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<document id="D3F3EFAA31F6E521262160DA7D8DBED5" ID-DOI="10.11646/zootaxa.4810.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3938177" ID-ZooBank="16464D12-3BF0-4345-B3CB-B2581E1C37B5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594366343418" checkinUser="plazi" docAuthor="Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. &amp; Wahab, Rodzay Bin Haji Abdul" docDate="2020" docId="0390913BA904FF9DFF0C2721FF65FEAF" docLanguage="en" docName="zootaxa.4810.2.2.pdf" docOrigin="Zootaxa 4810 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Varitrella (Cantotrella) suikei Tan, Japir &amp; Chung 2020, sp. nov." docType="treatment" docVersion="4" lastPageNumber="269" masterDocId="FFA9E943A910FF84FF9B227DFF8EFFC8" masterDocTitle="New taxa of crickets (Orthoptera: Grylloidea: Phaloriinae, Phalangopsinae Itarinae and Podoscirtinae) from Borneo (Brunei Darussalam and Sandakan)" masterLastPageNumber="270" masterPageNumber="244" pageNumber="264" updateTime="1698779676340" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="DA412B95A99F61B70BE4412C77B27047" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C3BC3987980FD68D28A3045FFADA7EDE">
<mods:title id="89E6209A79C89BA0E71437B3F32BCEF9">New taxa of crickets (Orthoptera: Grylloidea: Phaloriinae, Phalangopsinae Itarinae and Podoscirtinae) from Borneo (Brunei Darussalam and Sandakan)</mods:title>
</mods:titleInfo>
<mods:name id="933EF1CFEC0283D9DE508A300C2FB661" type="personal">
<mods:role id="27A01B5FE225CE548F008A9160870B51">
<mods:roleTerm id="660933403C66FA311FE002906CCA039F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="730026247766079AAC9A1FF1521436BB">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="684850A08009A2E35F2A7806BC19EB90">Institut de Systématique, Evolution et Biodiversité, Muséum national d’Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France</mods:affiliation>
</mods:name>
<mods:name id="47BC682E31E3B0E19953169DADBD398A" type="personal">
<mods:role id="D4654231A9F1B5D0E8CB358B0E806A34">
<mods:roleTerm id="BE25774562799A895E2FEE4512477742">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD95996B005065BD23E3333A7C4E7703">Japir, Razy</mods:namePart>
<mods:affiliation id="96F033EAD4D89985DF6CA2D94D464776">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah. Razy. Japir @ sabah. gov. my (RJ)</mods:affiliation>
</mods:name>
<mods:name id="3C2113F35B7107887CEB5287A3F3CE8B" type="personal">
<mods:role id="6F5469566445186681C183DD76C8B455">
<mods:roleTerm id="D89A26E33190B549E89452F9545C9227">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7A83AA4087D73CC4D628D0C587AC2FDD">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="52CC1B4B94B22E4D2BC9A867D48C8EE2">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah. Razy. Japir @ sabah. gov. my (RJ)</mods:affiliation>
</mods:name>
<mods:name id="BD69235D9B1B8214194763577FB730A3" type="personal">
<mods:role id="A19694F78762F4F955B69BBA7D663C7A">
<mods:roleTerm id="09C95CB2E838EADA9367F10B7057E752">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="488C9C814D73EB9218EC5F2F9AACC588">Wahab, Rodzay Bin Haji Abdul</mods:namePart>
<mods:affiliation id="35B2229DB880294425FDAC7501197DC3">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam. rodzay. wahab @ ubd. edu. bn; https: // orcid / org / 0000 - 0002 - 2151 - 7709</mods:affiliation>
</mods:name>
<mods:typeOfResource id="606E859E9DA6F0E7F09AFF28C831A9CE">text</mods:typeOfResource>
<mods:relatedItem id="4C370FAAF07B602393CD4FB042DA90B6" type="host">
<mods:titleInfo id="EF7A4AB3D9503DE43ED25871403DCBA3">
<mods:title id="B490702AFC5D5D52509BD8CEE6A3D53B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2E8536AEAEEF84FFD0E97D19D02AF98A">
<mods:date id="8C7F788C11F86E77E57581492D3E1AB3">2020</mods:date>
<mods:detail id="97F8F623CCAF149376E830C1683344EA" type="pubDate">
<mods:number id="B30F077C0D21312D368FB97F24821A7E">2020-07-10</mods:number>
</mods:detail>
<mods:detail id="DD9468AB87ED7D1B9B15657D7B74B6EB" type="volume">
<mods:number id="40AA6CC66B84FF9137B13129074AC645">4810</mods:number>
</mods:detail>
<mods:detail id="2211CB8A9EA57C4F0688E229A5213471" type="issue">
<mods:number id="697DEDFD2CF3DA35DC3C35DDD52805F1">2</mods:number>
</mods:detail>
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<mods:start id="073301E49D1FF0704FC0F09476CB163A">244</mods:start>
<mods:end id="C3CCA46F87E5C4CC30F087971CB69A63">270</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="47067BA04BD4281DC42C4B2A0414F8E9">journal article</mods:classification>
<mods:identifier id="5B637ED3BDD7484C00BB7896E5414351" type="DOI">10.11646/zootaxa.4810.2.2</mods:identifier>
<mods:identifier id="391F1EA3B89A45B5628986F300E4A7B1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="048EE8711EA51F634E3BC154B8541AC6" type="Zenodo-Dep">3938177</mods:identifier>
<mods:identifier id="3E50546E76D295D02B51CD43A59DFAB1" type="ZooBank">16464D12-3BF0-4345-B3CB-B2581E1C37B5</mods:identifier>
</mods:mods>
<treatment id="0390913BA904FF9DFF0C2721FF65FEAF" ID-DOI="http://doi.org/10.5281/zenodo.4332665" ID-Zenodo-Dep="4332665" LSID="urn:lsid:plazi:treatment:0390913BA904FF9DFF0C2721FF65FEAF" httpUri="http://treatment.plazi.org/id/0390913BA904FF9DFF0C2721FF65FEAF" lastPageId="25" lastPageNumber="269" pageId="20" pageNumber="264">
<subSubSection id="C32373A6A904FF90FF0C2721FCE2FABE" box="[151,876,1372,1399]" pageId="20" pageNumber="264" type="nomenclature">
<paragraph id="8B86202DA904FF90FF0C2721FCE2FABE" blockId="20.[151,876,1372,1434]" box="[151,876,1372,1399]" pageId="20" pageNumber="264">
<heading id="D0CE9741A904FF90FF0C2721FCE2FABE" bold="true" box="[151,876,1372,1399]" fontSize="11" level="1" pageId="20" pageNumber="264" reason="1">
<emphasis id="B94DFC3FA904FF90FF0C2721FCE2FABE" bold="true" box="[151,876,1372,1399]" pageId="20" pageNumber="264">
<taxonomicName id="4C395BAEA904FF90FF0C2721FC8FFABE" authority="Tan, Japir &amp; Chung" authorityName="Tan, Japir &amp; Chung" authorityYear="2020" box="[151,769,1372,1399]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="suikei" status="sp. nov." subGenus="Cantotrella">
<emphasis id="B94DFC3FA904FF90FF0C2721FE87FABF" bold="true" box="[151,265,1372,1399]" italics="true" pageId="20" pageNumber="264">Varitrella</emphasis>
(
<emphasis id="B94DFC3FA904FF90FE812721FE2DFABF" bold="true" box="[282,419,1372,1399]" italics="true" pageId="20" pageNumber="264">Cantotrella</emphasis>
)
<emphasis id="B94DFC3FA904FF90FE2F2721FE72FABF" bold="true" box="[436,508,1372,1399]" italics="true" pageId="20" pageNumber="264">suikei</emphasis>
Tan, Japir &amp; Chung
</taxonomicName>
,
<taxonomicNameLabel id="A27E4144A904FF90FC952721FCE2FABE" box="[782,876,1372,1398]" pageId="20" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32373A6A904FF90FF0C27FDFEA0FA52" box="[151,302,1407,1434]" pageId="20" pageNumber="264" type="description">
<paragraph id="8B86202DA904FF90FF0C27FDFEA0FA52" blockId="20.[151,876,1372,1434]" box="[151,302,1407,1434]" pageId="20" pageNumber="264">
(
<figureCitation id="13023CA8A904FF90FF0427FDFEABFA52" box="[159,293,1407,1434]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStart-3="FIGURE 18" captionStartId-0="21.[151,250,1916,1941]" captionStartId-1="22.[151,250,1945,1970]" captionStartId-2="23.[151,250,1233,1258]" captionStartId-3="24.[151,250,1932,1957]" captionTargetBox-0="[151,1436,561,1892]" captionTargetBox-1="[179,1408,181,1920]" captionTargetBox-2="[151,1436,181,1207]" captionTargetBox-3="[218,1374,326,1880]" captionTargetId-0="figure@21.[151,1436,561,1892]" captionTargetId-1="figure@22.[179,1408,181,1920]" captionTargetId-2="figure@23.[151,1436,181,1207]" captionTargetId-3="figure@24.[214,1374,309,1907]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="FIGURE 15. Varitrella (Cantotrella) suikei sp. nov. male holotype in natural environment (A), habitus in lateral (B) view. Scale bar: 5 mm." captionText-1="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." captionText-2="FIGURE 17. Varitrella (Cantotrella) suikei sp. nov. male holotype genitalia in dorsal (A), ventral (B) and lateral (C) views." captionText-3="FIGURE 18. Varitrella (Cantotrella) suikei sp. nov. male holotype’s calling song: oscillograms (A, C) and spectrogram (B) of four echemes (A, B) and one echeme (C) and power spectrum of one echeme (D)." figureDoi-0="http://doi.org/10.5281/zenodo.3938209" figureDoi-1="http://doi.org/10.5281/zenodo.3938211" figureDoi-2="http://doi.org/10.5281/zenodo.3938213" figureDoi-3="http://doi.org/10.5281/zenodo.3938215" httpUri-0="https://zenodo.org/record/3938209/files/figure.png" httpUri-1="https://zenodo.org/record/3938211/files/figure.png" httpUri-2="https://zenodo.org/record/3938213/files/figure.png" httpUri-3="https://zenodo.org/record/3938215/files/figure.png" pageId="20" pageNumber="264">Figs. 15–18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32373A6A904FF90FF0C27BAFF68F9E2" pageId="20" pageNumber="264" type="materials_examined">
<paragraph id="8B86202DA904FF90FF0C27BAFF68F9E2" blockId="20.[151,1437,1479,2011]" pageId="20" pageNumber="264">
<emphasis id="B94DFC3FA904FF90FF0C27BAFEF1FA2A" bold="true" box="[151,383,1479,1506]" pageId="20" pageNumber="264">Material examined.</emphasis>
<materialsCitation id="3B512A70A904FF90FE1E27B5FF6CF9E2" collectingDate="2019-10-01" collectionCode="FRC" collectorName="M. K. Tan &amp; Tan, R &amp; J. Lee Yukang" country="Malaysia" county="Male" elevation="8" latitude="5.86985" location="Sandakan" longLatPrecision="1" longitude="117.93805" municipality="East" pageId="20" pageNumber="264" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<collectingCounty id="62E758A1A904FF90FE1E27B5FE31FA2A" box="[389,447,1480,1506]" pageId="20" pageNumber="264">Male</collectingCounty>
<typeStatus id="54829E8FA904FF90FE5F27B5FDA8FA2A" box="[452,550,1480,1506]" pageId="20" pageNumber="264" type="holotype">holotype</typeStatus>
(SDK.19.78),
<emphasis id="B94DFC3FA904FF90FD5227B5FCFFFA2A" bold="true" box="[713,881,1479,1506]" pageId="20" pageNumber="264">
<collectingMunicipality id="6BE2BA57A904FF90FD5227B5FD70FA2A" box="[713,766,1480,1506]" pageId="20" pageNumber="264">East</collectingMunicipality>
<collectingCountry id="F32E60BDA904FF90FC9F27BAFCFFFA2A" box="[772,881,1479,1506]" name="Malaysia" pageId="20" pageNumber="264">Malaysia</collectingCountry>
</emphasis>
,
<collectingRegion id="49FDEECFA904FF90FCE527B5FC4BFA2A" box="[894,965,1480,1506]" country="Malaysia" name="Sabah" pageId="20" pageNumber="264">Sabah</collectingRegion>
,
<location id="8EE676F6A904FF90FC5427B5FBB1FA2A" LSID="urn:lsid:plazi:treatment:0390913BA904FF9DFF0C2721FF65FEAF:8EE676F6A904FF90FC5427B5FBB1FA2A" box="[975,1087,1480,1506]" country="Malaysia" county="Male" latitude="5.86985" longLatPrecision="1" longitude="117.93805" municipality="East" name="Sandakan" pageId="20" pageNumber="264" stateProvince="Sabah">Sandakan</location>
,
<location id="8EE676F6A904FF90FBD327B5FA16FA2B" LSID="urn:lsid:plazi:treatment:0390913BA904FF9DFF0C2721FF65FEAF:8EE676F6A904FF90FBD327B5FA16FA2B" box="[1096,1432,1480,1507]" country="Malaysia" county="Male" latitude="5.86985" longLatPrecision="1" longitude="117.93805" municipality="East" name="Kabili Sepilok Forest Reserve" pageId="20" pageNumber="264" stateProvince="Sabah">Kabili Sepilok Forest Reserve</location>
,
<geoCoordinate id="EE0D46EAA904FF90FF0C2796FE87F9CE" box="[151,265,1515,1542]" degrees="5.86985" direction="north" orientation="latitude" pageId="20" pageNumber="264" precision="1" value="5.86985">N5.86985</geoCoordinate>
,
<geoCoordinate id="EE0D46EAA904FF90FE8D2796FE2EF9CE" box="[278,416,1515,1542]" degrees="117.93805" direction="east" orientation="longitude" pageId="20" pageNumber="264" precision="1" value="117.93805">E117.93805</geoCoordinate>
, 69.8±
<quantity id="4CC18DC8A904FF90FE762791FDB8F9CE" box="[493,566,1516,1542]" metricMagnitude="0" metricUnit="m" metricValue="7.7" pageId="20" pageNumber="264" unit="m" value="7.7">
<elevation id="0014C71EA904FF90FE762791FDB8F9CE" box="[493,566,1516,1542]" metricMagnitude="0" metricUnit="m" metricValue="7.7" pageId="20" pageNumber="264" unit="m" value="7.7">7.7 m</elevation>
</quantity>
.a.s.l., on foliage,
<date id="FF8706EDA904FF90FD602791FC3EF9CE" box="[763,944,1516,1542]" pageId="20" pageNumber="264" value="2019-10-01">
<collectingDate id="EFC3FF05A904FF90FD602791FC3EF9CE" box="[763,944,1516,1542]" pageId="20" pageNumber="264" value="2019-10-01">1 October 2019</collectingDate>
</date>
, coll.
<collectorName id="26CC45FBA904FF90FC6D2790FBE0F9CF" box="[1014,1134,1517,1543]" pageId="20" pageNumber="264">M. K. Tan</collectorName>
, R. Japir &amp;
<collectorName id="26CC45FBA904FF90FB602790FA13F9CE" box="[1275,1437,1516,1543]" pageId="20" pageNumber="264">J. Lee Yukang</collectorName>
(
<collectionCode id="ED28B8E8A904FF90FF04246DFF54F9E2" box="[159,218,1552,1578]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="20" pageNumber="264">FRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32373A6A904FF90FF5C244EFACAF87F" pageId="20" pageNumber="264" type="diagnosis">
<paragraph id="8B86202DA904FF90FF5C244EFD24F95E" blockId="20.[151,1437,1479,2011]" pageId="20" pageNumber="264">
<emphasis id="B94DFC3FA904FF90FF5C244EFECCF986" bold="true" box="[199,322,1587,1614]" pageId="20" pageNumber="264">Diagnosis.</emphasis>
This new species differs from congeners by male genitalia with dorso-anterior epiphallic spine more laterally pointed; ectoparameres stout and strongly curved with apex acute; and rachis surpassing lateral lobe of epiphallus and with apex strongly emarginated.
</paragraph>
<paragraph id="8B86202DA904FF90FF5C24E2FACAF87F" blockId="20.[151,1437,1479,2011]" pageId="20" pageNumber="264">
<emphasis id="B94DFC3FA904FF90FF5C24E2FDACF972" bold="true" box="[199,546,1695,1722]" pageId="20" pageNumber="264">Comparison with congeners.</emphasis>
This new species resembles Bornean congeners but differs by dorso-anterior epiphallic spine (ds) more laterally pointed than
<taxonomicName id="4C395BAEA904FF90FD2D24B8FCBBF916" baseAuthorityName="Chopard" baseAuthorityYear="1930" box="[694,821,1732,1758]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="B94DFC3FA904FF90FD2D24B8FCBBF916" box="[694,821,1732,1758]" italics="true" pageId="20" pageNumber="264">V. mjobergi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C395BAEA904FF90FCD824B8FC26F916" authorityName="Gorochov" authorityYear="2014" box="[835,936,1733,1758]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="B94DFC3FA904FF90FCD824B8FC26F916" box="[835,936,1733,1758]" italics="true" pageId="20" pageNumber="264">V. striata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C395BAEA904FF90FC4524B8FBD3F916" authorityName="Gorochov" authorityYear="2014" box="[990,1117,1732,1758]" class="Insecta" family="Haglotettigoniidae" genus="Terrozacla" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="trusmadi">
<emphasis id="B94DFC3FA904FF90FC4524B8FBD3F916" box="[990,1117,1732,1758]" italics="true" pageId="20" pageNumber="264">V. trusmadi</emphasis>
</taxonomicName>
, appearing shorter in profile (instead of absent in
<taxonomicName id="4C395BAEA904FF90FEE52494FE71F8CA" baseAuthorityName="Haan" baseAuthorityYear="1844" box="[382,511,1768,1794]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="B94DFC3FA904FF90FEE52494FE71F8CA" box="[382,511,1768,1794]" italics="true" pageId="20" pageNumber="264">V. quadrata</emphasis>
</taxonomicName>
); ectoparameres (ec) stouter and more strongly curved in profile than
<taxonomicName id="4C395BAEA904FF90FA912494FAE3F8CA" authorityName="Gorochov" authorityYear="2014" box="[1290,1389,1769,1794]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="B94DFC3FA904FF90FA912494FAE3F8CA" box="[1290,1389,1769,1794]" italics="true" pageId="20" pageNumber="264">V. striata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C395BAEA904FF90FF0C2570FE98F8EE" authorityName="Gorochov" authorityYear="2014" box="[151,278,1804,1830]" class="Insecta" family="Haglotettigoniidae" genus="Terrozacla" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="trusmadi">
<emphasis id="B94DFC3FA904FF90FF0C2570FE98F8EE" box="[151,278,1804,1830]" italics="true" pageId="20" pageNumber="264">V. trusmadi</emphasis>
</taxonomicName>
; rachis (ra) more strongly emarginated at apex than
<taxonomicName id="4C395BAEA904FF90FCCE2570FC36F8EE" authorityName="Gorochov" authorityYear="2014" box="[853,952,1805,1830]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="B94DFC3FA904FF90FCCE2570FC36F8EE" box="[853,952,1805,1830]" italics="true" pageId="20" pageNumber="264">V. striata</emphasis>
</taxonomicName>
(instead of truncated to faintly emarginated in
<taxonomicName id="4C395BAEA904FF90FF2F254CFEA7F882" authorityName="Gorochov" authorityYear="2014" box="[180,297,1841,1866]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="amoena">
<emphasis id="B94DFC3FA904FF90FF2F254CFEA7F882" box="[180,297,1841,1866]" italics="true" pageId="20" pageNumber="264">V. amoena</emphasis>
</taxonomicName>
,
<taxonomicName id="4C395BAEA904FF90FEAC254CFE1AF882" authorityName="Gorochov" authorityYear="2014" box="[311,404,1840,1866]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="sukau">
<emphasis id="B94DFC3FA904FF90FEAC254CFE1AF882" box="[311,404,1840,1866]" italics="true" pageId="20" pageNumber="264">V. sukau</emphasis>
</taxonomicName>
,
<taxonomicName id="4C395BAEA904FF90FE38254CFD8DF882" authorityName="Gorochov" authorityYear="2014" box="[419,515,1841,1866]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="tawau">
<emphasis id="B94DFC3FA904FF90FE38254CFD8DF882" box="[419,515,1841,1866]" italics="true" pageId="20" pageNumber="264">V. tawau</emphasis>
</taxonomicName>
and
<taxonomicName id="4C395BAEA904FF90FDA0254CFD34F882" authorityName="Gorochov" authorityYear="2014" box="[571,698,1840,1866]" class="Insecta" family="Haglotettigoniidae" genus="Terrozacla" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="trusmadi">
<emphasis id="B94DFC3FA904FF90FDA0254CFD34F882" box="[571,698,1840,1866]" italics="true" pageId="20" pageNumber="264">V. trusmadi</emphasis>
</taxonomicName>
). It also differs from
<taxonomicName id="4C395BAEA904FF90FC36254CFBA1F882" authorityName="Gorochov" authorityYear="2014" box="[941,1071,1840,1866]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="manukan">
<emphasis id="B94DFC3FA904FF90FC36254CFBA1F882" box="[941,1071,1840,1866]" italics="true" pageId="20" pageNumber="264">V. manukan</emphasis>
</taxonomicName>
by presence of dorsal epiphallic spine (ds), lateral apical lobe of epiphallus (al) not pointing dorsad at apex, ectoparameres (ec) curved and pointing ventrad posteriorly (instead of dorsally) and rachis (ra) with apical emargination wider. The new species also differs from
<taxonomicName id="4C395BAEA904FF90FF4E25E0FEBFF87E" authorityName="Gorochov" authorityYear="2014" box="[213,305,1948,1974]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="264" phylum="Arthropoda" rank="species" species="sukau">
<emphasis id="B94DFC3FA904FF90FF4E25E0FEBFF87E" box="[213,305,1948,1974]" italics="true" pageId="20" pageNumber="264">V. sukau</emphasis>
</taxonomicName>
from Sandakan by distinctly shorter dorsal epiphallic spine (ds) and shape of ectoparameres.
</paragraph>
</subSubSection>
<subSubSection id="C32373A6A904FF9CFF5C25C2FEF7FED7" lastPageId="24" lastPageNumber="268" pageId="20" pageNumber="264" type="description">
<paragraph id="8B86202DA904FF91FF5C25C2FE9DFDD3" blockId="20.[151,1437,1479,2011]" lastBlockId="21.[151,1437,153,539]" lastPageId="21" lastPageNumber="265" pageId="20" pageNumber="264">
<emphasis id="B94DFC3FA904FF90FF5C25C2FED7F812" bold="true" box="[199,345,1983,2010]" pageId="20" pageNumber="264">Description.</emphasis>
Habitus very typical of the genus, medium sized (
<figureCitation id="13023CA8A904FF90FC0E25BDFC67F812" box="[917,1001,1983,2010]" captionStart="FIGURE 15" captionStartId="21.[151,250,1916,1941]" captionTargetBox="[151,1436,561,1892]" captionTargetId="figure@21.[151,1436,561,1892]" captionTargetPageId="21" captionText="FIGURE 15. Varitrella (Cantotrella) suikei sp. nov. male holotype in natural environment (A), habitus in lateral (B) view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3938209" httpUri="https://zenodo.org/record/3938209/files/figure.png" pageId="20" pageNumber="264">Fig. 15</figureCitation>
). Head with dorsum slightly flattened, very finely pubescent; rostrum 1.1 times as wide as scapus, with apex truncated (in dorsal view) (
<figureCitation id="13023CA8A905FF91FB4E22E4FAB3FF7C" box="[1237,1341,153,180]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="21" pageNumber="265">Fig. 16A</figureCitation>
); scapus large and elongated; maxillary palpi with apical segment longest, cylindrical and oblong with obtuse apex, with subapical segment shorter than third segment (
<figureCitation id="13023CA8A905FF91FDE4229CFD68FF34" box="[639,742,225,252]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="21" pageNumber="265">Fig. 16B</figureCitation>
); eyes projected anteriorly in dorsal view (
<figureCitation id="13023CA8A905FF91FB50229CFABAFF34" box="[1227,1332,225,252]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="21" pageNumber="265">Fig. 16A</figureCitation>
); median ocellus round and small; lateral ocelli oblong and large, located near eyes. Pronotal disc 1.4 times as wide as long, faintly widening posteriorly (posterior margin 1.2 times as wide as anterior margin); finely and densely pubescent; anterior margin of disc distinctly concave in middle; posterior margin of disc nearly straight to slightly convex (
<figureCitation id="13023CA8A905FF91FAF42330FF45FE44" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="21" pageNumber="265">Fig. 16A</figureCitation>
). Pronotal lateral lobe 1.4 times as long as high, with ventral margin straight (
<figureCitation id="13023CA8A905FF91FBBA230CFB09FE44" box="[1057,1159,369,396]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="21" pageNumber="265">Fig. 16B</figureCitation>
). Metanotal gland absent (
<figureCitation id="13023CA8A905FF91FF3B23E8FE88FE78" box="[160,262,405,432]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="21" pageNumber="265">Fig. 16C</figureCitation>
). Fore tibia slightly swollen, with inner tympanum slit-like, with outer tympanum open and having elongate, oval tympanal membrane; hind tibiae with 7 inner and 5 outer long spines in distal half, 2–6 denticles spines, with small denticles before most proximal spines, and with inner spurs (apical spines) longer than corresponding outer ones.
</paragraph>
<caption id="DF4670A5A905FF91FF0C2501FE89F871" ID-DOI="http://doi.org/10.5281/zenodo.3938209" ID-Zenodo-Dep="3938209" httpUri="https://zenodo.org/record/3938209/files/figure.png" pageId="21" pageNumber="265" startId="21.[151,250,1916,1941]" targetBox="[151,1436,561,1892]" targetPageId="21">
<paragraph id="8B86202DA905FF91FF0C2501FE89F871" blockId="21.[151,1436,1916,1977]" pageId="21" pageNumber="265">
<emphasis id="B94DFC3FA905FF91FF0C2501FE91F85D" bold="true" box="[151,287,1916,1941]" pageId="21" pageNumber="265">FIGURE 15.</emphasis>
<taxonomicName id="4C395BAEA905FF91FEBF2500FDC1F85D" authority="Tan &amp; Japir &amp; Chung &amp; Wahab, 2020" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab" authorityYear="2020" box="[292,591,1917,1941]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="265" phylum="Arthropoda" rank="species" species="suikei" status="sp. nov." subGenus="Cantotrella">
<emphasis id="B94DFC3FA905FF91FEBF2500FE0AF85D" box="[292,388,1917,1941]" italics="true" pageId="21" pageNumber="265">Varitrella</emphasis>
(
<emphasis id="B94DFC3FA905FF91FE092500FD88F85D" box="[402,518,1917,1941]" italics="true" pageId="21" pageNumber="265">Cantotrella</emphasis>
)
<emphasis id="B94DFC3FA905FF91FD8F2500FDC1F85D" box="[532,591,1917,1941]" italics="true" pageId="21" pageNumber="265">suikei</emphasis>
</taxonomicName>
<emphasis id="B94DFC3FA905FF91FDCF2500FD2AF85D" bold="true" box="[596,676,1917,1941]" pageId="21" pageNumber="265">
<taxonomicNameLabel id="A27E4144A905FF91FDCF2500FD2AF85D" box="[596,676,1917,1941]" pageId="21" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype in natural environment (A), habitus in lateral (B) view. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DF4670A5A906FF92FF0C25E4FE09F832" ID-DOI="http://doi.org/10.5281/zenodo.3938211" ID-Zenodo-Dep="3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="22" pageNumber="266" startId="22.[151,250,1945,1970]" targetBox="[179,1408,181,1920]" targetPageId="22">
<paragraph id="8B86202DA906FF92FF0C25E4FE09F832" blockId="22.[151,1436,1945,2042]" pageId="22" pageNumber="266">
<emphasis id="B94DFC3FA906FF92FF0C25E4FEACF87A" bold="true" box="[151,290,1945,1970]" pageId="22" pageNumber="266">FIGURE 16.</emphasis>
<taxonomicName id="4C395BAEA906FF92FEB125E7FDD4F87A" authority="Tan &amp; Japir &amp; Chung &amp; Wahab, 2020" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab" authorityYear="2020" box="[298,602,1946,1970]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="22" pageNumber="266" phylum="Arthropoda" rank="species" species="suikei" status="sp. nov." subGenus="Cantotrella">
<emphasis id="B94DFC3FA906FF92FEB125E7FE04F87A" box="[298,394,1946,1970]" italics="true" pageId="22" pageNumber="266">Varitrella</emphasis>
(
<emphasis id="B94DFC3FA906FF92FE0025E7FD81F87A" box="[411,527,1946,1970]" italics="true" pageId="22" pageNumber="266">Cantotrella</emphasis>
)
<emphasis id="B94DFC3FA906FF92FD8425E7FDD4F87A" box="[543,602,1946,1970]" italics="true" pageId="22" pageNumber="266">suikei</emphasis>
</taxonomicName>
<emphasis id="B94DFC3FA906FF92FDF825E7FD3BF87A" bold="true" box="[611,693,1946,1970]" pageId="22" pageNumber="266">
<taxonomicNameLabel id="A27E4144A906FF92FDF825E7FD3BF87A" box="[611,693,1946,1970]" pageId="22" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining).
</paragraph>
</caption>
<caption id="DF4670A5A907FF93FF0C26ACFA0CFB22" ID-DOI="http://doi.org/10.5281/zenodo.3938213" ID-Zenodo-Dep="3938213" box="[151,1410,1233,1258]" httpUri="https://zenodo.org/record/3938213/files/figure.png" pageId="23" pageNumber="267" startId="23.[151,250,1233,1258]" targetBox="[151,1436,181,1207]" targetPageId="23">
<paragraph id="8B86202DA907FF93FF0C26ACFA0CFB22" blockId="23.[151,1410,1233,1258]" box="[151,1410,1233,1258]" pageId="23" pageNumber="267">
<emphasis id="B94DFC3FA907FF93FF0C26ACFEAEFB22" bold="true" box="[151,288,1233,1258]" pageId="23" pageNumber="267">FIGURE 17.</emphasis>
<taxonomicName id="4C395BAEA907FF93FEBD26AFFDDDFB22" authority="Tan &amp; Japir &amp; Chung &amp; Wahab, 2020" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab" authorityYear="2020" box="[294,595,1234,1258]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="23" pageNumber="267" phylum="Arthropoda" rank="species" species="suikei" status="sp. nov." subGenus="Cantotrella">
<emphasis id="B94DFC3FA907FF93FEBD26AFFE08FB22" box="[294,390,1234,1258]" italics="true" pageId="23" pageNumber="267">Varitrella</emphasis>
(
<emphasis id="B94DFC3FA907FF93FE0E26AFFD87FB22" box="[405,521,1234,1258]" italics="true" pageId="23" pageNumber="267">Cantotrella</emphasis>
)
<emphasis id="B94DFC3FA907FF93FD8326AFFDDDFB22" box="[536,595,1234,1258]" italics="true" pageId="23" pageNumber="267">suikei</emphasis>
</taxonomicName>
<emphasis id="B94DFC3FA907FF93FDC226AFFD24FB22" bold="true" box="[601,682,1234,1258]" pageId="23" pageNumber="267">
<taxonomicNameLabel id="A27E4144A907FF93FDC226AFFD24FB22" box="[601,682,1234,1258]" pageId="23" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype genitalia in dorsal (A), ventral (B) and lateral (C) views.
</paragraph>
</caption>
<paragraph id="8B86202DA907FF93FF5C2765FE47FAB2" blockId="23.[151,1437,1303,2015]" pageId="23" pageNumber="267">
Tegmen (
<figureCitation id="13023CA8A907FF93FEB42765FE1AFAFB" box="[303,404,1304,1331]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="23" pageNumber="267">Fig. 16D</figureCitation>
) covering abdomen and surpassing apex of hind femur; with mirror 1.5 times as long as wide, dividing vein typical of subgenus; with 5 veins in harp area; with around 16 branches on Sc (
<figureCitation id="13023CA8A907FF93FB3F2741FA83FA9F" box="[1188,1293,1340,1367]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="23" pageNumber="267">Fig. 16D</figureCitation>
); hind wings clearly surpassing tegmina.
</paragraph>
<paragraph id="8B86202DA907FF93FF5C27F9FEF7FA0A" blockId="23.[151,1437,1303,2015]" pageId="23" pageNumber="267">
Anal plate broad, with two whitish oval parts at base, slightly indented at apex (
<figureCitation id="13023CA8A907FF93FBFC27F9FB41FA57" box="[1127,1231,1412,1439]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="23" pageNumber="267">Fig. 16E</figureCitation>
). Subgenital plate typical of subgenus.
</paragraph>
<paragraph id="8B86202DA907FF93FF5C27B1FBBAF92A" blockId="23.[151,1437,1303,2015]" pageId="23" pageNumber="267">
Male genitalia as shown in
<figureCitation id="13023CA8A907FF93FE6227B1FDC2FA2E" box="[505,588,1484,1510]" captionStart="FIGURE 17" captionStartId="23.[151,250,1233,1258]" captionTargetBox="[151,1436,181,1207]" captionTargetId="figure@23.[151,1436,181,1207]" captionTargetPageId="23" captionText="FIGURE 17. Varitrella (Cantotrella) suikei sp. nov. male holotype genitalia in dorsal (A), ventral (B) and lateral (C) views." figureDoi="http://doi.org/10.5281/zenodo.3938213" httpUri="https://zenodo.org/record/3938213/files/figure.png" pageId="23" pageNumber="267">Fig. 17</figureCitation>
: Epiphallus in dorsal view with dorso-anterior epiphallic spine (ds) strongly sclerotised, somewhat laterally pointed, lateral margin concave, posterior end produced into two latero-apical lobular parts (al), deeply V-emarginated between latero-apical lobules; in profile with posterior part rounded with a dorsoapical spinule and two smaller spinules on dorsal margin, dorsal margin straight. Ectoparameres (ec) hooked-like, curved tapering into an acute apex, and pointing ventrad posteriorly. Rachis (ra, guiding rod) rather long, but not surpassing epiphallus, produced into two lateral lobes (lo) at posterior end, with apical emargination wider between lateral lobes in dorsal view; lateral lobe (lo) strongly widened and roundly rectangular in profile. Endoparameral apodemes (ae) very long, narrowing and converging towards each other anteriorly.
</paragraph>
<paragraph id="8B86202DA907FF93FF5C2491FE10F8CE" blockId="23.[151,1437,1303,2015]" box="[199,414,1772,1798]" pageId="23" pageNumber="267">Female. Unknown.</paragraph>
<paragraph id="8B86202DA907FF9CFF5C256DFAC5FF1F" blockId="23.[151,1437,1303,2015]" lastBlockId="24.[151,1436,153,287]" lastPageId="24" lastPageNumber="268" pageId="23" pageNumber="267">
Colouration. Body generally grey-brown (
<figureCitation id="13023CA8A907FF93FD39256DFD78F8E2" box="[674,758,1807,1834]" captionStart="FIGURE 15" captionStartId="21.[151,250,1916,1941]" captionTargetBox="[151,1436,561,1892]" captionTargetId="figure@21.[151,1436,561,1892]" captionTargetPageId="21" captionText="FIGURE 15. Varitrella (Cantotrella) suikei sp. nov. male holotype in natural environment (A), habitus in lateral (B) view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3938209" httpUri="https://zenodo.org/record/3938209/files/figure.png" pageId="23" pageNumber="267">Fig. 15</figureCitation>
). Head and pronotum with few dark spots (more in pronotal lateral lobe), but not as plentiful as some congeners (
<figureCitation id="13023CA8A907FF93FD7B2549FC00F887" box="[736,910,1844,1871]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="23" pageNumber="267">Figs. 16A, 16B</figureCitation>
); scapus and pedicel slightly lighter in colouration; maxillary palps with apical segment dark distally. Tegminal dorsal field yellowish, with some darker infumation but not obvious, veins range from yellow to dark; with a dark patch where the stridulatory file, diagonal and chords converge and a dark strip between mirror and CuA (
<figureCitation id="13023CA8A907FF93FCB525DDFC19F873" box="[814,919,1952,1979]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="23" pageNumber="267">Fig. 16D</figureCitation>
). Lateral tegminal field paler, almost transparent with darker brown veins, with small black spots near Sc between branches of Sc (
<figureCitation id="13023CA8A907FF93FBC225B9FB31F817" box="[1113,1215,1988,2015]" captionStart="FIGURE 16" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure@22.[179,1408,181,1920]" captionTargetPageId="22" captionText="FIGURE 16. Varitrella (Cantotrella) suikei sp. nov. male holotype: head and pronotum in dorsal (A) and lateral (B) views, metanotal area in dorsal view (C), right tegmen in dorsal view (D) and abdominal apex in dorsal view (E). Scale bars: 5 mm (D), 1 mm (remaining)." figureDoi="http://doi.org/10.5281/zenodo.3938211" httpUri="https://zenodo.org/record/3938211/files/figure.png" pageId="23" pageNumber="267">Fig. 16E</figureCitation>
). Hind wings trans- parent. Legs generally pale brown with irregular and sparse dark spots. Thoracic and abdominal segments yellow brown to brown; anal plate brown with two whitish oval parts at base and another whitish spot before apex.
</paragraph>
<paragraph id="8B86202DA908FF9CFF5C229CFEF7FED7" blockId="24.[151,1436,153,287]" pageId="24" pageNumber="268">
<emphasis id="B94DFC3FA908FF9CFF5C229CFE6AFF33" bold="true" box="[199,484,224,251]" pageId="24" pageNumber="268">Measurements (in mm).</emphasis>
Male
<typeStatus id="54829E8FA908FF9CFDB0229CFD03FF33" box="[555,653,225,251]" pageId="24" pageNumber="268" type="holotype">holotype</typeStatus>
BL = 16.9; HL = 2.8; PL = 2.7; PW = 4.0; TL = 18.7; TW = 4.0; HFL = 14.0; HTL = 13.1.
</paragraph>
</subSubSection>
<caption id="DF4670A5A908FF9CFF0C25F1FC58F801" ID-DOI="http://doi.org/10.5281/zenodo.3938215" ID-Zenodo-Dep="3938215" httpUri="https://zenodo.org/record/3938215/files/figure.png" pageId="24" pageNumber="268" startId="24.[151,250,1932,1957]" targetBox="[218,1374,326,1880]" targetPageId="24">
<paragraph id="8B86202DA908FF9CFF0C25F1FC58F801" blockId="24.[151,1436,1932,1993]" pageId="24" pageNumber="268">
<emphasis id="B94DFC3FA908FF9CFF0C25F1FEAEF86D" bold="true" box="[151,288,1932,1957]" pageId="24" pageNumber="268">FIGURE 18.</emphasis>
<taxonomicName id="4C395BAEA908FF9CFEBD25F0FDDDF86D" authority="Tan &amp; Japir &amp; Chung &amp; Wahab, 2020" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab" authorityYear="2020" box="[294,595,1933,1957]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="24" pageNumber="268" phylum="Arthropoda" rank="species" species="suikei" status="sp. nov." subGenus="Cantotrella">
<emphasis id="B94DFC3FA908FF9CFEBD25F0FE08F86D" box="[294,390,1933,1957]" italics="true" pageId="24" pageNumber="268">Varitrella</emphasis>
(
<emphasis id="B94DFC3FA908FF9CFE0E25F0FD87F86D" box="[405,521,1933,1957]" italics="true" pageId="24" pageNumber="268">Cantotrella</emphasis>
)
<emphasis id="B94DFC3FA908FF9CFD8325F0FDDDF86D" box="[536,595,1933,1957]" italics="true" pageId="24" pageNumber="268">suikei</emphasis>
</taxonomicName>
<emphasis id="B94DFC3FA908FF9CFDC225F0FD24F86D" bold="true" box="[601,682,1933,1957]" pageId="24" pageNumber="268">
<taxonomicNameLabel id="A27E4144A908FF9CFDC225F0FD24F86D" box="[601,682,1933,1957]" pageId="24" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype’s calling song: oscillograms (A, C) and spectrogram (B) of four echemes (A, B) and one echeme (C) and power spectrum of one echeme (D).
</paragraph>
</caption>
<subSubSection id="C32373A6A909FF9DFF5C22E4FF65FEAF" pageId="25" pageNumber="269" type="etymology">
<paragraph id="8B86202DA909FF9DFF5C22E4FAB0FF7B" blockId="25.[151,1437,153,359]" box="[199,1342,153,179]" pageId="25" pageNumber="269">
<emphasis id="B94DFC3FA909FF9DFF5C22E4FEC0FF7B" bold="true" box="[199,334,153,179]" pageId="25" pageNumber="269">Etymology.</emphasis>
This new species is named after Sui-Kei (first name) Lee (last name), noun in apposition.
</paragraph>
<paragraph id="8B86202DA909FF9DFF5C22C1FF65FEAF" blockId="25.[151,1437,153,359]" pageId="25" pageNumber="269">
<emphasis id="B94DFC3FA909FF9DFF5C22C1FE5AFF1F" bold="true" box="[199,468,188,215]" pageId="25" pageNumber="269">
Calling song (
<figureCitation id="13023CA8A909FF9DFEF422C1FE48FF1F" box="[367,454,188,215]" captionStart="FIGURE 18" captionStartId="24.[151,250,1932,1957]" captionTargetBox="[218,1374,326,1880]" captionTargetId="figure@24.[214,1374,309,1907]" captionTargetPageId="24" captionText="FIGURE 18. Varitrella (Cantotrella) suikei sp. nov. male holotype’s calling song: oscillograms (A, C) and spectrogram (B) of four echemes (A, B) and one echeme (C) and power spectrum of one echeme (D)." figureDoi="http://doi.org/10.5281/zenodo.3938215" httpUri="https://zenodo.org/record/3938215/files/figure.png" pageId="25" pageNumber="269">Fig. 18</figureCitation>
).
</emphasis>
The song consists of doublets of syllables at length to form a continuous trill. Each syllable has a duration of 25.1±2.0 ms (21.9–28.1 ms); each doublet has a duration of 69.4±2.2 ms (67.2–73.4 ms) and period of 320.4±62.4 ms (246.9–417.2 ms). The spectrum shows a dominant peak at 6.44±0.03 kHz (also the fundamental frequency), followed by the second dominant peak at 12.85±0.13 kHz and several peaks harmonically related.
</paragraph>
</subSubSection>
</treatment>
</document>
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