Aetana kinabalu group. Female genitalia; untreated in ventral view, cleared in ventral and dorsal views. — 163165. A. kinabalu Huber, 2005. — 166168. A. lambir Huber, sp. nov. — 169 171. A. gaya Huber, sp. nov. — 172174. A. poring Huber, sp. nov. — 175177. A. indah Huber, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829578" httpUri="https://zenodo.org/record/3829578/files/figure.png" pageId="36" pageNumber="37">163164</figureCitation>
). Internal genitalia as in
<figureCitation id="130245B4601BFFB19638FED544CFFECA" box="[289,390,340,366]" captionStart="Figs 111112" captionStartId="37.[227,279,1997,2023]" captionTargetBox="[239,1430,1527,1927]" captionTargetId="figure@37.[227,1437,1519,1939]" captionTargetPageId="37" captionText="Figs 111112. Aetana kinabalu Huber, 2005. Cleared female genitalia, ventral and dorsal views (arrow in Fig. 112 points at membranous pocket). ip = internal sclerotized pocket. Scale line: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829560" httpUri="https://zenodo.org/record/3829560/files/figure.png" pageId="36" pageNumber="37">Figs 112</figureCitation>
and
<figureCitation id="130245B4601BFFB196A6FED544A4FECB" box="[447,493,340,367]" captionStart="Figs 163177" captionStartId="51.[227,279,1968,1994]" captionTargetBox="[227,1437,454,1927]" captionTargetId="figure@51.[227,1437,454,1927]" captionTargetPageId="51" captionText="Figs 163177. Aetana kinabalu group. Female genitalia; untreated in ventral view, cleared in ventral and dorsal views. — 163165. A. kinabalu Huber, 2005. — 166168. A. lambir Huber, sp. nov. — 169 171. A. gaya Huber, sp. nov. — 172174. A. poring Huber, sp. nov. — 175177. A. indah Huber, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829578" httpUri="https://zenodo.org/record/3829578/files/figure.png" pageId="36" pageNumber="37">165</figureCitation>
, with pair of sclerotized pockets and more lateral pair of membranous pockets.
</paragraph>
<paragraph id="8B865931601BFFB197FAFE174410FE14" blockId="36.[227,345,406,432]" box="[227,345,406,432]" pageId="36" pageNumber="37">
<heading id="D0CEEE5D601BFFB197FAFE174410FE14" bold="true" box="[227,345,406,432]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B94D8523601BFFB197FAFE174410FE14" bold="true" box="[227,345,406,432]" pageId="36" pageNumber="37">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8B865931601BFFB197FAFE444485FD81" blockId="36.[227,1436,452,549]" pageId="36" pageNumber="37">
In males from Gunung Mulu, the tiny cone-shaped process on the genital bulb (
<emphasis id="B94D8523601BFFB193B8FE4541F7FE7A" box="[1185,1214,452,478]" italics="true" pageId="36" pageNumber="37">cf.</emphasis>
<bibRefCitation id="EFA824C0601BFFB193DFFE45402CFE7B" author="Huber B. A." box="[1222,1381,452,479]" pageId="36" pageNumber="37" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber 2005a</bibRefCitation>
: figs 121 and 122) is absent. Males from Bario with small additional process at basis of prolatero-ventral apophysis of femur.
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA601BFFB097FAFDCD465FFE11" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="8B865931601BFFB197FAFDCD44EDFDC2" blockId="36.[227,420,588,614]" box="[227,420,588,614]" pageId="36" pageNumber="37">
<heading id="D0CEEE5D601BFFB197FAFDCD44EDFDC2" bold="true" box="[227,420,588,614]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B94D8523601BFFB197FAFDCD44EDFDC2" bold="true" box="[227,420,588,614]" pageId="36" pageNumber="37">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="8B865931601BFFB197FAFDFA40D5FD1C" blockId="36.[227,1436,635,696]" pageId="36" pageNumber="37">Webs were in most cases made of two sheets: an upper sheet in which the spider hung, and a lower sheet that had to be removed in order to catch the spider. At most localities, webs were found high among the</paragraph>
<caption id="DF4609B9601BFFB197FAF82F40D7F7AB" ID-DOI="http://doi.org/10.5281/zenodo.3829557" ID-Zenodo-Dep="3829557" httpUri="https://zenodo.org/record/3829557/files/figure.png" pageId="36" pageNumber="37" startId="36.[227,278,1966,1992]" targetBox="[228,1438,741,1921]" targetPageId="36">
<paragraph id="8B865931601BFFB197FAF82F40D7F84F" blockId="36.[227,1438,1966,2028]" pageId="36" pageNumber="37">
<emphasis id="B94D8523601BFFB197FAF82F44C1F86C" bold="true" box="[227,392,1966,1992]" pageId="36" pageNumber="37">Figs 105110.</emphasis>
Live specimens.
<taxonomicName id="4C3922B2601BFFB1954DF82E4658F86C" authorityName="Huber" authorityYear="2005" box="[596,785,1966,1993]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523601BFFB1954DF82E4658F86C" box="[596,785,1966,1993]" italics="true" pageId="36" pageNumber="37">Aetana kinabalu</emphasis>
</taxonomicName>
group. —
<emphasis id="B94D8523601BFFB19488F82F46BFF86C" bold="true" box="[913,1014,1966,1992]" pageId="36" pageNumber="37">105106</emphasis>
.
<taxonomicName id="4C3922B2601BFFB1931CF82E418DF86C" authority="Huber" authorityName="Huber" authorityYear="2015" box="[1029,1220,1966,1993]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lambir" status="sp. nov.">
<emphasis id="B94D8523601BFFB1931CF82E413BF86C" box="[1029,1138,1966,1993]" italics="true" pageId="36" pageNumber="37">A. lambir</emphasis>
Huber
</taxonomicName>
,
<taxonomicNameLabel id="A27E3858601BFFB193C9F82E4065F86D" box="[1232,1324,1967,1993]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ and ♀ with eggsac from Lambir, Sarawak. —
<emphasis id="B94D8523601BFFB195B8F850464FF84F" bold="true" box="[673,774,2001,2027]" pageId="36" pageNumber="37">107108</emphasis>
.
<taxonomicName id="4C3922B2601BFFB1940AF8534687F84F" authority="Huber" authorityName="Huber" authorityYear="2015" box="[787,974,2001,2028]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="poring" status="sp. nov.">
<emphasis id="B94D8523601BFFB1940AF8534637F84F" box="[787,894,2002,2028]" italics="true" pageId="36" pageNumber="37">A. poring</emphasis>
Huber
</taxonomicName>
,
<taxonomicNameLabel id="A27E3858601BFFB194C1F853417BF848" box="[984,1074,2002,2028]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
, ♂♂ from Mt. Kinabalu, Sabah.
</paragraph>
<paragraph id="8B865931601BFFB197FAF87440D7F7AB" blockId="36.[227,256,2037,2063]" box="[227,1438,2037,2063]" lastBlockId="36.[1346,1438,2037,2063]" pageId="36" pageNumber="37">
<emphasis id="B94D8523601BFFB19613F8744431F7AB" bold="true" box="[266,376,2037,2063]" pageId="36" pageNumber="37">109 110</emphasis>
.
<taxonomicName id="4C3922B2601BFFB19690F874470BF7AB" authority="Huber" authorityName="Huber" authorityYear="2015" box="[393,578,2037,2063]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="indah" status="sp. nov.">
<emphasis id="B94D8523601BFFB19690F87444A4F7AB" box="[393,493,2037,2063]" italics="true" pageId="36" pageNumber="37">A. indah</emphasis>
Huber
</taxonomicName>
,
<taxonomicNameLabel id="A27E3858601BFFB19549F87447E7F7AB" box="[592,686,2037,2063]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
, adult and penultimate instar,
<specimenCount id="9D3F92B8601BFFB1930FF8744108F7AB" box="[1046,1089,2037,2063]" pageId="36" pageNumber="37" type="male">♂♂</specimenCount>
from Crocker Range,
<collectingRegion id="49FD97D3601BFFB19249F87440D3F7AB" box="[1360,1434,2037,2063]" country="Malaysia" name="Sabah" pageId="36" pageNumber="37">Sabah</collectingRegion>
.
</paragraph>
</caption>
<paragraph id="8B865931601AFFB097FAFEB0465FFE11" blockId="37.[227,1437,305,437]" pageId="37" pageNumber="38">
vegetation, even in the sunlight. At Gunung Mulu, webs were found among mosses and low vegetation on perpendicular rock surfaces about
<quantity id="4CC1F4D4601AFFB095A8FED547ABFECB" box="[689,738,340,367]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="m" value="2.0">2 m</quantity>
above the ground. At several localities,
<taxonomicName id="4C3922B2601AFFB093CEFED4402AFECA" authorityName="Huber" authorityYear="2005" box="[1239,1379,340,367]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523601AFFB093CEFED4402AFECA" box="[1239,1379,340,367]" italics="true" pageId="37" pageNumber="38">A. kinabalu</emphasis>
</taxonomicName>
was sympatric with a ground-dwelling congener: with
<taxonomicName id="4C3922B2601AFFB09432FEF946A2FE35" authority="Huber" authorityName="Huber" authorityYear="2015" box="[811,1003,375,402]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="poring" status="sp. nov.">
<emphasis id="B94D8523601AFFB09432FEF946D0FE35" box="[811,921,376,402]" italics="true" pageId="37" pageNumber="38">A. poring</emphasis>
Huber
</taxonomicName>
,
<taxonomicNameLabel id="A27E3858601AFFB094EFFEF94119FE36" box="[1014,1104,376,402]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
at Mt. Kinabalu and Poring; with
<taxonomicName id="4C3922B2601AFFB09607FE1A449DFE11" authority="Huber" authorityName="Huber" authorityYear="2015" box="[286,468,411,437]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="indah" status="sp. nov.">
<emphasis id="B94D8523601AFFB09607FE1A44CBFE11" box="[286,386,411,437]" italics="true" pageId="37" pageNumber="38">A. indah</emphasis>
Huber
</taxonomicName>
,
<taxonomicNameLabel id="A27E3858601AFFB096F9FE1A4773FE11" box="[480,570,411,437]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
at Crocker Range.
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA601AFFB097FAFE5D46C0FD81" box="[227,905,476,549]" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8B865931601AFFB097FAFE5D4435FE52" blockId="37.[227,380,476,502]" box="[227,380,476,502]" pageId="37" pageNumber="38">
<heading id="D0CEEE5D601AFFB097FAFE5D4435FE52" bold="true" box="[227,380,476,502]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B94D8523601AFFB097FAFE5D4435FE52" bold="true" box="[227,380,476,502]" pageId="37" pageNumber="38">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B865931601AFFB097FAFD8A46C0FD81" blockId="37.[227,905,522,549]" box="[227,905,522,549]" pageId="37" pageNumber="38">
Widely distributed in
<collectingRegion id="49FD97D3601AFFB096FBFD8A4763FD81" box="[482,554,523,549]" country="Malaysia" name="Sabah" pageId="37" pageNumber="38">Sabah</collectingRegion>
and eastern
<collectingCountry id="F32E19A1601AFFB095A4FD8A466BFD81" box="[701,802,523,549]" name="Malaysia" pageId="37" pageNumber="38">Sarawak</collectingCountry>
(
<figureCitation id="130245B4601AFFB0942BFD8A4633FD81" box="[818,890,522,549]" captionStart="Fig" captionStartId="7.[227,270,2036,2062]" captionTargetBox="[227,1436,710,1988]" captionTargetId="figure@7.[227,1436,711,1987]" captionTargetPageId="7" captionText="Fig. 5. Known distributions of the Aetana kinabalu (Borneo) and A. omayan (Philippines) groups." figureDoi="http://doi.org/10.5281/zenodo.3829519" httpUri="https://zenodo.org/record/3829519/files/figure.png" pageId="37" pageNumber="38">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>